treatments-xml/data/78/04/03/780403081765BB32FF9210A505001183.xml

<document id="8BCBB982C6817C1D4F5AFEA16D25A985" ID-CLB-Dataset="285459" ID-DOI="10.24823/EJB.2022.410" ID-GBIF-Dataset="b44efdf4-f377-4fa4-a2ee-00776e4fd78c" ID-ISSN="1747-0036" ID-Zenodo-Dep="10523329" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1704825482796" checkinUser="felipe" docAuthor="Julia, S., Kiew, R. &amp; Ling, C. Y." docDate="2022" docId="780403081765BB32FF9210A505001183" docLanguage="en" docName="EdinbJBot.79.410.1-46.pdf" docOrigin="Edinburgh Journal of Botany 79 (410)" docSource="http://dx.doi.org/10.24823/ejb.2022.410" docStyle="DocumentStyle:73DE8CF451BB5BCBD336E07A8276E690.4:EdinbJBot.2021-.journal_article" docStyleId="73DE8CF451BB5BCBD336E07A8276E690" docStyleName="EdinbJBot.2021-.journal_article" docStyleVersion="4" docTitle="Begonia mentiens S. Julia &amp; Kiew 2022, sp. nov." docType="treatment" docVersion="4" lastPageNumber="25" masterDocId="843D7B701770BB2AFFEB141501571341" masterDocTitle="ADDITIONS TO THE BEGONIA FLORA OF SARAWAK, BORNEO, I: TWELVE NEW SPECIES AND A NEW RECORD" masterLastPageNumber="46" masterPageNumber="1" pageNumber="22" updateTime="1705508941419" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8B4EAAB2D4F58EBDE003ED7EFDB6CCB2">ADDITIONS TO THE BEGONIA FLORA OF SARAWAK, BORNEO, I: TWELVE NEW SPECIES AND A NEW RECORD</mods:title>
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<mods:namePart id="3422996B73F4E1F32FB10777E88B29FD">Julia, S.</mods:namePart>
<mods:affiliation id="4FCC2E32380700D17E642F99839E51D1">Sarawak Herbarium, Forest Department Sarawak, KM 10 Jalan Datuk Amar Kalong Ningkan, 93250 Kuching, Sarawak, Malaysia. &amp; Author for correspondence. E-mail: juliasang 12 @ gmail. com.</mods:affiliation>
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<mods:namePart id="7E4E46651478F7F51016C1FA9DCF3B59">Kiew, R.</mods:namePart>
<mods:affiliation id="74F7FB66C50EEE716493CADF74949477">Forest Research Institute Malaysia, 52109 Kepong, Selangor, Malaysia.</mods:affiliation>
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<mods:namePart id="82055DB1AACEAD5F17037533A6B4EEBA">Ling, C. Y.</mods:namePart>
<mods:affiliation id="257CA0BCE765DC44F823B9991E5638B5">Sarawak Herbarium, Forest Department Sarawak, KM 10 Jalan Datuk Amar Kalong Ningkan, 93250 Kuching, Sarawak, Malaysia.</mods:affiliation>
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<mods:title id="6BB629F1F390855A8E1DE41D1707AD78">Edinburgh Journal of Botany</mods:title>
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<mods:date id="2DD688C32C622E44AEF12F8AEE138C6D">2022</mods:date>
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<subSubSection id="B8B7E1951765BB3FFF9210A5033C1788" box="[121,619,1200,1225]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="F012B21E1765BB3FFF9210A5033C1788" blockId="21.[121,619,1200,1225]" box="[121,619,1200,1225]" pageId="21" pageNumber="22">
<heading id="AB5A05721765BB3FFF9210A5033C1788" box="[121,619,1200,1225]" fontSize="36" level="2" pageId="21" pageNumber="22" reason="2">
<emphasis id="C2D96E0C1765BB3FFF9210A501D91788" bold="true" box="[121,142,1200,1225]" pageId="21" pageNumber="22">9.</emphasis>
<taxonomicName id="37ADC99D1765BB3FFF7E10A5035D1788" ID-CoL="B8QXV" authority="S.Julia &amp; Kiew" authorityName="S. Julia &amp; Kiew" authorityYear="2022" box="[149,522,1200,1225]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="mentiens" status="sp. nov.">
<emphasis id="C2D96E0C1765BB3FFF7E10A500351788" bold="true" box="[149,354,1200,1225]" pageId="21" pageNumber="22">Begonia mentiens</emphasis>
S.Julia &amp; Kiew
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, 
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<taxonomicNameLabel id="D9EAD3771765BB3FFDFF10A5033C1788" box="[532,619,1200,1225]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="F012B21E1765BB3FFF7C10C00039157F" blockId="21.[151,1160,1237,1673]" pageId="21" pageNumber="22">
Similar to 
<taxonomicName id="37ADC99D1765BB3FFEE110C0033317AF" authority="S. Julia" authorityName="S. Julia" box="[266,612,1237,1262]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="jamilahanuiana">
<emphasis id="C2D96E0C1765BB3FFEE110C0034717AF" box="[266,528,1237,1262]" italics="true" pageId="21" pageNumber="22">Begonia jamilahanuiana</emphasis>
S.Julia
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in its habit and leaf characters, but 
<taxonomicName id="37ADC99D1765BB3FFC1C10C0052F17AF" authorityName="S. Julia &amp; Kiew" authorityYear="2022" box="[1015,1144,1237,1262]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="21" pageNumber="22" phylum="Tracheophyta" rank="species" species="mentiens">
<emphasis id="C2D96E0C1765BB3FFC1C10C0052F17AF" box="[1015,1144,1237,1262]" italics="true" pageId="21" pageNumber="22">B. mentiens</emphasis>
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can be distinguished by its wider lamina, 
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(versus 
<quantity id="37551FFB1765BB3FFCDF10EF02F71652" box="[820,928,1274,1299]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="21" pageNumber="22" unit="cm" value="3.75" valueMax="5.0" valueMin="2.5">2.5–5 cm</quantity>
wide); inflorescence a compound cyme, 
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long, produced from leaf axils from base to stem apex (versus paniculate inflorescence 
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long produced from upper leaf axils); bracts narrowly ovate to linear, c.3 × 
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, and bracteoles c.2 × 
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, margin entire (versus bracts ovate, 6–7 × 
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, bractoles 2–6 × 
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, margin studded with glandular knobs); smaller ovary, 9–10 × 
<quantity id="37551FFB1765BB3FFE0111A0033F168F" box="[490,616,1461,1486]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="21" pageNumber="22" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
, with equal wings (versus ovary 17–20 × 
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, with unequal wings); and capsule wider than long, 8–10 × 
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, with longer pedicel, 
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long (versus longer capsule, 11–17 × 
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, with pedicel 
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).
