237 lines
25 KiB
XML
237 lines
25 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.5061052" ID-GBIF-Dataset="c08dd8ae-6f1e-4522-a4a2-ced89fdd3530" ID-GBIF-Taxon="183237254" ID-ISSN="1879-0445" ID-Zenodo-Dep="5061052" LSID="urn:lsid:zoobank.org:act:A917002F-D2AA-4F13-A5E6-ABA9B5F57FC2" checkinTime="1625232919875" checkinUser="diego" docAuthor="Qvarnström, Martin, Fikáček, Martin, Vikberg Wernström, Joel, Huld, Sigrid, Beutel, Rolf G., Arriaga-Varela, Emmanuel, Ahlberg, Per E. & Niedźwiedzki, Grzegorz" docDate="2021" docId="195169142275FF80FCB644156950FB6A" docLanguage="en" docName="CurrBiol.31.1-8.pdf" docOrigin="Current Biology 31" docSource="http://dx.doi.org/10.1016/j.cub.2021.05.015" docTitle="Triamyxidae Qvarnström & Fikáček & Vikberg Wernström & Huld & Beutel & Arriaga-Varela & Ahlberg & Niedźwiedzki 2021, fam. nov." docType="treatment" docUuid="A917002F-D2AA-4F13-A5E6-ABA9B5F57FC2" docUuidSource="ZooBank" docVersion="6" lastPageId="5" lastPageNumber="3" masterDocId="E568116C2276FF85FF9D404A6854FFFB" masterDocTitle="Exceptionally preserved beetles in a Triassic coprolite of putative dinosauriform origin" masterLastPageNumber="8" masterPageNumber="1" pageId="3" pageNumber="3" updateTime="1643022678139" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Exceptionally preserved beetles in a Triassic coprolite of putative dinosauriform origin</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Qvarnström, Martin</mods:namePart>
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<mods:affiliation>Department of Organismal Biology, Evolutionary Biology Centre, Uppsala University, Norbyv € agen 18 A, 752 36 Uppsala, Sweden</mods:affiliation>
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<mods:nameIdentifier type="email">martin.qvarnstrom@ebc.uu.se</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fikáček, Martin</mods:namePart>
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<mods:affiliation>Department of Biological Sciences, National Sun Yat-sen University, no. 70, Lienhai Road, Kaohsiung 80424, Taiwan & Department of Entomology, National Museum, Praha 9, Czech Republic & Department of Zoology, Faculty of Science, Charles University, Praha 2, Czech Republic</mods:affiliation>
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<mods:nameIdentifier type="email">mfikacek@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vikberg Wernström, Joel</mods:namePart>
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<mods:affiliation>Department of Organismal Biology, Evolutionary Biology Centre, Uppsala University, Norbyv € agen 18 A, 752 36 Uppsala, Sweden</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huld, Sigrid</mods:namePart>
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<mods:affiliation>Geocentrum, Uppsala University, Villav € agen 16, 752 36 Uppsala, Sweden</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Beutel, Rolf G.</mods:namePart>
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<mods:affiliation>Institut fu ¨ r Zoologie und Evolutionsforschung, Friedrich-Schiller-Universit € at Jena, Jena, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Arriaga-Varela, Emmanuel</mods:namePart>
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<mods:affiliation>Centro de Estudios en Zoologı´a, CUCBA, Universidad de Guadalajara, Zapopan, Mexico</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ahlberg, Per E.</mods:namePart>
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<mods:affiliation>Department of Organismal Biology, Evolutionary Biology Centre, Uppsala University, Norbyv € agen 18 A, 752 36 Uppsala, Sweden</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Niedźwiedzki, Grzegorz</mods:namePart>
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<mods:affiliation>Department of Organismal Biology, Evolutionary Biology Centre, Uppsala University, Norbyv € agen 18 A, 752 36 Uppsala, Sweden</mods:affiliation>
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<mods:nameIdentifier type="email">grzegorz.niedzwiedzki@ebc.uu.se</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Current Biology</mods:title>
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||
</mods:titleInfo>
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||
<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-06-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>31</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>8</mods:end>
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||
</mods:extent>
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||
</mods:part>
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||
</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://dx.doi.org/10.1016/j.cub.2021.05.015</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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||
<mods:identifier type="DOI">10.1016/j.cub.2021.05.015</mods:identifier>
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||
<mods:identifier type="GBIF-Dataset">c08dd8ae-6f1e-4522-a4a2-ced89fdd3530</mods:identifier>
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||
<mods:identifier type="ISSN">1879-0445</mods:identifier>
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||
