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<document id="ABB06C30F60F4BF8D4B1045A7247E433" ID-DOI="10.1186/s13358-022-00259-x" ID-ISSN="1664-2384" ID-PMC="PMC9681704" ID-PubMed="36439694" ID-Zenodo-Dep="12000082" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1718653166208" checkinUser="felipe" docAuthor="Leu, Marc, Bucher, Hugo, Vennemann, Torsten, Bagherpour, Borhan, Ji, Cheng, Brosse, Morgane &amp; Goudemand, Nicolas" docDate="2022" docId="038B1D1D645BFF83FF1BEC9BD2EBFBC7" docLanguage="en" docName="SwissJPalaeontol.141.19.pdf" docOrigin="Swiss Journal of Palaeontology (19) 141 (1)" docSource="http://dx.doi.org/10.1186/s13358-022-00259-x" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Urdyella tridenta Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand 2022, n. sp." docType="treatment" docVersion="10" lastPageNumber="51" masterDocId="FFB265656469FFB1FF8CED10D031FFE4" masterDocTitle="A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China" masterLastPageNumber="61" masterPageNumber="1" pageNumber="51" updateTime="1722285166916" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="42C2C9DD3FC6D3ECADA9804619A24B92">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="04D8637E74B5AC41D8818F5191E84F29">Leu, Marc</mods:namePart>
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<mods:namePart id="E653D2069A4188141D8284948A3A3F60">Vennemann, Torsten</mods:namePart>
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<mods:date id="17A3E15B92B8BF4AA470BEA25EE8411A">2022</mods:date>
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<subSubSection id="C338FF80645BFF83FF1BEC9BD1A5FE47" box="[151,404,395,419]" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="8B9DAC0B645BFF83FF1BEC9BD1A5FE47" blockId="50.[151,404,395,451]" box="[151,404,395,419]" pageId="50" pageNumber="51">
<heading id="D0D51B67645BFF83FF1BEC9BD1A5FE47" box="[151,404,395,419]" fontSize="9" level="2" pageId="50" pageNumber="51" reason="8">
<taxonomicName id="4C22D788645BFF83FF1BEC9BD164FE47" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[151,341,395,419]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="tridenta" status="sp. nov.">
<emphasis id="B9567019645BFF83FF1BEC9BD164FE47" box="[151,341,395,419]" italics="true" pageId="50" pageNumber="51">Urdyella tridenta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62645BFF83FED7EC9CD1A5FE47" box="[347,404,396,419]" pageId="50" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C338FF80645BFF83FF1BECBBD2EBFBC7" pageId="50" pageNumber="51" type="description">
<paragraph id="8B9DAC0B645BFF83FF1BECBBD12EFE27" blockId="50.[151,404,395,451]" box="[151,287,427,451]" pageId="50" pageNumber="51">
<figureCitation id="1319B08E645BFF83FF1BECBBD0EEFE27" box="[151,223,427,451]" captionStart="Fig" captionStartId="46.[169,200,1717,1737]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@47.[228,1362,254,1763]" captionTargetPageId="47" captionText="Fig. 26 Cornudininae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AW Urdyella unicorna n. gen. n. sp.; A LAR225, PIMUZ 39345; B LAR225, PIMUZ 39346;C LAR229, PIMUZ 39347; D LAR227, PIMUZ 39348;E LAR229,PIMUZ 39349; F LAR229,PIMUZ 39350; G LAR225,PIMUZ 39351; H LAR229,PIMUZ 39352; I QIA123, PIMUZ 39353; J LAR227,PIMUZ 39354; K QIA123, PIMUZ 39355; L QIA123, PIMUZ 39356; M LAR227,PIMUZ 39357; N LAR225, PIMUZ 39358; O QIA120, PIMUZ 39359; P LAR225, PIMUZ 39360; Q QIA121, PIMUZ 39361; R LAR232, PIMUZ 39362; S LAR232, PIMUZ 39363; T QIA124,PIMUZ 39364; U LAR232, PIMUZ 39365; V QIA123, PIMUZ 39366; W SHA341, PIMUZ 39367.X-AA Urdyella tridenta n. gen. n. sp.; X LAR227, PIMUZ 39341; Y LAR227, PIMUZ 39342; Z LAR227, PIMUZ 39343; AA LAR227, PIMUZ 39344.AB, AC Discretella? n. sp. D; AB QIA124, PIMUZ 39118; AC QIA123, PIMUZ 39119. AD Neostrachanognathus n. sp. A; LAR232, PIMUZ 39183. AE Urdyella n. sp. A; LAR232, PIMUZ 39340" figureDoi="http://doi.org/10.5281/zenodo.13127516" httpUri="https://zenodo.org/record/13127516/files/figure.png" pageId="50" pageNumber="51">Fig. 26</figureCitation>
X-AA
</paragraph>
<paragraph id="8B9DAC0B645BFF83FF1BECFBD207FDC7" blockId="50.[151,775,491,611]" pageId="50" pageNumber="51">
<emphasis id="B9567019645BFF83FF1BECFBD139FDE7" box="[151,264,491,515]" italics="true" pageId="50" pageNumber="51">Etymology</emphasis>
: Te species is named after the trident, Poseidons weapon in the Greek mythology.