</paragraph>
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<paragraph id="F012B21E1765BB3FFE9E1230030815C8" blockId="21.[151,1160,1237,1673]" pageId="21" pageNumber="22">
<materialsCitation id="40C5B8431765BB3FFE9E1230001315C8" collectingDate="2016-05-25" collectionCode="SAR" collectorName="Ling &amp; Dino" country="Malaysia" county="Sungai Igan" latitude="1.7369444" location="Borneo" longLatPrecision="21" longitude="112.099724" municipality="Lanjak Entimau Wildlife Sanctuary" pageId="21" pageNumber="22" specimenCode="SFC 5966" specimenCount="1" typeStatus="holotype">
– Type: 
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, 
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, Sarawak, 
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, 
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, 
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, 
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, 
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, 
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<collectingDate id="94576D361765BB3FFCE7125F022A1522" box="[780,893,1610,1635]" pageId="21" pageNumber="22" value="2016-05-25">25 v 2016</collectingDate>
</date>
, 
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<collectorName id="5D58D7C81765BB3FFC6D125F02E21522" box="[902,949,1610,1635]" pageId="21" pageNumber="22">Ling</collectorName>
&amp; 
<collectorName id="5D58D7C81765BB3FFC39125F05531522" box="[978,1028,1610,1635]" pageId="21" pageNumber="22">Dino</collectorName>
</emphasis>
<specimenCode id="A00B1A651765BB3FFBE0125F052F1522" box="[1035,1144,1610,1635]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="21" pageNumber="22">SFC 5966</specimenCode>
(
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<collectionCode id="96BC2ADB1765BB3FFEE31265006A15C8" box="[264,317,1648,1673]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="21" pageNumber="22" type="Herbarium">SAR</collectionCode>
!,
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<materialsCitation id="40C5B8431765BB3FFEA11265030015C8" box="[330,599,1648,1673]" collectingDate="2016-05-25" collectionCode="KEP" collectorName="Ling &amp; Dino" country="Malaysia" county="Sungai Igan" latitude="1.7369444" location="Type" longLatPrecision="21" longitude="112.099724" municipality="Lanjak Entimau Wildlife Sanctuary" pageId="21" pageNumber="22" specimenCode="SFC 5966" specimenCount="1" typeStatus="isotype">
<typeStatus id="2F160CBC1765BB3FFEA1126500CB15C8" box="[330,412,1648,1673]" pageId="21" pageNumber="22" type="isotype">isotype</typeStatus>
<collectionCode id="96BC2ADB1765BB3FFE491265008D15C8" box="[418,474,1648,1673]" collectionName="KEP" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="21" pageNumber="22" type="Herbarium">KEP</collectionCode>
!). 
<figureCitation id="6896AE9B1765BB3FFE061265030015C8" box="[493,599,1648,1673]" captionStart="Figure 10" captionStartId="22.[121,185,1514,1536]" captionTargetBox="[121,1164,265,1493]" captionTargetId="figure-7@22.[118,1167,262,1495]" captionTargetPageId="22" captionText="Figure 10. Begonia mentiens S.Julia &amp; Kiew, sp. nov. A, Habit, showing spotted leaves; B, habit, showing individual with plain leaves; C, stipule; D, inflorescence on lower part of stem; E, male inflorescence; F, male flower; G and H, female flower; I, old capsule; J, cross-section of female flower. All photographs taken by C. Y. Ling: A, C and G–J, Ling &amp; Dino SFC 5918; B and E–G, Ling &amp; Dino SFC 5949; D, Ling &amp; Dino SFC 5966 (the type specimen)." figureDoi="http://doi.org/10.5281/zenodo.10523371" httpUri="https://zenodo.org/record/10523371/files/figure.png" pageId="21" pageNumber="22">
<emphasis id="C2D96E0C1765BB3FFE061265030015C8" bold="true" box="[493,599,1648,1673]" pageId="21" pageNumber="22">Figure 10</emphasis>
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<emphasis id="C2D96E0C1765BB3FFDB31265030815C8" bold="true" box="[600,607,1648,1673]" pageId="21" pageNumber="22">.</emphasis>
</paragraph>
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<emphasis id="C2D96E0C1766BB3CFF9211FF01881541" bold="true" box="[121,223,1514,1536]" pageId="22" pageNumber="23">Figure 10.</emphasis>
<taxonomicName id="37ADC99D1766BB3CFF0E11FF037C1541" authority="S.Julia &amp; Kiew" authorityName="S. Julia &amp; Kiew" authorityYear="2022" box="[229,555,1514,1536]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="species" species="mentiens" status="sp. nov.">
<emphasis id="C2D96E0C1766BB3CFF0E11FF00C11541" box="[229,406,1514,1536]" italics="true" pageId="22" pageNumber="23">Begonia mentiens</emphasis>
S.Julia &amp; Kiew
</taxonomicName>
, 
<taxonomicNameLabel id="D9EAD3771766BB3CFDDF11FF03D41541" box="[564,643,1514,1536]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
A, Habit, showing spotted leaves; B, habit, showing individual with plain leaves; C, stipule; D, inflorescence on lower part of stem; E, male inflorescence; F, male flower; G and H, female flower; I, old capsule; J, cross-section of female flower. All photographs taken by C. Y. Ling: A, C and G–J, 
<emphasis id="C2D96E0C1766BB3CFE26125B036A1525" box="[461,573,1614,1636]" italics="true" pageId="22" pageNumber="23">Ling &amp; Dino</emphasis>
SFC 5918; B and E–G, 
<emphasis id="C2D96E0C1766BB3CFCCD125B02C11525" box="[806,918,1614,1636]" italics="true" pageId="22" pageNumber="23">Ling &amp; Dino</emphasis>
SFC 5949; D, 
<emphasis id="C2D96E0C1766BB3CFBCA125B01F115C4" italics="true" pageId="22" pageNumber="23">Ling &amp; Dino</emphasis>
SFC 5966 (the type specimen).