<mods:identifier type="Zenodo-Dep">5061134</mods:identifier>
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<mods:identifier type="ZooBank">6FB8E4B4-51C6-493C-A1E6-1829B9258993</mods:identifier>
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||
</mods:mods>
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||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5061052" ID-GBIF-Taxon="183237254" ID-Zenodo-Dep="5061052" LSID="urn:lsid:zoobank.org:act:A917002F-D2AA-4F13-A5E6-ABA9B5F57FC2" httpUri="http://treatment.plazi.org/id/195169142275FF80FCB644156950FB6A" lastPageId="5" lastPageNumber="3" pageId="3" pageNumber="3">
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<subSubSection box="[811,1139,1119,1140]" pageId="3" pageNumber="3" type="nomenclature">
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<paragraph blockId="3.[811,1450,1119,1987]" box="[811,1139,1119,1140]" pageId="3" pageNumber="3">
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<heading bold="true" box="[811,1139,1119,1140]" fontSize="8" level="2" pageId="3" pageNumber="3" reason="6">
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<emphasis bold="true" box="[811,1139,1119,1140]" pageId="3" pageNumber="3">
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Family
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<taxonomicName authority="Qvarnström & Fikáček & Vikberg Wernström & Huld & Beutel & Arriaga-Varela & Ahlberg & Niedźwiedzki, 2021" authorityName="Qvarnström & Fikáček & Vikberg Wernström & Huld & Beutel & Arriaga-Varela & Ahlberg & Niedźwiedzki" authorityYear="2021" box="[891,1028,1119,1140]" class="Insecta" family="Triamyxidae" higherTaxonomySource="Manual Input" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family" status="fam. nov." subOrder="Myxophaga">Triamyxidae</taxonomicName>
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<taxonomicNameLabel box="[1035,1139,1119,1140]" pageId="3" pageNumber="3" rank="family">fam. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="3" type="type_taxon">
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<paragraph blockId="3.[811,1450,1119,1987]" box="[811,940,1148,1169]" pageId="3" pageNumber="3">
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<emphasis bold="true" box="[811,940,1148,1169]" italics="true" pageId="3" pageNumber="3">
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<typeStatus box="[811,865,1148,1169]" pageId="3" pageNumber="3">Type</typeStatus>
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genus
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</emphasis>
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</paragraph>
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<paragraph blockId="3.[811,1450,1119,1987]" box="[811,1002,1177,1198]" pageId="3" pageNumber="3">
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<taxonomicName authority="Qvarnström & Fikáček & Vikberg Wernström & Huld & Beutel & Arriaga-Varela & Ahlberg & Niedźwiedzki, 2021" authorityName="Qvarnström & Fikáček & Vikberg Wernström & Huld & Beutel & Arriaga-Varela & Ahlberg & Niedźwiedzki" authorityYear="2021" box="[811,901,1177,1198]" class="Insecta" family="Hydroscaphidae" genus="Triamyxa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis box="[811,901,1177,1198]" italics="true" pageId="3" pageNumber="3">Triamyxa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[907,1002,1177,1198]" pageId="3" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="5" pageId="3" pageNumber="3" type="diagnosis">
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<paragraph blockId="3.[811,1450,1119,1987]" box="[811,1052,1206,1227]" pageId="3" pageNumber="3">
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<emphasis bold="true" box="[811,1052,1206,1227]" italics="true" pageId="3" pageNumber="3">Differential diagnosis</emphasis>
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</paragraph>
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<paragraph blockId="3.[811,1450,1119,1987]" lastBlockId="5.[142,780,1119,1169]" lastPageId="5" lastPageNumber="5" pageId="3" pageNumber="3">
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Characters of the new family are put in contrast to [
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<emphasis bold="true" box="[1362,1381,1237,1256]" pageId="3" pageNumber="3">x</emphasis>
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] other myxophagan families: dorsal body surface without tubercles or ridges (
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<emphasis bold="true" box="[919,938,1296,1315]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Hinton" authorityYear="1936" box="[937,1062,1294,1315]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
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); eyes strongly protruding (
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<emphasis bold="true" box="[1347,1366,1296,1315]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Erichson" authorityYear="1845" class="Insecta" family="Sphaeriusidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Sphaeriusidae</taxonomicName>
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, most