</paragraph>
<paragraph id="8B9DAC0B645BFF83FF1BEF3BD24AFDA7" blockId="50.[151,775,491,611]" box="[151,635,555,579]" pageId="50" pageNumber="51">
<emphasis id="B9567019645BFF83FF1BEF3BD0CEFDA7" box="[151,255,555,579]" italics="true" pageId="50" pageNumber="51">
<typeStatus id="549912A9645BFF83FF1BEF3BD0CAFDA7" box="[151,251,555,579]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
.
</emphasis>
Specimen illustrated in
<figureCitation id="1319B08E645BFF83FD8AEF3BD27FFDA7" box="[518,590,555,579]" captionStart="Fig" captionStartId="46.[169,200,1717,1737]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@47.[228,1362,254,1763]" captionTargetPageId="47" captionText="Fig. 26 Cornudininae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AW Urdyella unicorna n. gen. n. sp.; A LAR225, PIMUZ 39345; B LAR225, PIMUZ 39346;C LAR229, PIMUZ 39347; D LAR227, PIMUZ 39348;E LAR229,PIMUZ 39349; F LAR229,PIMUZ 39350; G LAR225,PIMUZ 39351; H LAR229,PIMUZ 39352; I QIA123, PIMUZ 39353; J LAR227,PIMUZ 39354; K QIA123, PIMUZ 39355; L QIA123, PIMUZ 39356; M LAR227,PIMUZ 39357; N LAR225, PIMUZ 39358; O QIA120, PIMUZ 39359; P LAR225, PIMUZ 39360; Q QIA121, PIMUZ 39361; R LAR232, PIMUZ 39362; S LAR232, PIMUZ 39363; T QIA124,PIMUZ 39364; U LAR232, PIMUZ 39365; V QIA123, PIMUZ 39366; W SHA341, PIMUZ 39367.X-AA Urdyella tridenta n. gen. n. sp.; X LAR227, PIMUZ 39341; Y LAR227, PIMUZ 39342; Z LAR227, PIMUZ 39343; AA LAR227, PIMUZ 39344.AB, AC Discretella? n. sp. D; AB QIA124, PIMUZ 39118; AC QIA123, PIMUZ 39119. AD Neostrachanognathus n. sp. A; LAR232, PIMUZ 39183. AE Urdyella n. sp. A; LAR232, PIMUZ 39340" figureDoi="http://doi.org/10.5281/zenodo.13127516" httpUri="https://zenodo.org/record/13127516/files/figure.png" pageId="50" pageNumber="51">Fig. 26</figureCitation>
AA.
</paragraph>
<paragraph id="8B9DAC0B645BFF83FF1BEF5CD241FD87" blockId="50.[151,775,491,611]" box="[151,624,587,611]" pageId="50" pageNumber="51">
<emphasis id="B9567019645BFF83FF1BEF5CD131FD87" box="[151,256,588,611]" italics="true" pageId="50" pageNumber="51">
<typeStatus id="549912A9645BFF83FF1BEF5CD0CAFD87" box="[151,251,588,611]" pageId="50" pageNumber="51" type="paratype">Paratype</typeStatus>
.
</emphasis>
Specimens illustrated in
<figureCitation id="1319B08E645BFF83FD9DEF5BD25DFD87" box="[529,620,587,611]" captionStart="Fig" captionStartId="46.[169,200,1717,1737]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@47.[228,1362,254,1763]" captionTargetPageId="47" captionText="Fig. 26 Cornudininae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. AW Urdyella unicorna n. gen. n. sp.; A LAR225, PIMUZ 39345; B LAR225, PIMUZ 39346;C LAR229, PIMUZ 39347; D LAR227, PIMUZ 39348;E LAR229,PIMUZ 39349; F LAR229,PIMUZ 39350; G LAR225,PIMUZ 39351; H LAR229,PIMUZ 39352; I QIA123, PIMUZ 39353; J LAR227,PIMUZ 39354; K QIA123, PIMUZ 39355; L QIA123, PIMUZ 39356; M LAR227,PIMUZ 39357; N LAR225, PIMUZ 39358; O QIA120, PIMUZ 39359; P LAR225, PIMUZ 39360; Q QIA121, PIMUZ 39361; R LAR232, PIMUZ 39362; S LAR232, PIMUZ 39363; T QIA124,PIMUZ 39364; U LAR232, PIMUZ 39365; V QIA123, PIMUZ 39366; W SHA341, PIMUZ 39367.X-AA Urdyella tridenta n. gen. n. sp.; X LAR227, PIMUZ 39341; Y LAR227, PIMUZ 39342; Z LAR227, PIMUZ 39343; AA LAR227, PIMUZ 39344.AB, AC Discretella? n. sp. D; AB QIA124, PIMUZ 39118; AC QIA123, PIMUZ 39119. AD Neostrachanognathus n. sp. A; LAR232, PIMUZ 39183. AE Urdyella n. sp. A; LAR232, PIMUZ 39340" figureDoi="http://doi.org/10.5281/zenodo.13127516" httpUri="https://zenodo.org/record/13127516/files/figure.png" pageId="50" pageNumber="51">Fig. 26Z</figureCitation>
.
