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<mods:title id="5111C22E2A89D2EDF3765AB7E0077017">The genus Cerithidea Swainson, 1840 (Gastropoda: Potamididae) in the Indo-West Pacific region</mods:title>
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<paragraph id="8BCCB63565142868D1A0FD0DFDEE8CE4" blockId="46.[151,593,727,787]" box="[151,593,727,754]" pageId="46" pageNumber="47">
<heading id="D084015965142868D1A0FD0DFDEE8CE4" bold="true" box="[151,593,727,754]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<taxonomicName id="4C73CDB665142868D1A0FD0DFDEE8CE4" ID-CoL="SYLH" authority="Mabille, 1887" authorityName="Mabille" authorityYear="1887" box="[151,593,727,754]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="B9076A2765142868D1A0FD0DFDEE8CE4" bold="true" box="[151,593,727,754]" pageId="46" pageNumber="47">
<emphasis id="B9076A2765142868D1A0FD0DFE1E8CE7" bold="true" box="[151,417,727,753]" italics="true" pageId="46" pageNumber="47">Cerithidea tonkiniana</emphasis>
<bibRefCitation id="EFE2CBC465142868D09FFD02FDEE8CE4" author="Mabille" box="[424,593,728,754]" pageId="46" pageNumber="47" refString="Mabille, J. (1887) Sur quelques mollusques du Tonkin. Bulletins de la Societe Malacologique de France, 4, 73 - 164." type="journal article" year="1887">Mabille, 1887</bibRefCitation>
</emphasis>
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<paragraph id="8BCCB63565142868D1A9FD20FE7E8D05" blockId="46.[151,593,727,787]" box="[158,449,762,787]" pageId="46" pageNumber="47">
(
<figureCitation id="1348AAB065142868D192FD20FEAF8D05" box="[165,272,762,787]" captionStart="FIGURE 2" captionStartId="13.[151,250,1873,1895]" captionTargetBox="[188,1412,193,1848]" captionTargetId="figure@13.[188,1412,193,1849]" captionTargetPageId="13" captionText="FIGURE 2. Living animals of Cerithidea species (not to scale). A. C. charbonnieri, Tapi R. estuary, Surat Thani, Thailand. B. C. obtusa, W Johore, Malaysia. C. C. anticipata, Brunswick R., NSW, Australia. D. C. quoyii, Singapore. E. C. moerchii, Kumamoto City, Kumamoto Pref., Kyushu, Japan. F. C. tonkiniana, Kumamoto City, Kumamoto Pref., Kyushu, Japan. G. C. reidi, Port Smith, Western Australia. H. C. weyersi, Tha Pom, Krabi, Thailand. Photography credits: C, D. Riek; D, S. K. Tan; E, F, H. Fukuda (from Fukuda, 2000). H, B. Craig." figureDoi="http://doi.org/10.5281/zenodo.10846363" httpUri="https://zenodo.org/record/10846363/files/figure.png" pageId="46" pageNumber="47">Figures 2</figureCitation>
F, 13NY, 14A)
</paragraph>
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<paragraph id="8BCCB63565142868D1A0FC98FDCA8D4E" blockId="46.[151,1437,834,1286]" box="[151,629,834,856]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FC98FDCA8D4E" box="[151,629,834,856]" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FC98FDFE8D4E" ID-CoL="SYLH" authority="Mabille, 1887: 158" authorityName="Mabille" authorityPageNumber="158" authorityYear="1887" box="[151,577,834,856]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="B9076A2765142868D1A0FC98FEC88D4E" box="[151,375,834,856]" italics="true" pageId="46" pageNumber="47">Cerithidea tonkiniana</emphasis>
<treatmentCitation id="0AD2902465142868D049FC98FDFE8D4E" author="Mabille" box="[382,577,834,856]" page="158" pageId="46" pageNumber="47" year="1887">
<bibRefCitation id="EFE2CBC465142868D049FC98FDAE8D4E" author="Mabille" box="[382,529,834,856]" pageId="46" pageNumber="47" refString="Mabille, J. (1887) Sur quelques mollusques du Tonkin. Bulletins de la Societe Malacologique de France, 4, 73 - 164." type="journal article" year="1887">Mabille, 1887</bibRefCitation>
: 158
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159
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</paragraph>
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<subSubSection id="C369E5BE65142868D34BFC98FB5D8D4E" box="[636,1250,834,856]" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BCCB63565142868D34BFC98FB5D8D4E" blockId="46.[151,1437,834,1286]" box="[636,1250,834,856]" pageId="46" pageNumber="47">
(
<materialsCitation id="3B1BBC6865142868D3B5FC98FB688D4E" ID-GBIF-Occurrence="3944783311" box="[642,1239,834,856]" pageId="46" pageNumber="47" typeStatus="syntype">
<collectingCountry id="F364F6A565142868D3B5FC98FD748D4E" box="[642,715,834,856]" name="Vietnam" pageId="46" pageNumber="47">Tonkin</collectingCountry>
[N
<collectingCountry id="F364F6A565142868D3C2FC98FCEE8D4E" box="[757,849,834,856]" name="Vietnam" pageId="46" pageNumber="47">Vietnam</collectingCountry>
]; 3
<typeStatus id="54C8089765142868D241FC99FC708D4E" box="[886,975,835,856]" pageId="46" pageNumber="47" type="syntype">syntypes</typeStatus>
<collectionCode id="ED622EF065142868D2E1FC99FB9C8D4E" LSID="urn:lsid:biocol.org:col:34988" box="[982,1059,835,856]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="46" pageNumber="47">MNHN</collectionCode>
25690,
<figureCitation id="1348AAB065142868D54FFC98FB7E8D4E" box="[1144,1217,834,856]" captionStart="FIGURE 13. A M" captionStartId="44.[151,250,1749,1771]" captionTargetBox="[224,1358,193,1726]" captionTargetId="figure@44.[224,1358,193,1727]" captionTargetPageId="44" captionText="FIGURE 13. A M, Cerithidea decollata. N Y, C. tonkiniana. A, Port Alfred, South Africa (NHMUK 1927.2.9.362). B, Africa (NHMUK 20130256). C, D, I, SE Pemba Bay, Cabo Delgado Prov., Mozambique (NHMUK 20060485). J M, Murex decollatus Linnaeus, 1767, neotype, SE Pemba Bay, Cabo Delgado Prov., Mozambique (NHMUK 20130229). E, Zwartkops R., South Africa (NHMUK 20130257). F, Umgeni R., Durban, South Africa (NHMUK 20041433). G, Toliara, Madagascar (NHMUK 20130258). H, no locality (NHMUK 20130259). N, O, Ta Mai Tuk, Tolo Harbour, Hong Kong (NHMUK 20130261). P, Sai Kung, Hong Kong (NHMUK 20070380). Q, C. tonkiniana Mabille, 1887, syntype, Tonkin, Vietnam (MNHN 25690). R, Hai Phong, Vietnam (NHMUK 20130260). S W, Ukidono-Kobashi, Yorimo R., Touruta-Shinden, Usa, Oita Pref., Japan (NHMUK 20070383; T W views of same specimen). X, Y, Hikawa R., Yatsushiro-gun, Kumamoto Pref., Japan (NHMUK 20130262)." figureDoi="http://doi.org/10.5281/zenodo.10846383" httpUri="https://zenodo.org/record/10846383/files/figure.png" pageId="46" pageNumber="47">Fig. 13</figureCitation>
Q
</materialsCitation>
].
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<bibRefCitation id="EFE2CBC465142868D5DEFC98FA278D4E" author="Reid" box="[1257,1432,834,856]" pageId="46" pageNumber="47" refString="Reid, D. G., Claremont, M., Smith, L., Shamoto, M., Glaubrecht, M. &amp; Ozawa, T. (2013) Mosaics in the mangroves: allopatric diversification of tree-climbing mudwhelks (Gastropoda: Potamididae: Cerithidea) in the Indo-West Pacific. Biological Journal of the Linnean Society, 110, 564 - 580." type="journal article" year="2013">
Reid
<emphasis id="B9076A2765142868D416FC99FAF08D4E" box="[1313,1359,834,856]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
., 2013
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: figs 1 (phylogeny), 2 (map).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCCB63565142868D1A0FC5AFD568D80" blockId="46.[151,1437,834,1286]" box="[151,745,896,918]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FC5AFD568D80" box="[151,745,896,918]" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FC5AFDB28D80" ID-CoL="7PD76" box="[151,525,896,918]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Mollusca" rank="genus">
<emphasis id="B9076A2765142868D1A0FC5AFEC28D80" box="[151,381,896,918]" italics="true" pageId="46" pageNumber="47">Potamides (Cerithidea</emphasis>
?