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<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[984,1158,1323,1344]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
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); head anterior of eyes prolonged and distinctly narrowing (
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<emphasis bold="true" box="[1254,1273,1354,1373]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[1273,1446,1353,1374]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
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,
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<taxonomicName authorityName="Erichson" authorityYear="1845" box="[811,962,1382,1403]" class="Insecta" family="Sphaeriusidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Sphaeriusidae</taxonomicName>
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); antennal bases exposed dorsally between eyes (
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<emphasis bold="true" box="[876,895,1412,1431]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[895,1068,1411,1432]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
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,
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<taxonomicName authorityName="Hinton" authorityYear="1936" box="[1082,1207,1411,1432]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
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); antennal scapus and pedicel distinctly separated from each other (
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<emphasis bold="true" box="[1307,1326,1442,1461]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Steffan" authorityYear="1964" class="Insecta" family="Torridincolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Torridincolidae</taxonomicName>
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,
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<taxonomicName authorityName="Hinton" authorityYear="1936" box="[871,996,1470,1491]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
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), exposed in dorsal view (
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<emphasis bold="true" box="[1303,1322,1471,1490]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Hinton" authorityYear="1936" box="[1322,1446,1470,1491]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
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, modern
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<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[899,1073,1499,1520]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
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); terminal maxillary palpomere long, longer than penultimate (
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<emphasis bold="true" box="[1083,1102,1529,1548]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[1102,1275,1528,1549]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
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,
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<taxonomicName authorityName="Erichson" authorityYear="1845" box="[1289,1440,1528,1549]" class="Insecta" family="Sphaeriusidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Sphaeriusidae</taxonomicName>
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); mentum large, narrowing anteriad (
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<emphasis bold="true" box="[1198,1217,1559,1578]" pageId="3" pageNumber="3">x</emphasis>
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modern
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<taxonomicName authorityName="Le Conte" authorityYear="1874" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
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); prosternum long, with very short prosternal process (
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<emphasis bold="true" box="[817,836,1617,1636]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[836,1009,1616,1637]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
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,
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<taxonomicName authorityName="Hinton" authorityYear="1936" box="[1027,1147,1616,1637]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
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); propleuron wide, without posterior projection (
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<emphasis bold="true" box="[1049,1068,1646,1665]" pageId="3" pageNumber="3">x</emphasis>
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all modern families); mesoventrite only slightly shorter than metaventrite (
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<emphasis bold="true" box="[1254,1273,1676,1695]" pageId="3" pageNumber="3">x</emphasis>
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<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[1273,1446,1674,1695]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
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,
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<taxonomicName authorityName="Erichson" authorityYear="1845" box="[811,962,1703,1724]" class="Insecta" family="Sphaeriusidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Sphaeriusidae</taxonomicName>