</paragraph>
<paragraph id="8B9DAC0B645BFF83FF1BEF9BD1A7FD47" blockId="50.[151,775,651,1060]" box="[151,406,651,675]" pageId="50" pageNumber="51">
<emphasis id="B9567019645BFF83FF1BEF9BD14CFD47" box="[151,381,651,675]" italics="true" pageId="50" pageNumber="51">Number of specimens</emphasis>
. 5
</paragraph>
<paragraph id="8B9DAC0B645BFF83FF1BEFBCD17FFCC7" blockId="50.[151,775,651,1060]" pageId="50" pageNumber="51">
<emphasis id="B9567019645BFF83FF1BEFBCD139FD27" box="[151,264,684,707]" italics="true" pageId="50" pageNumber="51">Diagnosis.</emphasis>
Coniform P
<subScript id="17A6AE4E645BFF83FE15EFA7D193FD2C" attach="left" box="[409,418,695,712]" fontSize="7" pageId="50" pageNumber="51">1</subScript>
element with three subequally long and slightly recurved denticles all attached to the basal cup. Deep, teardrop shaped basal cavity. No clear anterior process.
</paragraph>
<paragraph id="8B9DAC0B645BFF83FF20EE3BD087FC07" blockId="50.[151,775,651,1060]" pageId="50" pageNumber="51">
<emphasis id="B9567019645BFF83FF20EE3BD125FCA7" box="[172,276,811,835]" italics="true" pageId="50" pageNumber="51">Remarks.</emphasis>
Te P
<subScript id="17A6AE4E645BFF83FED1EE27D157FCAC" attach="left" box="[349,358,823,840]" fontSize="7" pageId="50" pageNumber="51">1</subScript>
element of this species is very similar to the one of
<taxonomicName id="4C22D788645BFF83FED8EE5BD210FC87" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[340,545,843,867]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="unicorna" status="sp. nov.">
<emphasis id="B9567019645BFF83FED8EE5BD210FC87" box="[340,545,843,867]" italics="true" pageId="50" pageNumber="51">Urdyella unicorna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62645BFF83FDA5EE5CD255FC87" box="[553,612,844,867]" pageId="50" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
except for the lack of an anterior process and the fact that the single large cusp of
<taxonomicName id="4C22D788645BFF83FEB7EE9BD23DFC47" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[315,524,907,931]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="unicorna" status="sp. nov.">
<emphasis id="B9567019645BFF83FEB7EE9BD23DFC47" box="[315,524,907,931]" italics="true" pageId="50" pageNumber="51">Urdyella unicorna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62645BFF83FD94EE9CD269FC47" box="[536,600,908,931]" pageId="50" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
is replaced by three subequally long denticles in
<taxonomicName id="4C22D788645BFF83FDAFEEBBD2D9FC27" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[547,744,939,963]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="tridenta" status="sp. nov.">
<emphasis id="B9567019645BFF83FDAFEEBBD2D9FC27" box="[547,744,939,963]" italics="true" pageId="50" pageNumber="51">Urdyella tridenta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD62645BFF83FD7EEEBCD087FC07" pageId="50" pageNumber="51" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8B9DAC0B645BFF83FF21EEFCD2EBFBC7" blockId="50.[151,775,651,1060]" pageId="50" pageNumber="51">
<emphasis id="B9567019645BFF83FF21EEFCD119FBE7" box="[173,296,1004,1027]" italics="true" pageId="50" pageNumber="51">Occurrence</emphasis>
. Laren road cut, Luolou Formation, Smithian age, Nanpanjiang basin, South
<collectingCountry id="F335EC9B645BFF83FD81E91BD27EFBC7" box="[525,591,1035,1059]" name="China" pageId="50" pageNumber="51">China</collectingCountry>
(this paper).
</paragraph>
</subSubSection>
</treatment>
</document>