</paragraph>
</caption>
<subSubSection id="B8B7E1951767BB3DFF92151F00131679" pageId="23" pageNumber="24" type="description">
<paragraph id="F012B21E1767BB3DFF92151F00131679" blockId="23.[121,1165,266,1336]" pageId="23" pageNumber="24">
Cane-like herb to 
<quantity id="37551FFB1767BB3DFED6151F00D31262" box="[317,388,266,291]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" pageId="23" pageNumber="24" unit="cm" value="75.0">75 cm</quantity>
tall; whole plant glabrous. 
<emphasis id="C2D96E0C1767BB3DFD58151F03AE1262" box="[691,761,266,291]" italics="true" pageId="23" pageNumber="24">Stems</emphasis>
dark magenta, 
<quantity id="37551FFB1767BB3DFC4C151F05501262" box="[935,1031,266,291]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="23" pageNumber="24" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
thick, internodes 
<quantity id="37551FFB1767BB3DFF13153A00061209" box="[248,337,303,328]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="23" pageNumber="24" unit="cm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 cm</quantity>
, slightly thicker at the nodes. 
<emphasis id="C2D96E0C1767BB3DFD4B153A03AF1209" box="[672,760,303,328]" italics="true" pageId="23" pageNumber="24">Stipules</emphasis>
magenta, ovate, 7–10 × 
<quantity id="37551FFB1767BB3DFBE6153A05381209" box="[1037,1135,303,328]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="23" pageNumber="24" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
, margin entire, keeled, apex setose, seta 
<quantity id="37551FFB1767BB3DFDD0154103CC122C" box="[571,667,340,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="23" pageNumber="24" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, caducous. 
<emphasis id="C2D96E0C1767BB3DFCB0154102FF122C" box="[859,936,340,365]" italics="true" pageId="23" pageNumber="24">Leaves</emphasis>
alternate, distant, oblique, asymmetrical, held more or less horizontally, sometimes held at a 45° angle; petioles magenta, 
<quantity id="37551FFB1767BB3DFEA2158A009412F9" box="[329,451,415,440]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" metricValueMax="10.5" metricValueMin="6.0" pageId="23" pageNumber="24" unit="cm" value="8.25" valueMax="10.5" valueMin="6.0">6–10.5 cm</quantity>
long, grooved above; lamina glabrous with silvery elongated spots in between veins, sometimes with silvery spots along the margin and at the leaf apex or sometimes plain green above, pale green beneath, juvenile plants usually with pinkish spots, in life succulent, matt, ovate, 10–13 × 
<quantity id="37551FFB1767BB3DFD9A161A03881169" box="[625,735,527,552]" metricMagnitude="-2" metricUnit="m" metricValue="6.75" metricValueMax="9.0" metricValueMin="4.5" pageId="23" pageNumber="24" unit="cm" value="6.75" valueMax="9.0" valueMin="4.5">4.5–9 cm</quantity>
, broad side 
<quantity id="37551FFB1767BB3DFC8C161A02E91169" box="[871,958,527,552]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="23" pageNumber="24" unit="cm" value="6.0" valueMax="8.0" valueMin="4.0">4–8 cm</quantity>
wide, base shallowly cordate, basal lobes 
<quantity id="37551FFB1767BB3DFE3816210368110C" box="[467,575,564,589]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="8.5" metricValueMin="3.0" pageId="23" pageNumber="24" unit="cm" value="5.75" valueMax="8.5" valueMin="3.0">3–8.5 cm</quantity>
long, margin almost entire, apex acuminate to caudate, acumen 
<quantity id="37551FFB1767BB3DFEA9164F00F91132" box="[322,430,602,627]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="23" pageNumber="24" unit="cm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 cm</quantity>
long; venation palmate-pinnate, above veins green except near base, magenta beneath, slightly impressed above except near leaf base, prominent beneath, 6–8 veins radiating from the base of leaf, 7 or 8 along the midrib on the broad side, 3–5 veins in basal lobe. 