<emphasis id="B9076A2765142868D0BFFC5BFDB28D80" box="[392,525,896,918]" italics="true" pageId="46" pageNumber="47">) tonkinianus</emphasis>
</taxonomicName>
<treatmentCitation id="0AD2902465142868D310FC5AFD5A8D80" author="Fischer" box="[551,741,896,918]" page="163" pageId="46" pageNumber="47" year="1891">
<bibRefCitation id="EFE2CBC465142868D310FC5AFD0D8D80" author="Fischer" box="[551,690,896,918]" pageId="46" pageNumber="47" refString="Fischer, P. (1891) Catalogue et distribution geographique des mollusques terrestres, fluviatiles et marins d'une partie de l'Indo- Chine (Siam, Laos, Cambodge, Annam, Tonkin). Bulletin de la Societe d'Histoire Naturelle d'Autun, 4, 84 - 276." type="journal article" year="1891">Fischer, 1891</bibRefCitation>
: 163
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BCCB63565142868D1A0FC44FC3F8DA2" blockId="46.[151,1437,834,1286]" box="[151,896,926,948]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FC44FC3F8DA2" box="[151,896,926,948]" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FC44FDBE8DA2" ID-CoL="7PD76" box="[151,513,926,948]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="tonkinianus" subGenus="Cerithidea">
<emphasis id="B9076A2765142868D1A0FC44FDBE8DA2" box="[151,513,926,948]" italics="true" pageId="46" pageNumber="47">Potamides (Cerithidea) tonkinianus</emphasis>
</taxonomicName>
<treatmentCitation id="0AD2902465142868D32CFC44FCC38DA2" author="Fischer" box="[539,892,926,948]" page="416" pageId="46" pageNumber="47" year="1904">
<bibRefCitation id="EFE2CBC465142868D32CFC44FCF68DA2" author="Fischer" box="[539,841,926,948]" pageId="46" pageNumber="47" refString="Fischer, H. &amp; Dautzenberg, P. (1904) Catalogue des mollusqes terrestres et fluviatiles de l'Indo-Chine orientale cites jusqu'a ce jour. In: Pavie, A. (Ed.), Mission Pavie Indo-Chine 1879 - 1895. Etudes Diverses. III. Recherches sur l'Histoire Naturelle de l'Indo-Chine Orientale. E. Leroux, Paris, pp. 390 - 450." type="book chapter" year="1904">Fischer &amp; Dautzenberg, 1904</bibRefCitation>
: 416
</treatmentCitation>
.
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</paragraph>
<paragraph id="8BCCB63565142868D1A0FC67FB948DC5" blockId="46.[151,1437,834,1286]" box="[151,1067,957,979]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FC67FB948DC5" box="[151,1067,957,979]" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FC67FDC48DC5" ID-CoL="5XLDM" box="[151,635,957,979]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="sinensis" subGenus="Cerithidea">
<emphasis id="B9076A2765142868D1A0FC67FE628DC5" box="[151,477,957,979]" italics="true" pageId="46" pageNumber="47">Cerithidea (Cerithidea) sinensis</emphasis>
var.
<emphasis id="B9076A2765142868D338FC67FDC48DC5" box="[527,635,957,979]" italics="true" pageId="46" pageNumber="47">tonkiniana</emphasis>
</taxonomicName>
<treatmentCitation id="0AD2902465142868D3A2FC67FC4C8DC5" author="Dautzenberg" box="[661,1011,957,979]" page="137" pageId="46" pageNumber="47" year="1905">
<bibRefCitation id="EFE2CBC465142868D3A2FC67FC7D8DC5" author="Dautzenberg" box="[661,962,957,979]" pageId="46" pageNumber="47" refString="Dautzenberg, P. &amp; Fischer, H. (1905) Liste des mollusques recoltes par M. le Capitaine de Fregate Blaise au Tonkin, et description d'especes nouvelles. Journal de Conchyliologie, 53, 85 - 234." type="journal article" year="1905">Dautzenberg &amp; Fischer, 1905</bibRefCitation>
: 137
</treatmentCitation>
138.
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</paragraph>
<paragraph id="8BCCB63565142868D1A0FC06FBCA8A58" blockId="46.[151,1437,834,1286]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FC06FBCA8A58" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FC06FEF28DE4" ID-CoL="92WXD" box="[151,333,988,1010]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="ornata">
<emphasis id="B9076A2765142868D1A0FC06FEF28DE4" box="[151,333,988,1010]" italics="true" pageId="46" pageNumber="47">Cerithidea ornata</emphasis>
</taxonomicName>
<treatmentCitation id="0AD2902465142868D051FC06FDCF8DE4" author="Kuroda" box="[358,624,988,1010]" page="44" pageId="46" pageNumber="47" year="1952">
<bibRefCitation id="EFE2CBC465142868D051FC06FDF58DE4" author="Kuroda" box="[358,586,988,1010]" pageId="46" pageNumber="47" refString="Kuroda, T. &amp; Habe, T. (1952) Check List and Bibliography of the Recent Marine Mollusca of Japan. L. W. Stach, Tokyo, 210 pp." type="book" year="1952">Kuroda &amp; Habe, 1952</bibRefCitation>
: 44
</treatmentCitation>
.
<treatmentCitation id="0AD2902465142868D34DFC06FC398DE4" author="Habe" box="[634,902,988,1010]" page="34" pageId="46" pageNumber="47" year="1967">
<bibRefCitation id="EFE2CBC465142868D34DFC06FCDF8DE4" author="Habe" box="[634,864,988,1010]" pageId="46" pageNumber="47" refString="Habe, T. &amp; Kosuge, S. (1967) The Standard Book of Japanese Shells in Colour, Vol. III. Hoikusha, Osaka, xviii + 223 pp." type="book" year="1967">Habe &amp; Kosuge, 1967</bibRefCitation>
: 34
</treatmentCitation>
, pl. 13, fig. 17.
<treatmentCitation id="0AD2902465142868D511FC06FB7C8DE4" author="Habe" box="[1062,1219,988,1010]" page="38" pageId="46" pageNumber="47" year="1968">
<bibRefCitation id="EFE2CBC465142868D511FC06FB208DE4" author="Habe" box="[1062,1183,988,1010]" pageId="46" pageNumber="47" refString="Habe, T. (1968) Shells of the Western Pacific in Color. Vol. 2. 2 nd ed. Hoikusha, Osaka, 233 pp." type="book" year="1968">Habe, 1968</bibRefCitation>
: 38
</treatmentCitation>
, pl. 11, fig. 24.
<treatmentCitation id="0AD2902465142868D455FC06FE9C8A06" author="Higo" page="59" pageId="46" pageNumber="47" year="1973">
<bibRefCitation id="EFE2CBC465142868D455FC06FF428A06" author="Higo" pageId="46" pageNumber="47" refString="Higo, S. (1973) A Catalogue of Molluscan Fauna of the Japanese Islands and the Adjacent Area. Isahaya, 397 + 61 pp." type="book" year="1973">Higo, 1973</bibRefCitation>
: 59
</treatmentCitation>
.
<treatmentCitation id="0AD2902465142868D01AFC20FE658A06" author="Wells" box="[301,474,1018,1040]" page="145" pageId="46" pageNumber="47" year="1985">
<bibRefCitation id="EFE2CBC465142868D01AFC20FE178A06" author="Wells" box="[301,424,1018,1040]" pageId="46" pageNumber="47" refString="Wells, F. E. (1985) The Potamididae (Mollusca: Gastropoda) of Hong Kong, with an examination of habitat segregation in a small mangrove system. In: Morton, B. &amp; Dudgeon, D. (Eds) Proceedings of the Second International Workshop on the Malacofauna of Hong Kong and Southern China, Hong Kong. Hong Kong University Press, Hong Kong, pp. 139 - 154." type="book chapter" year="1985">Wells, 1985</bibRefCitation>
: 145
</treatmentCitation>
, pl. 1
<treatmentCitation id="0AD2902465142868D339FC20FD698A06" author="Choe" box="[526,726,1018,1040]" page="310" pageId="46" pageNumber="47" year="1992">
<bibRefCitation id="EFE2CBC465142868D339FC20FD1A8A06" author="Choe" box="[526,677,1018,1040]" pageId="46" pageNumber="47" refString="Choe, B. L. (1992) Mollusca (II). Illustrated Encyclopaedia of Fauna and Flora of Korea. Vol. 33. Ministry of Education, Republic of Korea, 860 pp." type="book" year="1992">G. Choe, 1992</bibRefCitation>
: 310
</treatmentCitation>
311, pl. 65, fig. 103a, b.
<treatmentCitation id="0AD2902465142868D2EDFC20FB6B8A06" author="Higo" box="[986,1236,1018,1040]" page="105" pageId="46" pageNumber="47" year="1993">
<bibRefCitation id="EFE2CBC465142868D2EDFC20FB1D8A06" author="Higo" box="[986,1186,1018,1040]" pageId="46" pageNumber="47" refString="Higo, S. &amp; Goto, Y. (1993) A Systematic List of Molluscan Shells from the Japanese Islands and the Adjacent Areas. Eru Malacological Publisher, Osaka, 3 + 22 + 693 + 13 + 149 pp." type="book" year="1993">Higo &amp; Goto, 1993</bibRefCitation>
: 105
</treatmentCitation>
.
<treatmentCitation id="0AD2902465142868D5E9FC20FED98A39" author="Higo" page="87" pageId="46" pageNumber="47" year="1999">
<bibRefCitation id="EFE2CBC465142868D5E9FC20FE818A39" author="Higo" pageId="46" pageNumber="47" refString="Higo, S., Callomon, P. &amp; Goto, Y. (1999) Catalogue and Bibliography of the Marine Shell-Bearing Mollusca of Japan. Elle Scientific Publications, Osaka, 749 pp." type="book" year="1999">Higo, Callomon &amp; Goto, 1999</bibRefCitation>
: 87
</treatmentCitation>
.
<treatmentCitation id="0AD2902465142868D045FBC0FDDF8A39" author="Lee" box="[370,608,1049,1071]" page="103" pageId="46" pageNumber="47" year="2002">
<bibRefCitation id="EFE2CBC465142868D045FBC0FD948A39" author="Lee" box="[370,555,1049,1071]" pageId="46" pageNumber="47" refString="Lee, J. - S. &amp; Min, D. - K. (2002) A catalogue of molluscan fauna in Korea. Korean Journal of Malacology, 18, 93 - 217." type="journal article" year="2002">Lee &amp; Min, 2002</bibRefCitation>
: 103
</treatmentCitation>
.
<treatmentCitation id="0AD2902465142868D35BFBC0FD468A39" author="Ma" box="[620,761,1049,1071]" page="38" pageId="46" pageNumber="47" year="2004">
<bibRefCitation id="EFE2CBC465142868D35BFBC0FD6D8A39" author="Ma" box="[620,722,1049,1071]" pageId="46" pageNumber="47" refString="Ma, X. (2004) Order Mesogastropoda. In: Qi, Z. (Ed.), Seashells of China. China Ocean Press, Beijing, pp. 31 - 81." type="book chapter" year="2004">Ma, 2004</bibRefCitation>
: 38
</treatmentCitation>
, pl. 016A (not 016C).
<treatmentCitation id="0AD2902465142868D2D0FBC3FB568A39" author="Kojima" box="[999,1257,1049,1071]" page="525" pageId="46" pageNumber="47" year="2006">
<bibRefCitation id="EFE2CBC465142868D2D0FBC3FB098A39" author="Kojima" box="[999,1206,1049,1071]" pageId="46" pageNumber="47" refString="Kojima, S., Kamimura, S., Iijima, A., Kimura, T., Kurozumi, T. &amp; Furota, T. (2006) Molecular phylogeny and population structure of tideland snails in the genus Cerithidea around Japan. Marine Biology, 149, 525 - 535. http: // dx. doi. org / 10.1007 / s 00227 - 005 - 0183 - 2" type="journal article" year="2006">
Kojima
<emphasis id="B9076A2765142868D50BFBC0FBD48A39" box="[1084,1131,1049,1071]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
., 2006
</bibRefCitation>
: 525
</treatmentCitation>
535.
<treatmentCitation id="0AD2902465142868D41FFBC3FE928A58" author="Reid" page="680" pageId="46" pageNumber="47" year="2008">
<bibRefCitation id="EFE2CBC465142868D41FFBC3FF428A58" author="Reid" pageId="46" pageNumber="47" refString="Reid, D. G., Dyal, P., Lozouet, P., Glaubrecht, M. &amp; Williams, S. T. (2008) Mudwhelks and mangroves: the evolutionary history of an ecological association (Gastropoda: Potamididae). Molecular Phylogenetics and Evolution, 47, 680 - 699. http: // dx. doi. org / 10.1016 / j. ympev. 2008.01.003" type="journal article" year="2008">
Reid
<emphasis id="B9076A2765142868D456FBC0FA308A39" box="[1377,1423,1049,1071]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
., 2008
</bibRefCitation>
: 680
</treatmentCitation>
699 (in part, includes
<taxonomicName id="4C73CDB665142868D320FBE3FD388A58" ID-CoL="SYK5" box="[535,647,1080,1102]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="balteata">
<emphasis id="B9076A2765142868D320FBE3FD388A58" box="[535,647,1080,1102]" italics="true" pageId="46" pageNumber="47">C. balteata</emphasis>
</taxonomicName>
).
<treatmentCitation id="0AD2902465142868D3ABFBE2FCFB8A58" author="Zhang" box="[668,836,1080,1102]" page="61" pageId="46" pageNumber="47" year="2008">
<bibRefCitation id="EFE2CBC465142868D3ABFBE2FC9E8A58" author="Zhang" box="[668,801,1080,1102]" pageId="46" pageNumber="47" refString="Zhang, S. (2008) Atlas of Marine Mollusks in China. Hai yang chu ban she, Beijing, iv + 383 pp." type="book" year="2008">Zhang, 2008</bibRefCitation>
: 61
</treatmentCitation>
, fig. (All not
<bibRefCitation id="EFE2CBC465142868D2F9FBE2FBD58A58" author="Sowerby" box="[974,1130,1080,1102]" pageId="46" pageNumber="47" refString="Sowerby, G. B. II (1855) Monograph of the genus Cerithium, Adanson. In: Sowerby, G. B. II. (Ed.), Thesaurus Conchyliorum, or Figures and Descriptions of Recent Shells. Vol. 2. part 16. Sowerby, London, pp. 847 - 899." type="book chapter" year="1855">Sowerby, 1855</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCCB63565142868D1A0FB8CFBD08ABC" blockId="46.[151,1437,834,1286]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FB8CFBD08ABC" bridged="true" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FB8CFE6B8A7B" ID-CoL="92WXD" box="[151,468,1110,1133]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="ornata" subGenus="Cerithidea">
<emphasis id="B9076A2765142868D1A0FB8CFE6B8A7B" box="[151,468,1110,1133]" italics="true" pageId="46" pageNumber="47">Cerithidea (Cerithidea) ornata</emphasis>
</taxonomicName>
<bibRefCitation id="EFE2CBC465142868D0D8FB8CFDC48A7A" author="Oyama" box="[495,635,1110,1132]" pageId="46" pageNumber="47" refString="Oyama, K. (1959) Cerithidea. In: Oyama, K. (Ed.), The Molluscan Shells, Vol. 2. Resources Exploitation Institute, Tokyo, unpaginated." type="book" year="1959">Oyama, 1959</bibRefCitation>
: figs 7, 14.
<treatmentCitation id="0AD2902465142868D3CEFB8CFC0E8A7A" author="Fukuda" box="[761,945,1110,1132]" page="21" pageId="46" pageNumber="47" year="1996">
<bibRefCitation id="EFE2CBC465142868D3CEFB8CFC358A7A" author="Fukuda" box="[761,906,1110,1132]" pageId="46" pageNumber="47" refString="Fukuda, H. (1996) Gastropoda. In: Hanawa, S. &amp; Sakuma, H. (Eds.), Present Status of Estuarine Locales and Benthic Invertebrates Occurring in Estuarine Environment in Japan. WWF Japan Science Report, 3, 11 - 63." type="journal article" year="1996">Fukuda, 1996</bibRefCitation>
: 21
</treatmentCitation>
, pl. 3, fig. 4.
<treatmentCitation id="0AD2902465142868D575FB8CFABB8A7A" author="Fukuda" box="[1090,1284,1110,1132]" page="106" pageId="46" pageNumber="47" year="2000">
<bibRefCitation id="EFE2CBC465142868D575FB8CFB6D8A7A" author="Fukuda" box="[1090,1234,1110,1132]" pageId="46" pageNumber="47" refString="Fukuda, H. (2000) Gastropods I: general view. In: Sato, M. (Ed.), Life in Ariake Sea: Biodiversity in Tidal Flats and Estuaries. Kaiyu-sha, Tokyo, pp. 213 - 252." type="book chapter" year="2000">Fukuda, 2000</bibRefCitation>
: 106
</treatmentCitation>
110, 135, fig. 5.4b (shell), fig. 5.5b (living animal).
<treatmentCitation id="0AD2902465142868D364FBACFC8B8A9D" author="Hasegawa" box="[595,820,1141,1163]" page="133" pageId="46" pageNumber="47" year="2000">
<bibRefCitation id="EFE2CBC465142868D364FBACFCBF8A9D" author="Hasegawa" box="[595,768,1141,1163]" pageId="46" pageNumber="47" refString="Hasegawa, K. (2000) Family Potamididae. In: Okutani, T., (Ed.), Marine Mollusks in Japan. Tokai University Press, Tokyo, pp. 133 - 134." type="book chapter" year="2000">Hasegawa, 2000</bibRefCitation>
: 133
</treatmentCitation>
, pl. 66, fig. 2.
<treatmentCitation id="0AD2902465142868D2E5FBACFAFB8A9D" author="Hong" box="[978,1348,1141,1163]" page="258" pageId="46" pageNumber="47" year="2010">
<bibRefCitation id="EFE2CBC465142868D2E5FBACFAAF8A9D" author="Hong" box="[978,1296,1141,1163]" pageId="46" pageNumber="47" refString="Hong, J. - S., Choi, J. - W. &amp; Tsutsumi, H. (2010) Concluding remarks on the joint survey of macrobenthic fauna on Suncheon Tidal Flats by the participants of &quot; Korea and Japan Joint Symposium on Biology of Tidal Flats 2009 &quot;. Plankton &amp; Benthos Research, 5 (suppl.), 255 - 263. http: // dx. doi. org / 10.3800 / pbr. 5.255" type="journal article" year="2010">Hong, Choi &amp; Tsutsumi, 2010</bibRefCitation>
: 258
</treatmentCitation>
, fig. 4b. Fukuda &amp; Kimura, 2012: 29, textfigs a (living animal), b, c (shell). (All not
<bibRefCitation id="EFE2CBC465142868D2FFFB4EFBDB8ABC" author="Sowerby" box="[968,1124,1172,1194]" pageId="46" pageNumber="47" refString="Sowerby, G. B. II (1855) Monograph of the genus Cerithium, Adanson. In: Sowerby, G. B. II. (Ed.), Thesaurus Conchyliorum, or Figures and Descriptions of Recent Shells. Vol. 2. part 16. Sowerby, London, pp. 847 - 899." type="book chapter" year="1855">Sowerby, 1855</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCCB63565142868D1A0FB68FAFC8ADE" blockId="46.[151,1437,834,1286]" box="[151,1347,1202,1224]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FB68FAFC8ADE" box="[151,1347,1202,1224]" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FB68FEE88ADE" box="[151,343,1202,1224]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="morchii">
<emphasis id="B9076A2765142868D1A0FB68FEE88ADE" box="[151,343,1202,1224]" italics="true" pageId="46" pageNumber="47">Cerithidea morchii</emphasis>
</taxonomicName>
<treatmentCitation id="0AD2902465142868D046FB69FDEB8ADE" author="Lee" box="[369,596,1202,1224]" page="103" pageId="46" pageNumber="47" year="2002">
<bibRefCitation id="EFE2CBC465142868D046FB69FD9B8ADE" author="Lee" box="[369,548,1202,1224]" pageId="46" pageNumber="47" refString="Lee, J. - S. &amp; Min, D. - K. (2002) A catalogue of molluscan fauna in Korea. Korean Journal of Malacology, 18, 93 - 217." type="journal article" year="2002">Lee &amp; Min, 2002</bibRefCitation>
: 103
</treatmentCitation>
(not
<bibRefCitation id="EFE2CBC465142868D3BEFB68FC9B8ADE" author="Sowerby" box="[649,804,1202,1224]" pageId="46" pageNumber="47" refString="Sowerby, G. B. II (1855) Monograph of the genus Cerithium, Adanson. In: Sowerby, G. B. II. (Ed.), Thesaurus Conchyliorum, or Figures and Descriptions of Recent Shells. Vol. 2. part 16. Sowerby, London, pp. 847 - 899." type="book chapter" year="1855">Sowerby, 1855</bibRefCitation>
).
<treatmentCitation id="0AD2902465142868D202FB68FBB28ADE" author="Min" box="[821,1037,1202,1224]" page="129" pageId="46" pageNumber="47" year="2004">
<bibRefCitation id="EFE2CBC465142868D202FB68FC648ADE" author="Min" box="[821,987,1202,1224]" pageId="46" pageNumber="47" refString="Min, D. - K., Lee, J. - S., Koh, D. - B. &amp; Je, J. - G. (2004) Mollusks in Korea. Min Molluscan Research Institute, Seoul, 566 pp." type="book" year="2004">
Min
<emphasis id="B9076A2765142868D251FB69FC2C8ADE" box="[870,915,1202,1224]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
., 2004
</bibRefCitation>
: 129
</treatmentCitation>
, fig. 220 (not
<bibRefCitation id="EFE2CBC465142868D5ABFB68FA888ADE" author="Sowerby" box="[1180,1335,1202,1224]" pageId="46" pageNumber="47" refString="Sowerby, G. B. II (1855) Monograph of the genus Cerithium, Adanson. In: Sowerby, G. B. II. (Ed.), Thesaurus Conchyliorum, or Figures and Descriptions of Recent Shells. Vol. 2. part 16. Sowerby, London, pp. 847 - 899." type="book chapter" year="1855">Sowerby, 1855</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCCB63565142868D1A0FB0BFBD68AF1" blockId="46.[151,1437,834,1286]" box="[151,1129,1233,1255]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FB0BFBD68AF1" box="[151,1129,1233,1255]" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FB0BFE658AF1" ID-CoL="SYLL" box="[151,474,1233,1255]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="weyersi" subGenus="Cerithidea">
<emphasis id="B9076A2765142868D1A0FB0BFE658AF1" box="[151,474,1233,1255]" italics="true" pageId="46" pageNumber="47">Cerithidea (Cerithidea) weyersi</emphasis>
</taxonomicName>
<treatmentCitation id="0AD2902465142868D0C2FB0BFD248AF1" author="Thach" box="[501,667,1233,1255]" page="60" pageId="46" pageNumber="47" year="2007">
<bibRefCitation id="EFE2CBC465142868D0C2FB0BFDCA8AF1" author="Thach" box="[501,629,1233,1255]" pageId="46" pageNumber="47" refString="Thach, N. N. (2007) Recently Collected Shells of Vietnam. L'Informatore Piceno, Ancona, Italy, 384 pp." type="book" year="2007">Thach, 2007</bibRefCitation>
: 60
</treatmentCitation>
, pl. 9, figs 173, 181 (not
<bibRefCitation id="EFE2CBC465142868D2AEFB0BFBE28AF1" author="Dautzenberg" box="[921,1117,1233,1255]" pageId="46" pageNumber="47" refString="Dautzenberg, P. (1899) Contribution a la faune malacologique de Sumatra (recoltes de M. J. - L. Weyers). Annales de la Societe Royale Malacologique de Belgique, 34, 1 - 26." type="journal article" year="1899">Dautzenberg, 1899</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCCB63565142868D1A0FB2AFB818B10" blockId="46.[151,1437,834,1286]" box="[151,1086,1264,1286]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="AB63911B65142868D1A0FB2AFB818B10" box="[151,1086,1264,1286]" pageId="46" pageNumber="47">
<taxonomicName id="4C73CDB665142868D1A0FB2AFEE68B10" ID-CoL="5XLDM" box="[151,345,1264,1286]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="sinensis">
<emphasis id="B9076A2765142868D1A0FB2AFEE68B10" box="[151,345,1264,1286]" italics="true" pageId="46" pageNumber="47">Cerithidea sinensis</emphasis>
</taxonomicName>
<treatmentCitation id="0AD2902465142868D044FB2AFD838B10" author="Li" box="[371,572,1264,1286]" page="452" pageId="46" pageNumber="47" year="2013">
<bibRefCitation id="EFE2CBC465142868D044FB2AFDB48B10" author="Li" box="[371,523,1264,1286]" pageId="46" pageNumber="47" refString="Li, Y. &amp; Xu, R. (2013) Brackish water snails from Qi'ao-Dan'gan Island in the Pearl River estuary, China. Turkish Journal of Zoology, 37, 449 - 457." type="journal article" year="2013">Li &amp; Xu, 2013</bibRefCitation>
: 452
</treatmentCitation>
453, figs 8 (shell), 9 (radula) (not
<bibRefCitation id="EFE2CBC465142868D2A8FB2AFB8C8B10" author="Philippi" box="[927,1075,1264,1286]" pageId="46" pageNumber="47" refString="Philippi, R. A. (1848) testaceorum novorum centuria (continuatio). Zeitschrift fur Malakozoologie, 1848, 17 - 27." type="journal article" year="1848">Philippi, 1848</bibRefCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BCCB63565142868D1A0FAE8FAA48BE9" blockId="46.[151,1436,1330,2003]" pageId="46" pageNumber="47">
<emphasis id="B9076A2765142868D1A0FAE8FE3D8B5D" bold="true" box="[151,386,1330,1355]" pageId="46" pageNumber="47">Taxonomic history.</emphasis>
The original description by
<bibRefCitation id="EFE2CBC465142868D3D5FAE8FC2D8B5D" author="Mabille" box="[738,914,1330,1355]" pageId="46" pageNumber="47" refString="Mabille, J. (1887) Sur quelques mollusques du Tonkin. Bulletins de la Societe Malacologique de France, 4, 73 - 164." type="journal article" year="1887">Mabille (1887)</bibRefCitation>
was rather vague and the species was not illustrated. For these reasons the name has not been used since the early twentieth century, until recently resurrected by
<bibRefCitation id="EFE2CBC465142868D077FAA0FDB18B85" box="[320,526,1402,1427]" pageId="46" pageNumber="47" refString="Reid, D. G., Claremont, M., Smith, L., Shamoto, M., Glaubrecht, M. &amp; Ozawa, T. (2013) Mosaics in the mangroves: allopatric diversification of tree-climbing mudwhelks (Gastropoda: Potamididae: Cerithidea) in the Indo-West Pacific. Biological Journal of the Linnean Society, 110, 564 - 580." type="journal article">
Reid
<emphasis id="B9076A2765142868D049FAA6FE0E8B85" box="[382,433,1402,1427]" italics="true" pageId="46" pageNumber="47">et al</emphasis>
. (2013)
</bibRefCitation>
.
<bibRefCitation id="EFE2CBC465142868D32CFAA0FCC98B85" author="Dautzenberg" box="[539,886,1402,1427]" pageId="46" pageNumber="47" refString="Dautzenberg, P. &amp; Fischer, H. (1905) Liste des mollusques recoltes par M. le Capitaine de Fregate Blaise au Tonkin, et description d'especes nouvelles. Journal de Conchyliologie, 53, 85 - 234." type="journal article" year="1905">Dautzenberg &amp; Fischer (1905)</bibRefCitation>
considered it to be a variety of
<taxonomicName id="4C73CDB665142868D5D1FAA1FADE8B85" ID-CoL="5XLDM" box="[1254,1377,1403,1427]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="sinensis">
<emphasis id="B9076A2765142868D5D1FAA1FADE8B85" box="[1254,1377,1403,1427]" italics="true" pageId="46" pageNumber="47">C. sinensis</emphasis>
</taxonomicName>
. The species has frequently been described and figured in the recent literature from
<collectingCountry id="F364F6A565142868D520FA45FBE58BA0" box="[1047,1114,1439,1462]" name="Japan" pageId="46" pageNumber="47">Japan</collectingCountry>
,
<collectingCountry id="F364F6A565142868D551FA44FB138BA1" box="[1126,1196,1438,1463]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
,
<collectingCountry id="F364F6A565142868D58FFA45FB438BA0" box="[1208,1276,1439,1462]" name="South Korea" pageId="46" pageNumber="47">Korea</collectingCountry>
and
<collectingCountry id="F364F6A565142868D401FA44FA278BA1" box="[1334,1432,1438,1463]" name="Vietnam" pageId="46" pageNumber="47">Vietnam</collectingCountry>
, although always misidentified. It has usually been named
<taxonomicName id="4C73CDB665142868D20CFA19FC158BCD" ID-CoL="92WXD" box="[827,938,1475,1499]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="ornata">
<emphasis id="B9076A2765142868D20CFA19FC158BCD" box="[827,938,1475,1499]" italics="true" pageId="46" pageNumber="47">C. ornata</emphasis>
</taxonomicName>
(a synonym of the superficially similar
<taxonomicName id="4C73CDB665142868D4B5FA19FF4C8BE9" ID-CoL="SYK5" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="46" pageNumber="47" phylum="Mollusca" rank="species" species="balteata">
<emphasis id="B9076A2765142868D4B5FA19FF4C8BE9" italics="true" pageId="46" pageNumber="47">C. balteata</emphasis>
</taxonomicName>
), following the identification by early Japanese workers (
<bibRefCitation id="EFE2CBC465142868D24FFA3CFBD08BE9" author="Kuroda" box="[888,1135,1510,1535]" pageId="46" pageNumber="47" refString="Kuroda, T. &amp; Habe, T. (1952) Check List and Bibliography of the Recent Marine Mollusca of Japan. L. W. Stach, Tokyo, 210 pp." type="book" year="1952">Kuroda &amp; Habe 1952</bibRefCitation>
;
<bibRefCitation id="EFE2CBC465142868D54CFA3DFAB18BE9" author="Oyama" box="[1147,1294,1510,1535]" pageId="46" pageNumber="47" refString="Oyama, K. (1959) Cerithidea. In: Oyama, K. (Ed.), The Molluscan Shells, Vol. 2. Resources Exploitation Institute, Tokyo, unpaginated." type="book" year="1959">Oyama 1959</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C369E5BE65142868D1F0F9D0FBA68899" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BCCB63565142868D1F0F9D0FBA68899" blockId="46.[151,1436,1330,2003]" pageId="46" pageNumber="47">
<emphasis id="B9076A2765142868D1F0F9D0FEFE8835" bold="true" box="[199,321,1546,1571]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
Shell: periphery rounded or slightly angled; aperture thickened and flared; 1125 rounded axial ribs on penultimate whorl, 411 ribs after ventrolateral varix; ventrolateral varix a strong rib at 180220(250)°; 5 weak spiral ridges above periphery (especially on spire if not eroded); banded or lined pattern. S
<collectingCountry id="F364F6A565142868D5FCF989FAB1887C" box="[1227,1294,1619,1642]" name="Japan" pageId="46" pageNumber="47">Japan</collectingCountry>
, S
<collectingCountry id="F364F6A565142868D401F989FAC2887C" box="[1334,1405,1619,1642]" name="South Korea" pageId="46" pageNumber="47">Korea</collectingCountry>
, S
<collectingCountry id="F364F6A565142868D1A0F9ACFF628899" box="[151,221,1654,1679]" name="China" pageId="46" pageNumber="47">China</collectingCountry>
, N
<collectingCountry id="F364F6A565142868D034F9ACFED98899" box="[259,358,1654,1679]" name="Vietnam" pageId="46" pageNumber="47">Vietnam</collectingCountry>
. COI GenBank
<accessionNumber id="94202BD665142868D32FF9ACFD238899" box="[536,668,1654,1679]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AM932766" pageId="46" pageNumber="47">AM932766</accessionNumber>
,
<accessionNumber id="94202BD665142868D390F9ACFC948899" box="[679,811,1654,1679]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AM932767" pageId="46" pageNumber="47">AM932767</accessionNumber>
,
<accessionNumber id="94202BD665142868D201F9ACFC0F8899" box="[822,944,1654,1679]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/HE680220" pageId="46" pageNumber="47">HE680220</accessionNumber>
680224.
</paragraph>
</subSubSection>
<subSubSection id="C369E5BE65142869D1F0F940FEF58A19" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="8BCCB63565142868D1F0F940FDB588A5" blockId="46.[151,1436,1330,2003]" box="[199,522,1690,1715]" pageId="46" pageNumber="47">
<emphasis id="B9076A2765142868D1F0F940FE0F88A5" bold="true" box="[199,432,1690,1715]" pageId="46" pageNumber="47">Material examined.</emphasis>
15 lots.
</paragraph>
<paragraph id="8BCCB63565142869D1F0F964FF458FBA" blockId="46.[151,1436,1330,2003]" lastBlockId="47.[151,1436,151,1904]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="B9076A2765142868D1F0F964FE0788C1" bold="true" box="[199,440,1726,1751]" pageId="46" pageNumber="47">
Shell (
<figureCitation id="1348AAB065142868D020F964FED088C1" box="[279,367,1726,1751]" captionStart="FIGURE 13. A M" captionStartId="44.[151,250,1749,1771]" captionTargetBox="[224,1358,193,1726]" captionTargetId="figure@44.[224,1358,193,1727]" captionTargetPageId="44" captionText="FIGURE 13. A M, Cerithidea decollata. N Y, C. tonkiniana. A, Port Alfred, South Africa (NHMUK 1927.2.9.362). B, Africa (NHMUK 20130256). C, D, I, SE Pemba Bay, Cabo Delgado Prov., Mozambique (NHMUK 20060485). J M, Murex decollatus Linnaeus, 1767, neotype, SE Pemba Bay, Cabo Delgado Prov., Mozambique (NHMUK 20130229). E, Zwartkops R., South Africa (NHMUK 20130257). F, Umgeni R., Durban, South Africa (NHMUK 20041433). G, Toliara, Madagascar (NHMUK 20130258). H, no locality (NHMUK 20130259). N, O, Ta Mai Tuk, Tolo Harbour, Hong Kong (NHMUK 20130261). P, Sai Kung, Hong Kong (NHMUK 20070380). Q, C. tonkiniana Mabille, 1887, syntype, Tonkin, Vietnam (MNHN 25690). R, Hai Phong, Vietnam (NHMUK 20130260). S W, Ukidono-Kobashi, Yorimo R., Touruta-Shinden, Usa, Oita Pref., Japan (NHMUK 20070383; T W views of same specimen). X, Y, Hikawa R., Yatsushiro-gun, Kumamoto Pref., Japan (NHMUK 20130262)." figureDoi="http://doi.org/10.5281/zenodo.10846383" httpUri="https://zenodo.org/record/10846383/files/figure.png" pageId="46" pageNumber="47">Fig. 13</figureCitation>
NY):
</emphasis>
H = 23.0
<quantity id="4C8B1BD065142868D373F965FD1088C0" box="[580,687,1727,1751]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" pageId="46" pageNumber="47" unit="mm" value="38.1">38.1 mm</quantity>
. Shape elongated conical (H/B = 2.042.31, SH = 2.592.99); decollate, 67 whorls remaining; spire whorls moderately rounded, suture distinct; spire profile mostly straight, but concave in apical whorls that are lost in adults and becoming convex at final whorl; periphery rounded to slightly angled; thin to moderate thickness. Adult lip flared, slightly thickened; rarely a previous lip on final whorl; apertural margin slightly sinuous in side view; moderate anterior projection adjacent to canal. Sculpture on spire of straight to slightly opisthocline axial ribs, becoming slightly curved (opisthocyrt) on final 12 whorls, ribs prominent, rounded, interspaces11.5 times width of ribs, 1125 ribs on penultimate whorl; axial ribs becoming more distant but usually remaining strong after ventrolateral varix, numbering 411; spiral sculpture only visible on spire whorls if well preserved, 5 weak spiral ridges on axial ribs only, plus strong outermost basal cord usually visible in suture; traces of 5 spiral ridges sometimes present above periphery on final whorl, but only in interspaces between axial ribs; base with 913 striae, of which outermost is largest and may form a weak peripheral keel, below which axial ribs do not continue. Ventrolateral varix a thickened rib at 180220(250)°. Surface with fine spiral microstriae on periostracum; ribs dull when worn. Colour: pale brown with dark spiral lines and bands; basic pattern is brown band on base, a line just below periphery and 4 bands at and above periphery, but these may be lost or fused; even in darkest shells there is a pale band at shoulder and below periphery; bands show through in aperture.
</paragraph>
<paragraph id="8BCCB63565152869D1F0FE6DFCB48C0E" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
<emphasis id="B9076A2765152869D1F0FE6DFE268FC6" bold="true" box="[199,409,439,464]" pageId="47" pageNumber="48">
Animal (
<figureCitation id="1348AAB065152869D007FE6DFEC88FC6" box="[304,375,439,464]" captionStart="FIGURE 2" captionStartId="13.[151,250,1873,1895]" captionTargetBox="[188,1412,193,1848]" captionTargetId="figure@13.[188,1412,193,1849]" captionTargetPageId="13" captionText="FIGURE 2. Living animals of Cerithidea species (not to scale). A. C. charbonnieri, Tapi R. estuary, Surat Thani, Thailand. B. C. obtusa, W Johore, Malaysia. C. C. anticipata, Brunswick R., NSW, Australia. D. C. quoyii, Singapore. E. C. moerchii, Kumamoto City, Kumamoto Pref., Kyushu, Japan. F. C. tonkiniana, Kumamoto City, Kumamoto Pref., Kyushu, Japan. G. C. reidi, Port Smith, Western Australia. H. C. weyersi, Tha Pom, Krabi, Thailand. Photography credits: C, D. Riek; D, S. K. Tan; E, F, H. Fukuda (from Fukuda, 2000). H, B. Craig." figureDoi="http://doi.org/10.5281/zenodo.10846363" httpUri="https://zenodo.org/record/10846363/files/figure.png" pageId="47" pageNumber="48">Fig. 2</figureCitation>
F):
</emphasis>
Head grey with cream spots; anterior half of snout blackish with yellow spots arranged in 23 transverse bands; tentacles grey with black rings and black base; sides of foot blackish; sole of foot grey; mantle pale grey (ethanol-preserved specimens similar).
</paragraph>
<paragraph id="8BCCB63565152869D1F0FDFEFDF58CE6" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
<emphasis id="B9076A2765152869D1F0FDFEFE248C2B" bold="true" box="[199,411,548,573]" pageId="47" pageNumber="48">
Range (
<figureCitation id="1348AAB065152869D015FDFEFE338C2B" box="[290,396,548,573]" captionStart="FIGURE 14. A" captionStartId="48.[151,250,1921,1943]" captionTargetPageId="48" captionText="FIGURE 14. A. Distribution of Cerithidea tonkiniana (squares). B. Distribution of C. rhizophorarum (triangles) and of C. moerchii (circles). Solid symbols, specimens examined; open symbols, literature records: A, Thach (2007); B, Ma (2004); C, Noseworthy et al. (2007); D, Lee &amp; Chao (2004); E, Hong et al. (2010); F, Springsteen &amp; Leobrera (1986)." figureDoi="http://doi.org/10.5281/zenodo.10846385" httpUri="https://zenodo.org/record/10846385/files/figure.png" pageId="47" pageNumber="48">Fig. 14A</figureCitation>
):
</emphasis>
S
<collectingCountry id="F364F6A565152869D080FDFFFE458C2A" box="[439,506,549,572]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
, S
<collectingCountry id="F364F6A565152869D32CFDFFFDDD8C2A" box="[539,610,549,572]" name="South Korea" pageId="47" pageNumber="48">Korea</collectingCountry>
, S
<collectingCountry id="F364F6A565152869D3B3FDFEFD758C2B" box="[644,714,548,573]" name="China" pageId="47" pageNumber="48">China</collectingCountry>
,
<collectingCountry id="F364F6A565152869D3E3FDFEFC888C2B" box="[724,823,548,573]" name="Vietnam" pageId="47" pageNumber="48">Vietnam</collectingCountry>
. Records:
<materialsCitation id="3B1BBC6865152869D29CFDFFFA238C2A" ID-GBIF-Occurrence="3944783409" box="[939,1436,548,573]" pageId="47" pageNumber="48">
<collectingCountry id="F364F6A565152869D29CFDFFFC508C2A" box="[939,1007,549,572]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
: Tosa Prov., Shikoku (
<collectionCode id="ED622EF065152869D5DFFDFEFA8E8C2A" LSID="urn:lsid:biocol.org:col:34784" box="[1256,1329,548,572]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34784" name="Academy of Natural Sciences of Philadelphia" pageId="47" pageNumber="48">ANSP</collectionCode>
<date id="FFCD90F565152869D40FFDFEFA2F8C2A" box="[1336,1424,548,572]" pageId="47" pageNumber="48" value="29-11-19">191129</date>
);
</materialsCitation>
<materialsCitation id="3B1BBC6865152869D1A0FD9DFAB98C76" ID-GBIF-Occurrence="3944783321" box="[151,1286,583,608]" pageId="47" pageNumber="48">
Ukidono-kobashi, Yorimo R., Touruta-shinden, Usa, Oita Pref., Kyushu (
<collectionCode id="ED622EF065152869D53DFD92FBC98C76" LSID="urn:lsid:biocol.org:col:34665" box="[1034,1142,584,608]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="47" pageNumber="48">NHMUK</collectionCode>
20070383);
</materialsCitation>
<materialsCitation id="3B1BBC6865152869D422FD9DFCD08C92" ID-GBIF-Occurrence="3944783378" pageId="47" pageNumber="48">
Hikawa R., Yatsushiro-gun, Kumamoto Pref., Kyushu (
<collectionCode id="ED622EF065152869D34BFDB6FD588C92" LSID="urn:lsid:biocol.org:col:34665" box="[636,743,620,644]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="47" pageNumber="48">NHMUK</collectionCode>
20070378).
</materialsCitation>
S
<collectingCountry id="F364F6A565152869D2BCFDB7FC6B8C92" box="[907,980,621,644]" name="South Korea" pageId="47" pageNumber="48">Korea</collectingCountry>
: Suncheon Bay (
<bibRefCitation id="EFE2CBC465152869D5B8FDB6FF6B8CBE" author="Hong" pageId="47" pageNumber="48" refString="Hong, J. - S., Choi, J. - W. &amp; Tsutsumi, H. (2010) Concluding remarks on the joint survey of macrobenthic fauna on Suncheon Tidal Flats by the participants of &quot; Korea and Japan Joint Symposium on Biology of Tidal Flats 2009 &quot;. Plankton &amp; Benthos Research, 5 (suppl.), 255 - 263. http: // dx. doi. org / 10.3800 / pbr. 5.255" type="journal article" year="2010">Hong, Choi &amp; Tsutsumi 2010</bibRefCitation>
).
<collectingCountry id="F364F6A565152869D1DFFD55FE908CBE" box="[232,303,655,680]" name="China" pageId="47" pageNumber="48">China</collectingCountry>
: Zhejiang to Guangxi (Qi 2004);
<materialsCitation id="3B1BBC6865152869D399FD55FA238CBE" ID-GBIF-Occurrence="3944783360" box="[686,1436,655,680]" pageId="47" pageNumber="48">
Tolo Harbour,
<collectingCountry id="F364F6A565152869D262FD4AFC658CBE" box="[853,986,656,680]" name="Hong Kong" pageId="47" pageNumber="48">Hong Kong</collectingCountry>
(
<collectionCode id="ED622EF065152869D2DDFD4AFBE98CBE" LSID="urn:lsid:biocol.org:col:34665" box="[1002,1110,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="47" pageNumber="48">NHMUK</collectionCode>
20130261;
<collectionCode id="ED622EF065152869D5E9FD4AFA908CBE" LSID="urn:lsid:biocol.org:col:34871" box="[1246,1327,656,680]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="47" pageNumber="48">USNM</collectionCode>
858380).
</materialsCitation>
<materialsCitation id="3B1BBC6865152869D1A0FD6EFBC58CDB" ID-GBIF-Occurrence="3944783393" box="[151,1146,692,717]" pageId="47" pageNumber="48">
<collectingCountry id="F364F6A565152869D1A0FD6EFF458CDB" box="[151,250,692,717]" name="Vietnam" pageId="47" pageNumber="48">Vietnam</collectingCountry>
: Quikim Brackish Water Research Station, Hai Phong (
<collectionCode id="ED622EF065152869D2B5FD6EFC528CDA" LSID="urn:lsid:biocol.org:col:34665" box="[898,1005,692,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="47" pageNumber="48">NHMUK</collectionCode>
20130260);
</materialsCitation>
<materialsCitation id="3B1BBC6865152869D5B2FD6FFA238CDA" ID-GBIF-Occurrence="3944783390" box="[1157,1436,692,717]" pageId="47" pageNumber="48">
Ha Long Bay (
<collectionCode id="ED622EF065152869D402FD6EFA308CDA" LSID="urn:lsid:biocol.org:col:34992" box="[1333,1423,692,716]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="47" pageNumber="48">RMNH</collectionCode>
);
</materialsCitation>
Long Hai (
<bibRefCitation id="EFE2CBC465152869D027FD0DFD868CE6" author="Thach" box="[272,569,727,752]" pageId="47" pageNumber="48" refString="Thach, N. N. (2007) Recently Collected Shells of Vietnam. L'Informatore Piceno, Ancona, Italy, 384 pp." type="book" year="2007">
Thach 2007, as
<taxonomicName id="4C73CDB665152869D0F4FD02FD868CE6" box="[451,569,728,752]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="weyersi">
<emphasis id="B9076A2765152869D0F4FD02FD868CE6" box="[451,569,728,752]" italics="true" pageId="47" pageNumber="48">C. weyersi</emphasis>
</taxonomicName>
</bibRefCitation>
).
</paragraph>
<paragraph id="8BCCB63565152869D1F0FD26FE7F8D4B" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
<emphasis id="B9076A2765152869D1F0FD26FE7B8D03" bold="true" box="[199,452,764,789]" pageId="47" pageNumber="48">Habitat and ecology.</emphasis>
On mangrove trunks and branches and on mud and stones under mangroves (
<collectingCountry id="F364F6A565152869D46BFD26FF658D2E" name="Hong Kong" pageId="47" pageNumber="48">Hong Kong</collectingCountry>
); intertidal mudflat with
<taxonomicName id="4C73CDB665152869D0C9FCC5FDC08D2E" box="[510,639,799,824]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="B9076A2765152869D0C9FCC5FDC08D2E" box="[510,639,799,824]" italics="true" pageId="47" pageNumber="48">Phragmites</emphasis>
</taxonomicName>
reeds at river mouth (
<collectingCountry id="F364F6A565152869D2BFFCFAFC718D2E" box="[904,974,800,824]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
); brackish salt marsh at edge of shrimp ponds (northern
<collectingCountry id="F364F6A565152869D067FC9EFE0B8D4B" box="[336,436,836,861]" name="Vietnam" pageId="47" pageNumber="48">Vietnam</collectingCountry>
).
</paragraph>
<paragraph id="8BCCB63565152869D1F0FCB2FEF58A19" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
In
<collectingCountry id="F364F6A565152869D1D1FCB2FED28D96" box="[230,365,872,896]" name="Hong Kong" pageId="47" pageNumber="48">Hong Kong</collectingCountry>
,
<bibRefCitation id="EFE2CBC465152869D04FFCBDFDB68D96" author="Wells" box="[376,521,871,896]" pageId="47" pageNumber="48" refString="Wells, F. E. (1985) The Potamididae (Mollusca: Gastropoda) of Hong Kong, with an examination of habitat segregation in a small mangrove system. In: Morton, B. &amp; Dudgeon, D. (Eds) Proceedings of the Second International Workshop on the Malacofauna of Hong Kong and Southern China, Hong Kong. Hong Kong University Press, Hong Kong, pp. 139 - 154." type="book chapter" year="1985">Wells (1985)</bibRefCitation>
reported it from mangroves at upper tidal levels, attached by mucus to trees, and described it as rare, except at Ting Kok.
<bibRefCitation id="EFE2CBC465152869D36FFC56FCA28DB3" author="McMahon" box="[600,797,908,933]" pageId="47" pageNumber="48" refString="McMahon, R. F. (1985) Interspecific relationships between morphometric parameters and the vertical distribution patterns of seven species of turbinate gastropods on mangrove trees in Hong Kong. In: Morton, B. &amp; Dudgeon, D. (Eds.), Proceedings of the Second International Workshop on the Malacofauna of Hong Kong and Southern China, Hong Kong. Hong Kong University Press, Hong Kong, pp. 199 - 215." type="book chapter" year="1985">McMahon (1985)</bibRefCitation>
likewise observed that it was restricted to the upper parts of the mangrove belt, above the mean tide mark; it was most common at the landward edge and found above 1.0 m on trunks and branches, attached by mucus when dry.
<bibRefCitation id="EFE2CBC465152869D227FC0FFC578DFA" box="[784,1000,980,1005]" pageId="47" pageNumber="48" refString="Hong, J. - S., Choi, J. - W. &amp; Tsutsumi, H. (2010) Concluding remarks on the joint survey of macrobenthic fauna on Suncheon Tidal Flats by the participants of &quot; Korea and Japan Joint Symposium on Biology of Tidal Flats 2009 &quot;. Plankton &amp; Benthos Research, 5 (suppl.), 255 - 263. http: // dx. doi. org / 10.3800 / pbr. 5.255" type="journal article">
Hong
<emphasis id="B9076A2765152869D26FFC0FFC338DFB" box="[856,908,980,1005]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. (2010)
</bibRefCitation>
found it on mud flats among
<taxonomicName id="4C73CDB665152869D47EFC0EFA248DFB" box="[1353,1435,980,1005]" class="Magnoliopsida" family="Amaranthaceae" genus="Suaeda" kingdom="Plantae" order="Caryophyllales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="B9076A2765152869D47EFC0EFA248DFB" box="[1353,1435,980,1005]" italics="true" pageId="47" pageNumber="48">Suaeda</emphasis>
</taxonomicName>
plants in
<collectingCountry id="F364F6A565152869D1CAFC22FEFA8A19" box="[253,325,1016,1039]" name="South Korea" pageId="47" pageNumber="48">Korea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C369E5BE65152869D1F0FBC6FCB78966" pageId="47" pageNumber="48" type="discussion">
<paragraph id="8BCCB63565152869D1F0FBC6FED68B26" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
<emphasis id="B9076A2765152869D1F0FBC6FE848A23" bold="true" box="[199,315,1052,1077]" pageId="47" pageNumber="48">Remarks.</emphasis>
The shell of this species closely resembles that of
<taxonomicName id="4C73CDB665152869D2B6FBC7FC408A23" box="[897,1023,1052,1077]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="balteata">
<emphasis id="B9076A2765152869D2B6FBC7FC408A23" box="[897,1023,1052,1077]" italics="true" pageId="47" pageNumber="48">C. balteata</emphasis>
</taxonomicName>
from the
<collectingCountry id="F364F6A565152869D546FBC6FB4D8A23" box="[1137,1266,1052,1077]" name="Philippines" pageId="47" pageNumber="48">Philippines</collectingCountry>
,
<collectingCountry id="F364F6A565152869D5C9FBC6FAD48A23" box="[1278,1387,1052,1077]" name="Indonesia" pageId="47" pageNumber="48">Indonesia</collectingCountry>
and New
<collectingCountry id="F364F6A565152869D1E4FBE5FE988A4E" box="[211,295,1087,1112]" name="Guinea-Bissau" pageId="47" pageNumber="48">Guinea</collectingCountry>
, and it has usually been identified as
<taxonomicName id="4C73CDB665152869D3F8FB9AFC848A4E" box="[719,827,1088,1112]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="ornata">
<emphasis id="B9076A2765152869D3F8FB9AFC848A4E" box="[719,827,1088,1112]" italics="true" pageId="47" pageNumber="48">C. ornata</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C73CDB665152869D25CFB9AFC588A4E" box="[875,999,1087,1112]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="balteata">
<emphasis id="B9076A2765152869D25CFB9AFC588A4E" box="[875,999,1087,1112]" italics="true" pageId="47" pageNumber="48">C. balteata</emphasis>
</taxonomicName>
). Molecular data show that the two are distinct and belong to separate clades (
<bibRefCitation id="EFE2CBC465152869D365FBBEFCB08A6A" author="Reid" box="[594,783,1124,1149]" pageId="47" pageNumber="48" refString="Reid, D. G., Dyal, P., Lozouet, P., Glaubrecht, M. &amp; Williams, S. T. (2008) Mudwhelks and mangroves: the evolutionary history of an ecological association (Gastropoda: Potamididae). Molecular Phylogenetics and Evolution, 47, 680 - 699. http: // dx. doi. org / 10.1016 / j. ympev. 2008.01.003" type="journal article" year="2008">
Reid
<emphasis id="B9076A2765152869D3A5FBBFFD7B8A6B" box="[658,708,1124,1149]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. 2008
</bibRefCitation>
,
<bibRefCitation id="EFE2CBC465152869D22BFBBEFCE68A6A" author="Reid" box="[796,857,1124,1148]" pageId="47" pageNumber="48" refString="Reid, D. G., Claremont, M., Smith, L., Shamoto, M., Glaubrecht, M. &amp; Ozawa, T. (2013) Mosaics in the mangroves: allopatric diversification of tree-climbing mudwhelks (Gastropoda: Potamididae: Cerithidea) in the Indo-West Pacific. Biological Journal of the Linnean Society, 110, 564 - 580." type="journal article" year="2013">2013</bibRefCitation>
). The most useful characters for separation are the broader shape of
<taxonomicName id="4C73CDB665152869D05BFB52FDB48AB6" box="[364,523,1159,1184]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="B9076A2765152869D05BFB52FDB48AB6" box="[364,523,1159,1184]" italics="true" pageId="47" pageNumber="48">C. tonkiniana</emphasis>
</taxonomicName>
with stronger and more numerous (411, cf.
<quantity id="4C8B1BD065152869D50EFB5DFB388AB6" box="[1081,1159,1159,1184]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="15.24" metricValueMin="0.0" pageId="47" pageNumber="48" unit="in" value="3.0" valueMax="6.0" valueMin="0.0">06 in</quantity>
<taxonomicName id="4C73CDB665152869D5A3FB52FAA88AB6" box="[1172,1303,1159,1184]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="balteata">
<emphasis id="B9076A2765152869D5A3FB52FAA88AB6" box="[1172,1303,1159,1184]" italics="true" pageId="47" pageNumber="48">C. balteata</emphasis>
</taxonomicName>
) axial ribs continuing after the ventrolateral varix; the frequent position of the ventrolateral varix at about 180° (usually about 270° in
<taxonomicName id="4C73CDB665152869D1CFFB0AFEC78AFE" box="[248,376,1231,1256]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="balteata">
<emphasis id="B9076A2765152869D1CFFB0AFEC78AFE" box="[248,376,1231,1256]" italics="true" pageId="47" pageNumber="48">C. balteata</emphasis>
</taxonomicName>
); and the weak spiral ribs on the spire whorls (absent in
<taxonomicName id="4C73CDB665152869D514FB0AFB1B8AFE" box="[1059,1188,1231,1256]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="balteata">
<emphasis id="B9076A2765152869D514FB0AFB1B8AFE" box="[1059,1188,1231,1256]" italics="true" pageId="47" pageNumber="48">C. balteata</emphasis>
</taxonomicName>
). However, the great variability of
<taxonomicName id="4C73CDB665152869D003FB2FFE0D8B1B" box="[308,434,1268,1293]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="balteata">
<emphasis id="B9076A2765152869D003FB2FFE0D8B1B" box="[308,434,1268,1293]" italics="true" pageId="47" pageNumber="48">C. balteata</emphasis>
</taxonomicName>
across its wide range and the often eroded condition of the shells make morphological diagnosis difficult.
</paragraph>
<paragraph id="8BCCB63565152869D1F0FAE6FE2C883B" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
<taxonomicName id="4C73CDB665152869D1F0FAE6FE7F8B43" box="[199,448,1340,1365]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="B9076A2765152869D1F0FAE6FE7F8B43" box="[199,448,1340,1365]" italics="true" pageId="47" pageNumber="48">Cerithidea tonkiniana</emphasis>
</taxonomicName>
is sympatric over much of its range in
<collectingCountry id="F364F6A565152869D2B9FAE6FC6E8B43" box="[910,977,1340,1365]" name="China" pageId="47" pageNumber="48">China</collectingCountry>
and northern
<collectingCountry id="F364F6A565152869D542FAE6FB6B8B43" box="[1141,1236,1340,1365]" name="Vietnam" pageId="47" pageNumber="48">Vietnam</collectingCountry>
with
<taxonomicName id="4C73CDB665152869D42EFAE7FA2A8B42" box="[1305,1429,1341,1364]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="sinensis">
<emphasis id="B9076A2765152869D42EFAE7FA2A8B42" box="[1305,1429,1341,1364]" italics="true" pageId="47" pageNumber="48">C. sinensis</emphasis>
</taxonomicName>
, with a somewhat similar shell (see Remarks on
<taxonomicName id="4C73CDB665152869D39BFABAFC998B6E" box="[684,806,1376,1400]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="sinensis">
<emphasis id="B9076A2765152869D39BFABAFC998B6E" box="[684,806,1376,1400]" italics="true" pageId="47" pageNumber="48">C. sinensis</emphasis>
</taxonomicName>
for discrimination). Its range also overlaps broadly with that of another potamidid,
<taxonomicName id="4C73CDB665152869D0EDFA5EFD418B8B" box="[474,766,1412,1437]" class="Gastropoda" family="Potamididae" genus="Cerithideopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="largillierti">
<emphasis id="B9076A2765152869D0EDFA5EFD418B8B" box="[474,766,1412,1437]" italics="true" pageId="47" pageNumber="48">Cerithideopsis largillierti</emphasis>
</taxonomicName>
. Although superficially similar, the latter species is not decollate, lacks a distinct ventrolateral varix, the aperture is sinuous in side view and is not flared, the periostracum is thicker with strong slightly bristly striae, the whorls are more rounded and the axial ribs are often more numerous (2026 on penultimate whorl); the living animal is black with yellow snout tip and tentacle bases, and does not climb onto vegetation.
</paragraph>
<paragraph id="8BCCB63565152869D1F0F9E3FBB488AB" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
In molecular analyses
<taxonomicName id="4C73CDB665152869D0F3F9E2FDE18846" box="[452,606,1591,1616]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="B9076A2765152869D0F3F9E2FDE18846" box="[452,606,1591,1616]" italics="true" pageId="47" pageNumber="48">C. tonkiniana</emphasis>
</taxonomicName>
forms a clade with
<taxonomicName id="4C73CDB665152869D208F9E2FC7C8846" box="[831,963,1591,1616]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="moerchii">
<emphasis id="B9076A2765152869D208F9E2FC7C8846" box="[831,963,1591,1616]" italics="true" pageId="47" pageNumber="48">C. moerchii</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFE2CBC465152869D2E5F9EDFB2C8846" author="Reid" box="[978,1171,1591,1616]" pageId="47" pageNumber="48" refString="Reid, D. G., Dyal, P., Lozouet, P., Glaubrecht, M. &amp; Williams, S. T. (2008) Mudwhelks and mangroves: the evolutionary history of an ecological association (Gastropoda: Potamididae). Molecular Phylogenetics and Evolution, 47, 680 - 699. http: // dx. doi. org / 10.1016 / j. ympev. 2008.01.003" type="journal article" year="2008">
Reid
<emphasis id="B9076A2765152869D526F9E3FBFC8846" box="[1041,1091,1591,1616]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
., 2008
</bibRefCitation>
,
<bibRefCitation id="EFE2CBC465152869D5A8F9E2FB658846" author="Reid" box="[1183,1242,1592,1616]" pageId="47" pageNumber="48" refString="Reid, D. G., Claremont, M., Smith, L., Shamoto, M., Glaubrecht, M. &amp; Ozawa, T. (2013) Mosaics in the mangroves: allopatric diversification of tree-climbing mudwhelks (Gastropoda: Potamididae: Cerithidea) in the Indo-West Pacific. Biological Journal of the Linnean Society, 110, 564 - 580." type="journal article" year="2013">2013</bibRefCitation>
;
<figureCitation id="1348AAB065152869D5D0F9EDFA988846" box="[1255,1319,1591,1616]" captionStart="FIGURE 1" captionStartId="11.[151,250,1936,1958]" captionTargetBox="[203,1385,201,1916]" captionTargetId="figure@11.[196,1387,193,1917]" captionTargetPageId="11" captionText="FIGURE 1. Molecular phylogeny of Cerithidea species produced by Bayesian analysis of partial COI sequences, using Cerithideopsilla species as outgroup (after Reid et al. 2013). Localities details given by Reid et al. (2013). Support values are posterior probabilities; only values ≥ 0.95 (i. e. with strong support) are shown." httpUri="https://zenodo.org/record/285732/files/figure.png" pageId="47" pageNumber="48">Fig. 1</figureCitation>
) although the sister species of the latter is probably the unsampled
<taxonomicName id="4C73CDB665152869D267F987FB9F8863" box="[848,1056,1628,1653]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="rhizophorarum">
<emphasis id="B9076A2765152869D267F987FB9F8863" box="[848,1056,1628,1653]" italics="true" pageId="47" pageNumber="48">C. rhizophorarum</emphasis>
</taxonomicName>
(see Remarks on that species).
<taxonomicName id="4C73CDB665152869D1A0F9A5FE2D888E" box="[151,402,1663,1688]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="tonkiniana">
<emphasis id="B9076A2765152869D1A0F9A5FE2D888E" box="[151,402,1663,1688]" italics="true" pageId="47" pageNumber="48">Cerithidea tonkiniana</emphasis>
</taxonomicName>
and
<taxonomicName id="4C73CDB665152869D0E7F95AFDE9888E" box="[464,598,1663,1688]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="moerchii">
<emphasis id="B9076A2765152869D0E7F95AFDE9888E" box="[464,598,1663,1688]" italics="true" pageId="47" pageNumber="48">C. moerchii</emphasis>
</taxonomicName>
are broadly sympatric from southern
<collectingCountry id="F364F6A565152869D522F95BFBEA888E" box="[1045,1109,1665,1688]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
to
<collectingCountry id="F364F6A565152869D5B6F9A5FB5C888E" box="[1153,1251,1663,1688]" name="Vietnam" pageId="47" pageNumber="48">Vietnam</collectingCountry>
, but are readily distinguished by the much stronger spiral ridges of
<taxonomicName id="4C73CDB665152869D3E1F97FFCE688AB" box="[726,857,1700,1725]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="moerchii">
<emphasis id="B9076A2765152869D3E1F97FFCE688AB" box="[726,857,1700,1725]" italics="true" pageId="47" pageNumber="48">C. moerchii</emphasis>
</taxonomicName>
(
<figureCitation id="1348AAB065152869D25EF97EFC0888AB" box="[873,951,1700,1725]" captionStart="FIGURE 15. A I" captionStartId="50.[151,250,1692,1714]" captionTargetBox="[245,1337,193,1669]" captionTargetId="figure@50.[245,1337,193,1671]" captionTargetPageId="50" captionText="FIGURE 15. A I, Cerithidea rhizophorarum. J AA, C. moerchii. A, C. rhizophorarum A. Adams, 1855, syntype, Cagayan de Oro, Mindanao, Philippines (NHMUK 1996432 / 4). B, no locality (NHMUK 1996432 / 7; not a syntype of C. rhizophorarum). C H, Guijulngan, Negros Oriental, Philippines (USNM 232846; E H views of same specimen). I, Batangas, Luzon, Philippines (USNM 243643). J, Tai Mei Tuk, Tolo Harbour, Hong Kong (NHMUK 20130263). K N, Shiu Hau, Tong Fuk Min Wan, Lantau I., Hong Kong (NHMUK 20130264); L, M views of same specimen). O, P, Yoshida R., Oiehama, Kasaoka City, Hiroshima Pref., Japan (NHMUK 20130265). Q, Kushida R., Matsunase, Mie Pref., Japan (NHMUK 20130266). R, Yonekura, Kiire-nukumi-cho, Kagoshima City, Kagoshima Pref., Japan (NHMUK 20130267). S, Tsuyazaki, Fukuoka Pref., Japan (NHMUK 20130268). T, Hikawa R., Kumamoto Pref., Japan (NHMUK 20130269). U, Cerithium moerchii A. Adams in G. B. Sowerby II, 1855, syntype, no locality (NHMUK 20060321). V, Funaura Bay, Iriomote I., Ryukyu Is, Japan (NHMUK 20130270). W Y, Ura, Tatsugou-cho, Amami Is, Kagoshima Pref., Japan (NHMUK 20130271; X, Y views of same specimen). Z, AA, Tamshui R., Bali, Taiwan (NHMUK 20070387)." figureDoi="http://doi.org/10.5281/zenodo.10846387" httpUri="https://zenodo.org/record/10846387/files/figure.png" pageId="47" pageNumber="48">Fig. 15</figureCitation>
JAA).
</paragraph>
<paragraph id="8BCCB63565152869D1F0F91DFECE893E" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
The larval development has not been explicitly reported, but
<bibRefCitation id="EFE2CBC465152869D2B3F91DFBD088F6" box="[900,1135,1735,1760]" pageId="47" pageNumber="48" refString="Kojima, S., Kamimura, S., Iijima, A., Kimura, T., Kurozumi, T. &amp; Furota, T. (2006) Molecular phylogeny and population structure of tideland snails in the genus Cerithidea around Japan. Marine Biology, 149, 525 - 535. http: // dx. doi. org / 10.1007 / s 00227 - 005 - 0183 - 2" type="journal article">
Kojima
<emphasis id="B9076A2765152869D2D6F913FBA488F6" box="[993,1051,1735,1760]" italics="true" pageId="47" pageNumber="48">et al.</emphasis>
(2006)
</bibRefCitation>
claimed a planktotrophic stage of 1220 days (quoting
<bibRefCitation id="EFE2CBC465152869D323F936FCBC8912" author="Kimura" box="[532,771,1772,1797]" pageId="47" pageNumber="48" refString="Kimura, T., Fujioka, E., Kimura, S. &amp; Aoki, S. (2002) A comparative morphology of eggs and larvae of nine gastropods in tidal flat and marsh of the reed. Venus, 61, 114 - 115. [in Japanese]" type="journal article" year="2002">
Kimura
<emphasis id="B9076A2765152869D34EF937FD0D8913" box="[633,690,1772,1797]" italics="true" pageId="47" pageNumber="48">et al</emphasis>
. 2002
</bibRefCitation>
, who mentioned planktonic larvae in
<taxonomicName id="4C73CDB665152869D5D6F937FAEA8912" box="[1249,1365,1773,1796]" class="Gastropoda" family="Potamididae" genus="Cerithidea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="47" pageNumber="48" phylum="Mollusca" rank="species" species="ornata">
<emphasis id="B9076A2765152869D5D6F937FAEA8912" box="[1249,1365,1773,1796]" italics="true" pageId="47" pageNumber="48">C. ornata</emphasis>
</taxonomicName>
, and unpublished work).
</paragraph>
<paragraph id="8BCCB63565152869D1F0F8EEFCB78966" blockId="47.[151,1436,151,1904]" pageId="47" pageNumber="48">
This species is listed as endangered in
<collectingCountry id="F364F6A565152869D3AEF8EFFD66895A" box="[665,729,1845,1868]" name="Japan" pageId="47" pageNumber="48">Japan</collectingCountry>
(
<bibRefCitation id="EFE2CBC465152869D3DDF8EEFC3B895B" author="Fukuda" box="[746,900,1844,1869]" pageId="47" pageNumber="48" refString="Fukuda, H. (1996) Gastropoda. In: Hanawa, S. &amp; Sakuma, H. (Eds.), Present Status of Estuarine Locales and Benthic Invertebrates Occurring in Estuarine Environment in Japan. WWF Japan Science Report, 3, 11 - 63." type="journal article" year="1996">Fukuda 1996</bibRefCitation>
; Fukuda &amp; Kimura 2012) as a consequence of threats to its habitat and its narrow range in the country.
</paragraph>
</subSubSection>
</treatment>
</document>