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,
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<taxonomicName authorityName="Steffan" authorityYear="1964" box="[974,1132,1703,1724]" class="Insecta" family="Torridincolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Torridincolidae</taxonomicName>
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); mesoventrite with wide subpentagonal elevation (
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<emphasis bold="true" box="[1038,1057,1734,1753]" pageId="3" pageNumber="3">x</emphasis>
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modern
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<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[1143,1316,1733,1754]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
|
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,
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<taxonomicName authorityName="Steffan" authorityYear="1964" class="Insecta" family="Torridincolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Torridincolidae</taxonomicName>
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); elytra only slightly shortened, exposing 1 to 2 terminal abdominal tergites (
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<emphasis bold="true" box="[1021,1040,1792,1811]" pageId="3" pageNumber="3">x</emphasis>
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all modern families), truncated posteriorly (
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<emphasis bold="true" box="[866,885,1822,1841]" pageId="3" pageNumber="3">x</emphasis>
|
||
<taxonomicName authorityName="Erichson" authorityYear="1845" box="[885,1036,1820,1841]" class="Insecta" family="Sphaeriusidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Sphaeriusidae</taxonomicName>
|
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,
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||
<taxonomicName authorityName="Steffan" authorityYear="1964" box="[1049,1206,1820,1841]" class="Insecta" family="Torridincolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Torridincolidae</taxonomicName>
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,
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||
<taxonomicName authorityName="Hinton" authorityYear="1936" box="[1219,1344,1820,1841]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
|
||
); metanepisternum moderately wide anteriorly (
|
||
<emphasis bold="true" box="[1223,1242,1851,1870]" pageId="3" pageNumber="3">x</emphasis>
|
||
<taxonomicName authorityName="Erichson" authorityYear="1845" box="[1241,1392,1849,1870]" class="Insecta" family="Sphaeriusidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Sphaeriusidae</taxonomicName>
|
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,
|
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<taxonomicName authorityName="Steffan" authorityYear="1964" class="Insecta" family="Torridincolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Torridincolidae</taxonomicName>
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||
,
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||
<taxonomicName authorityName="Hinton" authorityYear="1936" box="[947,1072,1879,1900]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
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||
); metacoxal plates present, narrow (
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||
<emphasis bold="true" box="[817,836,1909,1928]" pageId="3" pageNumber="3">x</emphasis>
|
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<taxonomicName authorityName="Erichson" authorityYear="1845" box="[836,987,1908,1929]" class="Insecta" family="Sphaeriusidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Sphaeriusidae</taxonomicName>
|
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,
|
||
<taxonomicName authorityName="Steffan" authorityYear="1964" box="[1013,1170,1908,1929]" class="Insecta" family="Torridincolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Torridincolidae</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Hinton" authorityYear="1936" box="[1195,1320,1908,1929]" class="Insecta" family="Lepiceridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Lepiceridae</taxonomicName>
|
||
); abdomen with 5 to 6 exposed ventrites (
|
||
<figureCitation box="[1165,1255,1937,1958]" captionStart="Figure 3" captionStartId="4.[159,219,1537,1556]" captionTargetBox="[257,1369,275,1497]" captionTargetId="figure-79@4.[257,1370,271,1497]" captionTargetPageId="4" captionText="Figure 3. Phylogenetic position and morphology of Triamyxa coprolithica (A and B) Results of Bayesian analysis of Coleoptera dataset, with fossil position in Polyphaga excluded following the prothorax morphology:(A) complete beetle tree with color-coded posterior probabilities (PPs) of Triamyxa position; (B) maximum credibility tree with Polyphaga not shown. (C–E) Reconstruction of Triamyxa morphology: (C) dorsal view; (D) ventral view of a specimen with 5-segmented abdomen; (E) specimen with exposed 6th abdominal ventrite (see Figure 4 for comparison of all specimens). (F–H) Alternative positions of Triamyxa revealed by Myxophaga analyses: (F) Bayesian, molecular topology constrained; (G) maximum parsimony, molecular topology constrained; (H) maximum parsimony, topology unconstrained. (I–Q) Comparative morphology of the prothorax in ventral view:(I) Olibrus; (J) Acrotrichis; (K) Scaphidium; (L) Nebria; (M) Beutelius; (N) Lepicerus; (O) Satonius; (P) Sphaerius; (Q) Hydroscapha. (R–T) Prothoracic morphology of Triamyxa revealed by superimposition of disarticulated remains: (R) isolated pronotum; (S) isolated pronotum with superimposed prosternum from another specimen; (T) final reconstruction with the position of propleuron. See also Figures S2 and S3." figureDoi="http://doi.org/10.5281/zenodo.5061140" httpUri="https://zenodo.org/record/5061140/files/figure.png" pageId="3" pageNumber="3">Figure 4</figureCitation>