<emphasis id="C2D96E0C1767BB3DFEB316DF00AF11A2" box="[344,504,714,739]" italics="true" pageId="23" pageNumber="24">Inflorescences</emphasis>
protogynous, in the upper leaf axils, compound cyme, 
<quantity id="37551FFB1767BB3DFF9116FA01861049" box="[122,209,751,776]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="23" pageNumber="24" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 cm</quantity>
long, up to 
<specimenCount id="E6AB79971767BB3DFEBE16FA00E01049" box="[341,439,751,776]" count="7" pageId="23" pageNumber="24" type="female">7 female</specimenCount>
inflorescences in the lower part of the leafless stem; male inflorescence towards the apex; bracts reddish, narrowly ovate to linear, c.3 × 
<quantity id="37551FFB1767BB3DFC0917010572106C" box="[994,1061,788,813]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="mm" value="1.0">1 mm</quantity>
, margin entire, apex setose, seta c. 
<quantity id="37551FFB1767BB3DFE43172F00B21012" box="[424,485,826,851]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="mm" value="1.0">1 mm</quantity>
long, caducous; bracteoles similar to bracts but smaller, c.2 × 
<quantity id="37551FFB1767BB3DFF5C174A01AD1039" box="[183,250,863,888]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="23" pageNumber="24" unit="mm" value="1.0">1 mm</quantity>
, caducous. 
<emphasis id="C2D96E0C1767BB3DFE6B174A035A1039" box="[384,525,863,888]" italics="true" pageId="23" pageNumber="24">Male flowers</emphasis>
: pedicel pink, 
<quantity id="37551FFB1767BB3DFD47174A024C1039" box="[684,795,863,888]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="23" pageNumber="24" unit="mm" value="8.5" valueMax="10.0" valueMin="7.0">7–10 mm</quantity>
long; tepals 2, white or pale pink tinged reddish towards base, ovate, 6–7.5 × 
<quantity id="37551FFB1767BB3DFD491791024E10DC" box="[674,793,900,925]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" pageId="23" pageNumber="24" unit="mm" value="5.25" valueMax="6.5" valueMin="4.0">4–6.5 mm</quantity>
, margin entire, apex rounded; stamens 19–26, cluster subsessile, globose, filaments pale yellow, c. 
<quantity id="37551FFB1767BB3DFC9517BF02A31082" box="[894,1012,938,963]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="23" pageNumber="24" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
long; anthers pale yellow, obovate, c.0.8–1 × 
<quantity id="37551FFB1767BB3DFE3317DA031810A9" box="[472,591,975,1000]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="23" pageNumber="24" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
, apex emarginate. 
<emphasis id="C2D96E0C1767BB3DFCCF17DA029C10A9" box="[804,971,975,1000]" italics="true" pageId="23" pageNumber="24">Female flowers</emphasis>
: pedicel pinkish, 
<quantity id="37551FFB1767BB3DFF9117E101BF174C" box="[122,232,1012,1037]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="23" pageNumber="24" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
long, glabrous; ovary pale green, ovate to triangular in outline, ovary including the wings 9–10 × 
<quantity id="37551FFB1767BB3DFEF1100F00CF1772" box="[282,408,1050,1075]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="23" pageNumber="24" unit="mm" value="14.0" valueMax="15.0" valueMin="13.0">13–15 mm</quantity>
, wings 3, equal, sometimes tinged pink, locules 3, placenta 1 per locule; tepals 5, pale pink, deeper pinkish at base, outer four tepals elliptic, 6–9 × 
<quantity id="37551FFB1767BB3DFBE0102A053A1719" box="[1035,1133,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="23" pageNumber="24" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
, margin entire, apex rounded, inner tepal narrowly elliptic, 6 × 
<quantity id="37551FFB1767BB3DFCCF107102CC173C" box="[804,923,1124,1149]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="23" pageNumber="24" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
; styles 3, pale yellow green, 
<quantity id="37551FFB1767BB3DFF2E109F007217E2" box="[197,293,1162,1187]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="23" pageNumber="24" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
long, deeply divided to base, widely Y-shaped; stigma pale yellow, papillose cells large and pale yellow forming a continuous twisted band. 
<emphasis id="C2D96E0C1767BB3DFCAB10BA02F21789" box="[832,933,1199,1224]" italics="true" pageId="23" pageNumber="24">Capsules</emphasis>
glabrous, 8–10 × 
<quantity id="37551FFB1767BB3DFF9110C101AF17AC" box="[122,248,1236,1261]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="1.6" metricValueMin="1.2" pageId="23" pageNumber="24" unit="mm" value="14.0" valueMax="16.0" valueMin="12.0">12–16 mm</quantity>
, locules 3, wings 3, equal, rounded proximally, truncate distally, 
<quantity id="37551FFB1767BB3DFC2A10C1057617AC" box="[961,1057,1236,1261]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="23" pageNumber="24" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
wide, thinly fibrous, dehiscing between locule and wing; pedicel pendent, 
<quantity id="37551FFB1767BB3DFC8610EF02BE1652" box="[877,1001,1274,1299]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="23" pageNumber="24" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
long, reddish towards the base.
</paragraph>
</subSubSection>
<subSubSection id="B8B7E1951767BB3DFF921146020216D3" pageId="23" pageNumber="24" type="distribution">
<paragraph id="F012B21E1767BB3DFF921146020216D3" blockId="23.[121,1051,1363,1426]" pageId="23" pageNumber="24">
<emphasis id="C2D96E0C1767BB3DFF92114601A0162D" box="[121,247,1363,1388]" italics="true" pageId="23" pageNumber="24">Distribution</emphasis>
. 
<collectingCountry id="88BAF28E1767BB3DFEEE1146003D162D" box="[261,362,1363,1388]" name="Malaysia" pageId="23" pageNumber="24">Malaysia</collectingCountry>
(
<collectingCountry id="88BAF28E1767BB3DFE9D11460088162D" box="[374,479,1363,1388]" name="Malaysia" pageId="23" pageNumber="24">Sarawak</collectingCountry>
). To date, known only from Lanjak Entimau Wildlife Sanctuary (see 
<figureCitation id="6896AE9B1767BB3DFEC2116C00D216D3" box="[297,389,1401,1426]" captionStart="Figure 1" captionStartId="2.[121,185,1535,1557]" captionTargetBox="[122,1161,268,1511]" captionTargetId="figure-7@2.[121,1164,265,1512]" captionTargetPageId="2" captionText="Figure 1. Distribution of Begonia papyraptera and the 12 new species of Begonia recorded from Sarawak and described in this article." figureDoi="http://doi.org/10.5281/zenodo.10523331" httpUri="https://zenodo.org/record/10523331/files/figure.png" pageId="23" pageNumber="24">
<emphasis id="C2D96E0C1767BB3DFEC2116C00D216D3" bold="true" box="[297,389,1401,1426]" pageId="23" pageNumber="24">Figure 1</emphasis>
</figureCitation>
), where the species is widely distributed.
</paragraph>
</subSubSection>
<subSubSection id="B8B7E1951767BB3DFF9211B8036716AA" pageId="23" pageNumber="24" type="biology_ecology">
<paragraph id="F012B21E1767BB3DFF9211B8036716AA" blockId="23.[121,1134,1453,1515]" pageId="23" pageNumber="24">
<emphasis id="C2D96E0C1767BB3DFF9211B8019E1687" box="[121,201,1453,1478]" italics="true" pageId="23" pageNumber="24">Habitat</emphasis>
. Riparian forest along stream bank with large sandstone or mudstone boulders, on alluvial soils, at elevations 
<quantity id="37551FFB1767BB3DFE4D11C7037B16AA" box="[422,556,1490,1515]" metricMagnitude="2" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.4" metricValueMin="1.24" pageId="23" pageNumber="24" unit="m" value="132.0" valueMax="140.0" valueMin="124.0">
124– 
<specimenCount id="E6AB79971767BB3DFE3411C7037B16AA" box="[479,556,1490,1515]" count="140" pageId="23" pageNumber="24" type="generic">140 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="B8B7E1951767BB3DFF92121200D11504" pageId="23" pageNumber="24" type="etymology">
<paragraph id="F012B21E1767BB3DFF92121200D11504" blockId="23.[121,1102,1543,1605]" pageId="23" pageNumber="24">
<emphasis id="C2D96E0C1767BB3DFF92121201BB1561" box="[121,236,1543,1568]" italics="true" pageId="23" pageNumber="24">Etymology</emphasis>
. Latin, 
<emphasis id="C2D96E0C1767BB3DFED6121200F51561" box="[317,418,1543,1568]" italics="true" pageId="23" pageNumber="24">mentiens</emphasis>
(‘imitating’), referring to the sterile plant resembling those of 
<taxonomicName id="37ADC99D1767BB3DFF91123900281504" box="[122,383,1580,1605]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="jamilahanuiana">
<emphasis id="C2D96E0C1767BB3DFF91123900281504" box="[122,383,1580,1605]" italics="true" pageId="23" pageNumber="24">Begonia jamilahanuiana</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="B8B7E1951767BB32FF92124A05D01223" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="F012B21E1767BB32FF92124A05D01223" blockId="23.[121,1121,1631,1686]" lastBlockId="24.[121,1159,265,354]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="C2D96E0C1767BB3DFF92124A00E51534" box="[121,434,1631,1653]" italics="true" pageId="23" pageNumber="24">Additional specimens examined</emphasis>
. 
<materialsCitation id="40C5B8431767BB3DFE55124A027515D7" collectingDate="2016-05-20" collectionCode="KEP, SAR" collectorName="Ling &amp; Dino" country="Malaysia" latitude="1.7055556" location="Sungai Sunpan" longLatPrecision="21" longitude="112.08" municipality="Ulu Mujok" pageId="23" pageNumber="24" specimenCode="SFC 5918" specimenCount="1" stateProvince="Sarawak">
<collectingCountry id="88BAF28E1767BB3DFE55124A03481534" box="[446,543,1631,1653]" name="Malaysia" pageId="23" pageNumber="24">MALAYSIA</collectingCountry>
. 
<emphasis id="C2D96E0C1767BB3DFDC0124A03D51534" bold="true" box="[555,642,1631,1653]" pageId="23" pageNumber="24">
<collectingRegion id="32697CFC1767BB3DFDC0124A03D51534" box="[555,642,1631,1653]" country="Malaysia" name="Sarawak" pageId="23" pageNumber="24">Sarawak</collectingRegion>
</emphasis>
: Julau District – 
<collectingMunicipality id="107628641767BB3DFCC5124A02C21534" box="[814,917,1631,1653]" pageId="23" pageNumber="24">Ulu Mujok</collectingMunicipality>
, 
<location id="F572E4C51767BB3DFC75124A05601534" LSID="urn:lsid:plazi:treatment:780403081765BB32FF9210A505001183:F572E4C51767BB3DFC75124A05601534" box="[926,1079,1631,1653]" country="Malaysia" latitude="1.7055556" longLatPrecision="21" longitude="112.08" municipality="Ulu Mujok" name="Sungai Sunpan" pageId="23" pageNumber="24" stateProvince="Sarawak">Sungai Sunpan</location>
, 
<geoCoordinate id="9599D4D91767BB3DFF911295018915D7" box="[122,222,1664,1686]" degrees="1" direction="north" minutes="42" orientation="latitude" pageId="23" pageNumber="24" precision="15" seconds="20" value="1.7055556">
1 
<emphasis id="C2D96E0C1767BB3DFF6D129501D915D7" box="[134,142,1664,1686]" italics="true" pageId="23" pageNumber="24">°</emphasis>
42′20′′N
</geoCoordinate>
, 
<geoCoordinate id="9599D4D91767BB3DFF0D1295003615D7" box="[230,353,1664,1686]" degrees="112" direction="east" minutes="04" orientation="longitude" pageId="23" pageNumber="24" precision="15" seconds="48" value="112.08">
112 
<emphasis id="C2D96E0C1767BB3DFEE71295004315D7" box="[268,276,1664,1686]" italics="true" pageId="23" pageNumber="24">°</emphasis>
04′48′′E
</geoCoordinate>
, 
<date id="841394DE1767BB3DFE811295008715D7" box="[362,464,1664,1686]" pageId="23" pageNumber="24" value="2016-05-20">
<collectingDate id="94576D361767BB3DFE811295008715D7" box="[362,464,1664,1686]" pageId="23" pageNumber="24" value="2016-05-20">20 v 2016</collectingDate>
</date>
, 
<emphasis id="C2D96E0C1767BB3DFE331295031F15D7" box="[472,584,1664,1686]" italics="true" pageId="23" pageNumber="24">
<collectorName id="5D58D7C81767BB3DFE331295035515D7" box="[472,514,1664,1686]" pageId="23" pageNumber="24">Ling</collectorName>
&amp; 
<collectorName id="5D58D7C81767BB3DFDF71295031F15D7" box="[540,584,1664,1686]" pageId="23" pageNumber="24">Dino</collectorName>
</emphasis>
<specimenCode id="A00B1A651767BB3DFDA5129503E715D7" box="[590,688,1664,1686]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="23" pageNumber="24">SFC 5918</specimenCode>
(
<collectionCode id="96BC2ADB1767BB3DFD50129503B215D7" box="[699,741,1664,1686]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="23" pageNumber="24" type="Herbarium">KEP</collectionCode>
, 
<collectionCode id="96BC2ADB1767BB3DFD051295024A15D7" box="[750,797,1664,1686]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="23" pageNumber="24" type="Herbarium">SAR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="40C5B8431767BB32FCC012950572125E" collectingDate="2016-05-21" collectionCode="KEP, SAR" collectorName="Ling &amp; Dino" country="Malaysia" county="Mujok" lastPageId="24" lastPageNumber="25" latitude="1.681389" location="Sungai Segak" longLatPrecision="21" longitude="112.16111" municipality="Lanjak Entimau Wildlife Sanctuary" pageId="23" pageNumber="24" specimenCode="SFC 5929" specimenCount="1" stateProvince="Sarawak">
<collectingCounty id="1973CA921767BB3DFCC01295023C15D7" box="[811,875,1664,1686]" pageId="23" pageNumber="24">Mujok</collectingCounty>
, 
<collectingMunicipality id="107628641767BB32FC9F12950188125E" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">Lanjak Entimau Wildlife Sanctuary</collectingMunicipality>
, 
<location id="F572E4C51768BB32FF0C151C0024125E" LSID="urn:lsid:plazi:treatment:780403081765BB32FF9210A505001183:F572E4C51768BB32FF0C151C0024125E" box="[231,371,265,287]" country="Malaysia" county="Mujok" latitude="1.681389" longLatPrecision="21" longitude="112.16111" municipality="Lanjak Entimau Wildlife Sanctuary" name="Sungai Segak" pageId="24" pageNumber="25" stateProvince="Sarawak">Sungai Segak</location>
, 
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1 
<emphasis id="C2D96E0C1768BB32FE63151C00C7125E" box="[392,400,265,287]" italics="true" pageId="24" pageNumber="25">°</emphasis>
40′53′′N
</geoCoordinate>
, 
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112 
<emphasis id="C2D96E0C1768BB32FDE4151C0340125E" box="[527,535,265,287]" italics="true" pageId="24" pageNumber="25">°</emphasis>
09′40′′E
</geoCoordinate>
, 
<date id="841394DE1768BB32FD87151C0385125E" box="[620,722,265,287]" pageId="24" pageNumber="25" value="2016-05-21">
<collectingDate id="94576D361768BB32FD87151C0385125E" box="[620,722,265,287]" pageId="24" pageNumber="25" value="2016-05-21">21 v 2016</collectingDate>
</date>
, 
<emphasis id="C2D96E0C1768BB32FD31151C021D125E" box="[730,842,265,287]" italics="true" pageId="24" pageNumber="25">
<collectorName id="5D58D7C81768BB32FD31151C0253125E" box="[730,772,265,287]" pageId="24" pageNumber="25">Ling</collectorName>
&amp; 
<collectorName id="5D58D7C81768BB32FCF5151C021D125E" box="[798,842,265,287]" pageId="24" pageNumber="25">Dino</collectorName>
</emphasis>
<specimenCode id="A00B1A651768BB32FCBB151C02E5125E" box="[848,946,265,287]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="24" pageNumber="25">SFC 5929</specimenCode>
(
<collectionCode id="96BC2ADB1768BB32FC55151C02BF125E" box="[958,1000,265,287]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="24" pageNumber="25" type="Herbarium">KEP</collectionCode>
, 
<collectionCode id="96BC2ADB1768BB32FC1A151C0577125E" box="[1009,1056,265,287]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="24" pageNumber="25" type="Herbarium">SAR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="40C5B8431768BB32FBC5151C01FA1223" collectingDate="2016-05-23" collectionCode="KEP, SAR" collectorName="Ling &amp; Dino" country="Malaysia" latitude="1.69" location="Sungai Mujok" longLatPrecision="21" longitude="112.11694" municipality="Sungai Nyamuk" pageId="24" pageNumber="25" specimenCode="SFC 5949" specimenCount="1" stateProvince="Sarawak">
<collectingMunicipality id="107628641768BB32FBC5151C01981200" pageId="24" pageNumber="25">Sungai Nyamuk</collectingMunicipality>
, tributary of 
<location id="F572E4C51768BB32FEA7153E00801200" LSID="urn:lsid:plazi:treatment:780403081765BB32FF9210A505001183:F572E4C51768BB32FEA7153E00801200" box="[332,471,299,321]" country="Malaysia" latitude="1.69" longLatPrecision="21" longitude="112.11694" municipality="Sungai Nyamuk" name="Sungai Mujok" pageId="24" pageNumber="25" stateProvince="Sarawak">Sungai Mujok</location>
, 
<geoCoordinate id="9599D4D91768BB32FE0B153E03121200" box="[480,581,299,321]" degrees="1" direction="north" minutes="41" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="24" value="1.69">
1 
<emphasis id="C2D96E0C1768BB32FE07153E00A31200" box="[492,500,299,321]" italics="true" pageId="24" pageNumber="25">°</emphasis>
41′24′′N
</geoCoordinate>
, 
<geoCoordinate id="9599D4D91768BB32FDA6153E03901200" box="[589,711,299,321]" degrees="112" direction="east" minutes="07" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="01" value="112.11694">
112 
<emphasis id="C2D96E0C1768BB32FD98153E032C1200" box="[627,635,299,321]" italics="true" pageId="24" pageNumber="25">°</emphasis>
07′01′′E
</geoCoordinate>
, 
<date id="841394DE1768BB32FD3B153E02611200" box="[720,822,299,321]" pageId="24" pageNumber="25" value="2016-05-23">
<collectingDate id="94576D361768BB32FD3B153E02611200" box="[720,822,299,321]" pageId="24" pageNumber="25" value="2016-05-23">23 v 2016</collectingDate>
</date>
, 
<emphasis id="C2D96E0C1768BB32FCD4153E02F91200" box="[831,942,299,321]" italics="true" pageId="24" pageNumber="25">
<collectorName id="5D58D7C81768BB32FCD4153E023E1200" box="[831,873,299,321]" pageId="24" pageNumber="25">Ling</collectorName>
&amp; 
<collectorName id="5D58D7C81768BB32FC69153E02F91200" box="[898,942,299,321]" pageId="24" pageNumber="25">Dino</collectorName>
</emphasis>
<specimenCode id="A00B1A651768BB32FC5E153E05401200" box="[949,1047,299,321]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="24" pageNumber="25">SFC 5949</specimenCode>
(
<collectionCode id="96BC2ADB1768BB32FBC9153E051B1200" box="[1058,1100,299,321]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="24" pageNumber="25" type="Herbarium">KEP</collectionCode>
, 
<collectionCode id="96BC2ADB1768BB32FF91155901FE1223" box="[122,169,332,354]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="24" pageNumber="25" type="Herbarium">SAR</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="40C5B8431768BB32FF5C155905D41223" box="[183,1155,332,354]" collectingDate="2016-05-24" collectionCode="KEP, SAR" collectorName="Ling &amp; Dino" country="Malaysia" latitude="1.7344445" location="Sungai Sugai" longLatPrecision="21" longitude="112.099724" municipality="Sungai Mujok" pageId="24" pageNumber="25" specimenCode="SFC 5958" specimenCount="1" stateProvince="Sarawak">
<collectingMunicipality id="107628641768BB32FF5C155900151223" box="[183,322,332,354]" pageId="24" pageNumber="25">Sungai Mujok</collectingMunicipality>
, 
<location id="F572E4C51768BB32FEA1155900871223" LSID="urn:lsid:plazi:treatment:780403081765BB32FF9210A505001183:F572E4C51768BB32FEA1155900871223" box="[330,464,332,354]" country="Malaysia" latitude="1.7344445" longLatPrecision="21" longitude="112.099724" municipality="Sungai Mujok" name="Sungai Sugai" pageId="24" pageNumber="25" stateProvince="Sarawak">Sungai Sugai</location>
, 
<geoCoordinate id="9599D4D91768BB32FE321559036A1223" box="[473,573,332,354]" degrees="1" direction="north" minutes="44" orientation="latitude" pageId="24" pageNumber="25" precision="15" seconds="04" value="1.7344445">
1 
<emphasis id="C2D96E0C1768BB32FE0E155900BA1223" box="[485,493,332,354]" italics="true" pageId="24" pageNumber="25">°</emphasis>
44′04′′N
</geoCoordinate>
, 
<geoCoordinate id="9599D4D91768BB32FDAE155903971223" box="[581,704,332,354]" degrees="112" direction="east" minutes="05" orientation="longitude" pageId="24" pageNumber="25" precision="15" seconds="59" value="112.099724">
112 
<emphasis id="C2D96E0C1768BB32FD87155903231223" box="[620,628,332,354]" italics="true" pageId="24" pageNumber="25">°</emphasis>
05′59′′E
</geoCoordinate>
, 
<date id="841394DE1768BB32FD22155902781223" box="[713,815,332,354]" pageId="24" pageNumber="25" value="2016-05-24">
<collectingDate id="94576D361768BB32FD22155902781223" box="[713,815,332,354]" pageId="24" pageNumber="25" value="2016-05-24">24 v 2016</collectingDate>
</date>
, 
<emphasis id="C2D96E0C1768BB32FCDC155902F01223" box="[823,935,332,354]" italics="true" pageId="24" pageNumber="25">
<collectorName id="5D58D7C81768BB32FCDC155902361223" box="[823,865,332,354]" pageId="24" pageNumber="25">Ling</collectorName>
&amp; 
<collectorName id="5D58D7C81768BB32FC90155902F01223" box="[891,935,332,354]" pageId="24" pageNumber="25">Dino</collectorName>
</emphasis>
<specimenCode id="A00B1A651768BB32FC46155905581223" box="[941,1039,332,354]" collectionCode="SFC" country="China" httpUri="http://grbio.org/cool/myie-bfyd" name="Laboratory of Fishes" pageId="24" pageNumber="25">SFC 5958</specimenCode>
(
<collectionCode id="96BC2ADB1768BB32FBF0155905121223" box="[1051,1093,332,354]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="24" pageNumber="25" type="Herbarium">KEP</collectionCode>
, 
<collectionCode id="96BC2ADB1768BB32FBA5155905291223" box="[1102,1150,332,354]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12931" lsid="urn:lsid:biocol.org:col:12931" name="Department of Forestry" pageId="24" pageNumber="25" type="Herbarium">SAR</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="B8B7E1951768BB32FF92156A05001183" pageId="24" pageNumber="25" type="discussion">
<paragraph id="F012B21E1768BB32FF92156A02691113" blockId="24.[121,1149,383,706]" pageId="24" pageNumber="25">
<taxonomicName id="37ADC99D1768BB32FF92156A006912D9" authorityName="S. Julia &amp; Kiew" authorityYear="2022" box="[121,318,383,408]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="mentiens">
<emphasis id="C2D96E0C1768BB32FF92156A006912D9" box="[121,318,383,408]" italics="true" pageId="24" pageNumber="25">Begonia mentiens</emphasis>
</taxonomicName>
is rather common and occurs in a few river systems in Ulu Mujok. In the field, individual plants are scattered (not growing in clumps or close together). In comparison, plants of 
<taxonomicName id="37ADC99D1768BB32FE9D15DC032012A3" box="[374,631,457,482]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="jamilahanuana">
<emphasis id="C2D96E0C1768BB32FE9D15DC032012A3" box="[374,631,457,482]" italics="true" pageId="24" pageNumber="25">Begonia jamilahanuana</emphasis>
</taxonomicName>
usually grow close together to form a large clump. The irregular spots on the upper leaf surface of 
<taxonomicName id="37ADC99D1768BB32FD0315FA02FA1149" authorityName="S. Julia &amp; Kiew" authorityYear="2022" box="[744,941,495,520]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="mentiens">
<emphasis id="C2D96E0C1768BB32FD0315FA02FA1149" box="[744,941,495,520]" italics="true" pageId="24" pageNumber="25">Begonia mentiens</emphasis>
</taxonomicName>
are pink in the juvenile stage but turn white with age or sometimes completely disappear, whereas the spots on 
<taxonomicName id="37ADC99D1768BB32FF08162C00F71113" box="[227,416,569,594]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="jamilahanuana">
<emphasis id="C2D96E0C1768BB32FF08162C00F71113" box="[227,416,569,594]" italics="true" pageId="24" pageNumber="25">B. jamilahanuana</emphasis>
</taxonomicName>
leaves are persistent in adult plants.
</paragraph>
<paragraph id="F012B21E1768BB32FF72164A05001183" blockId="24.[121,1149,383,706]" pageId="24" pageNumber="25">
<taxonomicName id="37ADC99D1768BB32FF72164A00CE1139" box="[153,409,607,632]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="jamilahanuana">
<emphasis id="C2D96E0C1768BB32FF72164A00CE1139" box="[153,409,607,632]" italics="true" pageId="24" pageNumber="25">Begonia jamilahanuana</emphasis>
</taxonomicName>
and 
<taxonomicName id="37ADC99D1768BB32FE3A164A03041139" authorityName="S. Julia &amp; Kiew" authorityYear="2022" box="[465,595,607,632]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="mentiens">
<emphasis id="C2D96E0C1768BB32FE3A164A03041139" box="[465,595,607,632]" italics="true" pageId="24" pageNumber="25">B. mentiens</emphasis>
</taxonomicName>
seem to occupy different microhabitats. 
<taxonomicName id="37ADC99D1768BB32FBCF164A004D11DC" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="jamilahanuana">
<emphasis id="C2D96E0C1768BB32FBCF164A004D11DC" italics="true" pageId="24" pageNumber="25">Begonia jamilahanuana</emphasis>
</taxonomicName>
is usually found along shade ridges of hill mixed dipterocarp forest with a thick layer of leaf litter, in contrast to 
<taxonomicName id="37ADC99D1768BB32FDF716BC03CA1183" authorityName="S. Julia &amp; Kiew" authorityYear="2022" box="[540,669,681,706]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="mentiens">
<emphasis id="C2D96E0C1768BB32FDF716BC03CA1183" box="[540,669,681,706]" italics="true" pageId="24" pageNumber="25">B. mentiens</emphasis>
</taxonomicName>
, which usually grows by stream banks.
</paragraph>
</subSubSection>
</treatment>
</document>