|
||
;
|
||
<emphasis bold="true" box="[1276,1295,1939,1958]" pageId="3" pageNumber="3">x</emphasis>
|
||
<taxonomicName authorityName="Erichson" authorityYear="1845" box="[1295,1446,1937,1958]" class="Insecta" family="Sphaeriusidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Sphaeriusidae</taxonomicName>
|
||
, some
|
||
<taxonomicName authorityName="Steffan" authorityYear="1964" box="[876,1034,1966,1987]" class="Insecta" family="Torridincolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="3" phylum="Arthropoda" rank="family">Torridincolidae</taxonomicName>
|
||
); all abdominal segments with separate tergite and sternite (
|
||
<emphasis bold="true" box="[369,388,1880,1899]" pageId="4" pageNumber="4">x</emphasis>
|
||
<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[388,562,1879,1900]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
|
||
). For more details, see Table S1. From its habitus,
|
||
<taxonomicName authority="Qvarnström & Fikáček & Vikberg Wernström & Huld & Beutel & Arriaga-Varela & Ahlberg & Niedźwiedzki, 2021" authorityName="Qvarnström & Fikáček & Vikberg Wernström & Huld & Beutel & Arriaga-Varela & Ahlberg & Niedźwiedzki" authorityYear="2021" box="[458,583,1908,1929]" family="Triamyxidae" higherTaxonomySource="Manual Input" pageId="4" pageNumber="3" rank="family" status="fam. nov.">Triamyxidae</taxonomicName>
|
||
is similar to
|
||
<taxonomicName authorityName="Le Conte" authorityYear="1874" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
|
||
but differs from this family in a series of characters listed above, most notably the lack of fused tergites and sternites of abdominal segments III–VI. This abdominal configuration, a very uncommon shared derived feature and synapomorphy of extant
|
||
<taxonomicName authorityName="Le Conte" authorityYear="1874" box="[1031,1200,1937,1958]" class="Insecta" family="Hydroscaphidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="family">Hydroscaphidae</taxonomicName>
|
||
and the Triassic
|
||
<taxonomicName class="Insecta" family="Staphylinidae" genus="Leehermania" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="4" pageNumber="4">Leehermania</emphasis>
|
||
</taxonomicName>
|
||
, is clearly absent from
|
||
<taxonomicName authorityName="Qvarnström & Fikáček & Vikberg Wernström & Huld & Beutel & Arriaga-Varela & Ahlberg & Niedźwiedzki" authorityYear="2021" box="[1155,1247,1966,1987]" class="Insecta" family="Hydroscaphidae" genus="Triamyxa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1155,1247,1966,1987]" italics="true" pageId="4" pageNumber="4">Triamyxa</emphasis>
|
||
</taxonomicName>
|
||
, where tergites and sternites of all abdominal segments are distinctly separated (
|
||
<figureCitation box="[148,247,1148,1169]" captionStart="Figure 1" captionStartId="2.[159,219,1501,1520]" captionTargetBox="[175,1450,271,1460]" captionTargetId="figure-82@2.[175,1450,271,1460]" captionTargetPageId="2" captionText="Figure 1. Contents of coprolite fragment ZPAL AbIII/3520 (A) The coprolite rendered semi-transparent with inclusions, such as beetle remains and fibrous networks, representing fungal colonies or algae visible. Top corner: silhouette of Silesaurus opolensis, the most probable coprolite producer. (B–D) The holotype specimen of Triamyxa coprolithica in ventral (B), lateral (C), and dorsal (D) views. (E and F) The second complete specimen in ventral (E) and anterior (F) views. Individual ventrites are indicated by roman numerals. (G) Triamyxa coprolithica preserved in various degrees of disarticulation. (H) An isolated head and two elytra that do not belong to Triamyxa coprolithica but to slightly larger beetles that were also ingested by the coprolite producer. (I) Examples of individual remains of Triamyxa coprolithica (meso- and metaventrite, head, head attached to pronotum, and three pronota). (J) Two of numerous elytra of similar size and morphology attributed to T. coprolithica. (K) Fibrous structures interpreted as fungi or algae. (L) A possible decomposed wood fragment. Color coding: blue,eyes; purple, antennae;light purple,legs. Abbreviations:abd,abdomen;ant,antenna; cox, coxa;ey, eye; fem, femur; hd, head;lgs, legs;pp, propleuron; thx, thorax; tib, tibia; troch, trochanter; trs, tarsi; ventr, ventrites. See also Figure S1." figureDoi="http://doi.org/10.5281/zenodo.5061136" httpUri="https://zenodo.org/record/5061136/files/figure.png" pageId="5" pageNumber="5">FigureS1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |