treatments-xml/data/6A/5A/87/6A5A87CEFFDAB030FF7D7A9CFD20DCC0.xml

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<mods:title id="42281699133CECC44BB3A57CCAF9CD7F">Redescription of Rhyacophila yukii Tsuda 1942 and description of three new Rhyacophila species from Japan (Trichoptera: Rhyacophilidae)</mods:title>
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<mods:namePart id="C4203A7959D5017032377230E9511FF3">Torii, Takaaki</mods:namePart>
<mods:affiliation id="FCF8D461BD68D9AA987FF56655D19C98">Laboratory of Molecular Reproductive Biology, Graduate Division of Nutritional and Environmental Sciences, University of Shizuoka, 52 - 1, Yada, Suruga-Ku, Shizuoka, 422 - 8526, Japan &amp; Idea Consultant Inc., 1334 - 5 Riemon, Yaizu City, Shizuoka, 425 - 0084, Japan</mods:affiliation>
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<mods:namePart id="5B978C450CB349E6A447D348BFE633EA">Kobayashi, Tohru</mods:namePart>
<mods:affiliation id="85849B7FCFBDCD66EB12622B8237C669">Laboratory of Molecular Reproductive Biology, Graduate Division of Nutritional and Environmental Sciences, University of Shizuoka, 52 - 1, Yada, Suruga-Ku, Shizuoka, 422 - 8526, Japan</mods:affiliation>
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<paragraph id="E24C36D8FFDAB035FF7D7A9CFDA4D9AA" blockId="3.[151,543,1471,1534]" box="[151,543,1471,1498]" pageId="3" pageNumber="540">
<heading id="B90481B4FFDAB035FF7D7A9CFDA4D9AA" bold="true" box="[151,543,1471,1498]" fontSize="11" level="1" pageId="3" pageNumber="540" reason="1">
<emphasis id="D087EACAFFDAB035FF7D7A9CFDA4D9AA" bold="true" box="[151,543,1471,1498]" pageId="3" pageNumber="540">
<taxonomicName id="25F34D5BFFDAB035FF7D7A9CFE6DD9AA" authority="Torii" authorityName="Torii &amp; Kobayashi" authorityYear="2021" box="[151,470,1471,1498]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="hattorii" status="sp. nov.">
<emphasis id="D087EACAFFDAB035FF7D7A9CFE29D9AA" bold="true" box="[151,402,1471,1498]" italics="true" pageId="3" pageNumber="540">Rhyacophila hattorii</emphasis>
Torii
</taxonomicName>
<taxonomicNameLabel id="CBB457B1FFDAB035FE377AE3FDA4D9AA" box="[477,543,1472,1498]" pageId="3" pageNumber="540" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="E24C36D8FFDAB035FF7D7AC7FEA6D98E" blockId="3.[151,543,1471,1534]" box="[151,285,1508,1534]" pageId="3" pageNumber="540">
<figureCitation id="7AC82A5DFFDAB035FF7D7AC7FEA6D98E" box="[151,285,1508,1534]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="3" pageNumber="540">Figs 2A–2L</figureCitation>
</paragraph>
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<subSubSection id="AAE96553FFDAB035FF7D7900FEA8DA08" pageId="3" pageNumber="540" type="reference_group">
<paragraph id="E24C36D8FFDAB035FF7D7900FEA8DA08" blockId="3.[151,1437,1571,1656]" pageId="3" pageNumber="540">
<taxonomicName id="25F34D5BFFDAB035FF7D7900FE57DA4B" ID-CoL="4SP8S" authority=": Schmid 1970" authorityName="Schmid" authorityYear="1970" box="[151,492,1571,1595]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDAB035FF7D7900FEEADA4B" box="[151,337,1571,1595]" italics="true" pageId="3" pageNumber="540">Rhyacophila yukii</emphasis>
: 
<bibRefCitation id="86624B29FFDAB035FE8B7900FE57DA4B" author="Schmid, F." box="[353,492,1571,1595]" pageId="3" pageNumber="540" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid 1970</bibRefCitation>
</taxonomicName>
, 197, figs 1–3, male, 
<emphasis id="D087EACAFFDAB035FD387907FD43DA4B" box="[722,760,1572,1595]" italics="true" pageId="3" pageNumber="540">non</emphasis>
<bibRefCitation id="86624B29FFDAB035FCEA7900FC30DA4B" author="Tsuda, M." box="[768,907,1571,1595]" pageId="3" pageNumber="540" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda (1942)</bibRefCitation>
; 
<bibRefCitation id="86624B29FFDAB035FC727907FBE4DA4B" author="Kagaya, T. &amp; Nozaki, T. &amp; Kuranishi, R. B." box="[920,1119,1571,1596]" pageId="3" pageNumber="540" pagination="1 - 266" refId="ref7712" refString="Kagaya, T., Nozaki, T. &amp; Kuranishi, R. B. (1998) Fauna and distribution of Trichoptera in the Tama-River system. In: Katagiri, K. (Ed.), Fauna and Distribution of Trichoptera in the Tama-River System. Tokyu Foundation for Better Environment, Tokyo, pp. 1 - 266. [in Japanese]" type="book chapter" year="1998">Kagaya et al. 1998</bibRefCitation>
, pp. 22, 95, list. 
<bibRefCitation id="86624B29FFDAB035FAF97900FA23DA4B" author="Hattori, T." box="[1299,1432,1571,1595]" pageId="3" pageNumber="540" pagination="474 - 497" refId="ref7502" refString="Hattori, T. (2005) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations. Tokai University Press, Kanagawa, pp. 474 - 497. [in Japanese]" type="book chapter" year="2005">Hattori 2005</bibRefCitation>
, pp. 427, 435, fig. 5, male, female; 
<bibRefCitation id="86624B29FFDAB035FDC77961FCA8DA2A" author="Inazu, K. &amp; Nishida, A." box="[557,787,1602,1626]" pageId="3" pageNumber="540" pagination="169 - 212" refId="ref7610" refString="Inazu, K. &amp; Nishida, A. (2011) Trichoptera. In: Editorial office of Hyogo Fresh Water Biology (Ed.), Hyogo no kawa no ikimono zukan. Hyogo Rikusui-seibutsu Kenkyu-kai, Himeji, Hyogo, pp. 169 - 212. [in Japanese]" type="book chapter" year="2011">Inazu &amp; Nishida 2011</bibRefCitation>
, p. 176, figs 21, male; 
<bibRefCitation id="86624B29FFDAB035FBEB7961FB3DDA2A" author="Hattori, T." box="[1025,1158,1602,1626]" pageId="3" pageNumber="540" pagination="414 - 437" refId="ref7554" refString="Hattori, T. (2018) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations, 2 nd Edition. Tokai University Press, Kanagawa, pp. 414 - 437. [in Japanese]" type="book chapter" year="2018">Hattori 2018</bibRefCitation>
, pp. 487, 495, fig. 5, male, female.
</paragraph>
</subSubSection>
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<emphasis id="D087EACAFFDAB035FF7D7982FEA9DACB" bold="true" box="[151,274,1697,1723]" pageId="3" pageNumber="540">Diagnosis.</emphasis>
This new species is similar to other members of the 
<taxonomicName id="25F34D5BFFDAB035FC817982FBEDDACB" authorityName="R McLachlan" authorityYear="1879" box="[875,1110,1697,1723]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="sibirica">
<emphasis id="D087EACAFFDAB035FC817982FBEDDACB" box="[875,1110,1697,1723]" italics="true" pageId="3" pageNumber="540">Rhyacophila sibirica</emphasis>
</taxonomicName>
Group (
<taxonomicName id="25F34D5BFFDAB035FB5D7981FF69DAAF" authority="Tsuda 1940" authorityName="Tsuda" authorityYear="1940" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="transquilla">
<emphasis id="D087EACAFFDAB035FB5D7981FB73DACB" box="[1207,1224,1698,1723]" italics="true" pageId="3" pageNumber="540">R</emphasis>
. 
<emphasis id="D087EACAFFDAB035FB3D7982FAE9DACB" box="[1239,1362,1697,1723]" italics="true" pageId="3" pageNumber="540">transquilla</emphasis>
<bibRefCitation id="86624B29FFDAB035FAB07982FF69DAAF" author="Tsuda, M." pageId="3" pageNumber="540" pagination="119 - 135" refId="ref9082" refString="Tsuda, M. (1940) Zur kenntnis der Japanischen Rhyacophilinen (Rhyacophilidae, Trichoptera). Annotationes Zoologicae Japo- nenses, 19, 119 - 135. [in German]" type="journal article" year="1940">Tsuda 1940</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="25F34D5BFFDAB035FF3479E5FD83DAAF" authority="Lukyanchenko 1993" authorityName="Lukyanchenko" authorityYear="1993" box="[222,568,1733,1760]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="arefini">
<emphasis id="D087EACAFFDAB035FF3479E5FF54DAAF" box="[222,239,1734,1759]" italics="true" pageId="3" pageNumber="540">R</emphasis>
. 
<emphasis id="D087EACAFFDAB035FF1779E6FEFCDAAF" box="[253,327,1733,1759]" italics="true" pageId="3" pageNumber="540">arefini</emphasis>
<bibRefCitation id="86624B29FFDAB035FEA579E6FD83DAAF" author="Lukyanchenko, T. I." box="[335,568,1733,1759]" pageId="3" pageNumber="540" pagination="5 - 6" refId="ref8239" refString="Lukyanchenko, T. I. (1993) A new species of caddisfly of the genus Rhyacophila Pictet (Trichoptera: Rhyacophilidae) from eastern Asia. Braueria, 20, 5 - 6." type="journal article" year="1993">Lukyanchenko 1993</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="25F34D5BFFDAB035FDA979E5FCEFDAAF" authority="Tsuda 1940" authorityName="Tsuda" authorityYear="1940" box="[579,852,1733,1760]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="kisoensis">
<emphasis id="D087EACAFFDAB035FDA979E5FDEFDAAF" box="[579,596,1734,1759]" italics="true" pageId="3" pageNumber="540">R</emphasis>
. 
<emphasis id="D087EACAFFDAB035FD8979E6FD73DAAF" box="[611,712,1733,1759]" italics="true" pageId="3" pageNumber="540">kisoensis</emphasis>
<bibRefCitation id="86624B29FFDAB035FD2579E6FCEFDAAF" author="Tsuda, M." box="[719,852,1733,1759]" pageId="3" pageNumber="540" pagination="119 - 135" refId="ref9082" refString="Tsuda, M. (1940) Zur kenntnis der Japanischen Rhyacophilinen (Rhyacophilidae, Trichoptera). Annotationes Zoologicae Japo- nenses, 19, 119 - 135. [in German]" type="journal article" year="1940">Tsuda 1940</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="25F34D5BFFDAB035FC7A79E5FBCBDAAF" authority="Tsuda 1942" authorityName="Tsuda" authorityYear="1942" box="[912,1136,1733,1760]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDAB035FC7A79E5FC1ADAAF" box="[912,929,1734,1759]" italics="true" pageId="3" pageNumber="540">R</emphasis>
. 
<emphasis id="D087EACAFFDAB035FC4579E6FC5EDAAF" box="[943,997,1733,1759]" italics="true" pageId="3" pageNumber="540">yukii</emphasis>
<bibRefCitation id="86624B29FFDAB035FC0779E6FBCBDAAF" author="Tsuda, M." box="[1005,1136,1733,1759]" pageId="3" pageNumber="540" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda 1942</bibRefCitation>
</taxonomicName>
) in male genitalia, but can be diagnosed easily from them by the following combination of segment X characters: (1) in dorsal view somewhat hexagonal to oval; (2) in lateral view with the upper and lower portions more swollen than the center; (3) in caudal view hexagonal with its dorsal apex incised mesally, elongate downward, ventral apex incised and sclerotized. The female vaginal apparatus of this species is similar to that of 
<taxonomicName id="25F34D5BFFDAB035FC837875FB36DB1F" authority="Arefina 1993" authorityName="Arefina" authorityYear="1993" box="[873,1165,1877,1904]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="implicata">
<emphasis id="D087EACAFFDAB035FC837875FCC1DB1F" box="[873,890,1878,1903]" italics="true" pageId="3" pageNumber="540">R</emphasis>
. 
<emphasis id="D087EACAFFDAB035FC637876FC48DB1F" box="[905,1011,1877,1903]" italics="true" pageId="3" pageNumber="540">implicata</emphasis>
Arefina 1993
</taxonomicName>
(in 
<bibRefCitation id="86624B29FFDAB035FB507876FA34DB1F" author="Schmid, F. &amp; Arefina, T. J. &amp; Levanidova, I. M." box="[1210,1423,1877,1904]" pageId="3" pageNumber="540" pagination="161 - 172" refId="ref8796" refString="Schmid, F., Arefina, T. J. &amp; Levanidova, I. M. (1993) Contribution to the knowledge of the Rhyacophia (Trichoptera) of the sibirica group. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Entomologie, 63, 161 - 172." type="journal article" year="1993">Schmid et al. 1993</bibRefCitation>
), distributed in 
<collectingCountry id="9AE47648FFDAB035FEDF785AFE38DBE3" box="[309,387,1913,1939]" name="Russia" pageId="3" pageNumber="540">Russia</collectingCountry>
. However, it can be distinguished by the posterior process of the vaginal apparatus, which is shorter than that of 
<taxonomicName id="25F34D5BFFDAB035FE9878BDFE47DBC7" authorityName="Arefina" authorityYear="1993" box="[370,508,1949,1975]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="implicata">
<emphasis id="D087EACAFFDAB035FE9878BDFE47DBC7" box="[370,508,1949,1975]" italics="true" pageId="3" pageNumber="540">R. implicata</emphasis>
</taxonomicName>
.
</paragraph>
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<paragraph id="E24C36D8FFDDB032FF7D799CFCF4DB87" blockId="4.[151,1437,1727,2039]" pageId="4" pageNumber="541">
<emphasis id="D087EACAFFDDB032FF7D799CFEE3DAA7" bold="true" box="[151,344,1727,1751]" pageId="4" pageNumber="541">FIGURES 2A−2L</emphasis>
. 
<taxonomicName id="25F34D5BFFDDB032FE8F799CFD8CDAA7" authority="Torii &amp; Kobayashi, 2021" authorityName="Torii &amp; Kobayashi" authorityYear="2021" box="[357,567,1727,1751]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="541" phylum="Arthropoda" rank="species" species="hattorii" status="sp. nov.">
<emphasis id="D087EACAFFDDB032FE8F799CFD8CDAA7" box="[357,567,1727,1751]" italics="true" pageId="4" pageNumber="541">Rhyacophila hattorii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CBB457B1FFDDB032FDD779E3FD02DAA7" box="[573,697,1727,1751]" pageId="4" pageNumber="541" rank="species">new species</taxonomicNameLabel>
. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by 
<bibRefCitation id="86624B29FFDDB032FF7D786CFE95DB17" author="Schmid, F." box="[151,302,1871,1895]" pageId="4" pageNumber="541" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid (1970)</bibRefCitation>
and labelled as “ 
<taxonomicName id="25F34D5BFFDDB032FE347873FD90DB17" authorityName="Tsuda" authorityYear="1942" box="[478,555,1871,1895]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="541" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDDB032FE347873FE56DB17" box="[478,493,1872,1895]" italics="true" pageId="4" pageNumber="541">R</emphasis>
. 
<emphasis id="D087EACAFFDDB032FE10786CFD90DB17" box="[506,555,1871,1895]" italics="true" pageId="4" pageNumber="541">yukii</emphasis>
</taxonomicName>
”; 2L, dorsal, by 
<bibRefCitation id="86624B29FFDDB032FD32786CFCD4DB17" author="Schmid, F." box="[728,879,1871,1895]" pageId="4" pageNumber="541" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid (1970)</bibRefCitation>
and labelled as “ 
<taxonomicName id="25F34D5BFFDDB032FBF47873FBD7DB17" authorityName="Tsuda" authorityYear="1942" box="[1054,1132,1871,1895]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="541" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDDB032FBF47873FB96DB17" box="[1054,1069,1872,1895]" italics="true" pageId="4" pageNumber="541">R</emphasis>
. 
<emphasis id="D087EACAFFDDB032FBD1786CFBD7DB17" box="[1083,1132,1871,1895]" italics="true" pageId="4" pageNumber="541">yukii</emphasis>
</taxonomicName>
”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus.
</paragraph>
</caption>
<paragraph id="E24C36D8FFDCB033FF2D7FB4FA95DCA5" blockId="5.[151,1437,150,1038]" pageId="5" pageNumber="542">
<emphasis id="D087EACAFFDCB033FF2D7FB4FEBFDCC1" bold="true" box="[199,260,151,177]" pageId="5" pageNumber="542">Male</emphasis>
. Length of each male forewing 
<quantity id="250B9B3DFFDCB033FD817FB5FD49DCC1" box="[619,754,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" metricValueMax="8.5" metricValueMin="7.6" pageId="5" pageNumber="542" unit="mm" value="8.05" valueMax="8.5" valueMin="7.6">7.6–8.5 mm</quantity>
(n = 27). Specimens in alcohol with head, thorax, and wings dark brown; legs yellowish brown, with spurs dark brown. Segments VI and VII with thin spur ventrally.
</paragraph>
<paragraph id="E24C36D8FFDCB033FF2D7FFCFDF5DE4D" blockId="5.[151,1437,150,1038]" pageId="5" pageNumber="542">
<emphasis id="D087EACAFFDCB033FF2D7FFCFECADC88" bold="true" box="[199,369,222,249]" pageId="5" pageNumber="542">Male genitalia</emphasis>
(
<figureCitation id="7AC82A5DFFDCB033FE6A7FFCFDB0DC89" box="[384,523,223,249]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Figs 2A–2F</figureCitation>
). Segment IX in lateral view subrectangular, with upper portion longer than lower one, rectangular in dorsal view, twice as wide as long. Segment X in lateral view with upper and lower portions swollen and club-shaped, in dorsal view somewhat hexagonal to oval with its apex incised mesally, in caudal view hexagonal with dorsal apex incised mesally, elongate downward, ventral apex incised and sclerotized. Apical band (a.b.) in lateral view long and narrow, bar shaped. Ventral branch of aedeagus (v.b.a.) in lateral view cylindrical and protruding; aedeagus (aed.) in lateral view curved about 80°, resembling long and narrow spine; ventral lobe of aedeagus (v.l.a.) elongate subelliptical in dorsal view. Basal segment of each inferior appendage (b.i.a.) somewhat trapezoidal in lateral view, in ventral view with its inner and outer margins nearly straight; apical segment of each inferior appendage (a.i.a.) in lateral view with apex depressed obliquely, in ventral view with its inner and outer margins nearly straight, round apically.
</paragraph>
<paragraph id="E24C36D8FFDCB033FF2D7D64FCC0DE11" blockId="5.[151,1437,150,1038]" box="[199,891,583,609]" pageId="5" pageNumber="542">
<emphasis id="D087EACAFFDCB033FF2D7D64FEA5DE11" bold="true" box="[199,286,583,609]" pageId="5" pageNumber="542">Female</emphasis>
. Length of each female forewing 
<quantity id="250B9B3DFFDCB033FD707D64FC9ADE11" box="[666,801,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.8" metricValueMin="7.8" pageId="5" pageNumber="542" unit="mm" value="8.3" valueMax="8.8" valueMin="7.8">7.8–8.8 mm</quantity>
(n = 8).
</paragraph>
<paragraph id="E24C36D8FFDCB033FF2D7D49FB41DF2B" blockId="5.[151,1437,150,1038]" pageId="5" pageNumber="542">
<emphasis id="D087EACAFFDCB033FF2D7D49FE35DEF4" bold="true" box="[199,398,618,644]" pageId="5" pageNumber="542">Female genitalia</emphasis>
(
<figureCitation id="7AC82A5DFFDCB033FE747D49FD9DDEF4" box="[414,550,618,644]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Figs 2G–2J</figureCitation>
). Segment VI with thin spur ventrally. Corners of tergum and sternum of segment VII rounded in outline. Segment VIII cone shaped. Segments IX, X, XI (
<figureCitation id="7AC82A5DFFDCB033FC147DADFBEEDED8" box="[1022,1109,654,680]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Fig. 2H</figureCitation>
) often retracted into segment VIII or VII and VIII (
<figureCitation id="7AC82A5DFFDCB033FE797D91FE4BDEBC" box="[403,496,690,716]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Fig. 2G</figureCitation>
). Segment IX membranous, trapezoidal, indistinguishable from the intersegmental membrane VIII–X, proximal part slightly wider than distal edge. Segment X cylindrical. Segment XI with pair of cerci (c.). Apodemal rods from segment VIII (ap.r.VIII) extend forward to segment VII. Apodemal rods from segment XI (ap.r.XI) extending forward to segment VI or VII. Posterior process of vaginal apparatus (p.pr.) in dorsal view long elliptical with apex slightly incised (
<figureCitation id="7AC82A5DFFDCB033FD4F7C62FD48DF2B" box="[677,755,833,859]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Fig. 2I</figureCitation>
), in lateral view elongate bar shaped (
<figureCitation id="7AC82A5DFFDCB033FB777C62FB55DF2B" box="[1181,1262,833,859]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Fig. 2J</figureCitation>
).
</paragraph>
<paragraph id="E24C36D8FFDCB033FF2D7C46FDA8DF0E" blockId="5.[151,1437,150,1038]" box="[199,531,868,895]" pageId="5" pageNumber="542">
<emphasis id="D087EACAFFDCB033FF2D7C46FE34DF0F" bold="true" box="[199,399,869,895]" pageId="5" pageNumber="542">Immature stages</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="AAE96553FFDCB033FF2D7CABFCBBDA73" pageId="5" pageNumber="542" type="materials_examined">
<paragraph id="E24C36D8FFDCB033FF2D7CABFDF4DFB6" blockId="5.[151,1437,150,1038]" pageId="5" pageNumber="542">
<materialsCitation id="529B3C85FFDCB033FF2D7CABFDF4DFB6" collectingDate="1991-06-06" collectionCode="T, SEHU" county="Male" elevation="1350" latitude="35.8432" location="Kohshu-shi" longLatPrecision="6" longitude="138.8336" municipality="Hondani" pageId="5" pageNumber="542" specimenCount="1" stateProvince="Yamanashi" typeStatus="holotype">
<emphasis id="D087EACAFFDCB033FF2D7CABFE8FDFD2" bold="true" box="[199,308,904,930]" pageId="5" pageNumber="542">
<typeStatus id="3D48887AFFDCB033FF2D7CABFE8FDFD2" box="[199,308,904,930]" pageId="5" pageNumber="542" type="holotype">Holotype</typeStatus>
</emphasis>
. 
<collectingCounty id="0B2D4E54FFDCB033FEAE7CABFE3BDFD2" box="[324,384,904,930]" pageId="5" pageNumber="542">Male</collectingCounty>
, 
<collectingMunicipality id="0228ACA2FFDCB033FE677CABFE4BDFD2" box="[397,496,904,930]" pageId="5" pageNumber="542">Hondani</collectingMunicipality>
, Enzan-Icinose-Takahashi, 
<location id="E72C6003FFDCB033FCC27CABFC17DFD2" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FCC27CABFC17DFD2" box="[808,940,904,930]" county="Male" latitude="35.8432" longLatPrecision="6" longitude="138.8336" municipality="Hondani" name="Kohshu-shi" pageId="5" pageNumber="542" stateProvince="Yamanashi">Kohshu-shi</location>
, 
<collectingRegion id="2037F83AFFDCB033FC527CABFB88DFD2" box="[952,1075,904,930]" country="Japan" name="Yamanashi" pageId="5" pageNumber="542">Yamanashi</collectingRegion>
(
<geoCoordinate id="87C7501FFFDCB033FBA97CABFB79DFD3" box="[1091,1218,904,931]" degrees="35.8432" direction="north" orientation="latitude" pageId="5" pageNumber="542" precision="5" value="35.8432">35.8432°N</geoCoordinate>
, 
<geoCoordinate id="87C7501FFFDCB033FB257CABFAECDFD2" box="[1231,1367,904,930]" degrees="138.8336" direction="east" orientation="longitude" pageId="5" pageNumber="542" precision="5" value="138.8336">138.8336°E</geoCoordinate>
, 
<elevation id="69DED1EBFFDCB033FA8E7CABFF52DFB7" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="5" pageNumber="542" unit="m" value="1350.0">
<quantity id="250B9B3DFFDCB033FA8E7CABFF16DFB7" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="5" pageNumber="542" unit="m" value="1350.0">1350 m</quantity>
a.s.l.
</elevation>
), 
<date id="964D1018FFDCB033FF157C8FFED0DFB6" box="[255,363,940,966]" pageId="5" pageNumber="542" value="1991-06-06">
<collectingDate id="8609E9F0FFDCB033FF157C8FFED0DFB6" box="[255,363,940,966]" pageId="5" pageNumber="542" value="1991-06-06">6.vi.1991</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FE9D7C8EFE32DFB7" box="[375,393,941,967]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
. Nozaki (
<collectionCode id="84E2AE1DFFDCB033FE1E7C8FFDF8DFB6" box="[500,579,940,966]" pageId="5" pageNumber="542">SEHU</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="E24C36D8FFDCB033FF2D7CF3FD71D87E" blockId="5.[151,1437,150,1038]" pageId="5" pageNumber="542">
<materialsCitation id="529B3C85FFDCB033FF2D7CF3FAF9DF9A" box="[199,1346,976,1002]" collectingDate="1994-06-11" collectionCode="T" county="Kawamata-gawa" location="Hokuto-shi" municipality="Ohizumi-cho" pageId="5" pageNumber="542" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Yamanashi" typeStatus="paratype">
<emphasis id="D087EACAFFDCB033FF2D7CF3FE84DF9A" bold="true" box="[199,319,976,1002]" pageId="5" pageNumber="542">
<typeStatus id="3D48887AFFDCB033FF2D7CF3FE84DF9A" box="[199,319,976,1002]" pageId="5" pageNumber="542" type="paratype">Paratypes</typeStatus>
</emphasis>
. 
<specimenCount id="F4F5FD51FFDCB033FEA77CF3FE1DDF9A" box="[333,422,976,1002]" pageId="5" pageNumber="542" type="male">4 males</specimenCount>
, 
<specimenCount id="F4F5FD51FFDCB033FE5B7CF3FD9BDF9A" box="[433,544,976,1002]" pageId="5" pageNumber="542" type="female">3 females</specimenCount>
, 
<collectingCounty id="0B2D4E54FFDCB033FDC17CF3FD51DF9A" box="[555,746,976,1002]" pageId="5" pageNumber="542">Kawamata-gawa</collectingCounty>
, 
<collectingMunicipality id="0228ACA2FFDCB033FD1C7CF3FC36DF9A" box="[758,909,976,1002]" pageId="5" pageNumber="542">Ohizumi-cho</collectingMunicipality>
, 
<location id="E72C6003FFDCB033FC7D7CF3FBA3DF9A" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FC7D7CF3FBA3DF9A" box="[919,1048,976,1002]" county="Kawamata-gawa" municipality="Ohizumi-cho" name="Hokuto-shi" pageId="5" pageNumber="542" stateProvince="Yamanashi">Hokuto-shi</location>
, 
<collectingRegion id="2037F83AFFDCB033FBC87CF3FB1BDF9A" box="[1058,1184,976,1002]" country="Japan" name="Yamanashi" pageId="5" pageNumber="542">Yamanashi</collectingRegion>
, 
<date id="964D1018FFDCB033FB467CF3FA9EDF9A" box="[1196,1317,976,1002]" pageId="5" pageNumber="542" value="1994-06-11">
<collectingDate id="8609E9F0FFDCB033FB467CF3FA9EDF9A" box="[1196,1317,976,1002]" pageId="5" pageNumber="542" value="1994-06-11">11.vi.1994</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FADA7CF3FAF9DF9A" box="[1328,1346,976,1002]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="529B3C85FFDCB033FAA77CF3FF4DD87E" collectionCode="SEHU" location="Hattori" pageId="5" pageNumber="542" specimenCount="1" stateProvince="Yamanashi" typeStatus="paratype">
<location id="E72C6003FFDCB033FAA77CF3FA27DF9A" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FAA77CF3FA27DF9A" box="[1357,1436,976,1002]" name="Hattori" pageId="5" pageNumber="542" stateProvince="Yamanashi">Hattori</location>
(
<collectionCode id="84E2AE1DFFDCB033FF757CD7FF55D87E" box="[159,238,1012,1038]" pageId="5" pageNumber="542">SEHU</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="529B3C85FFDCB033FEE87CD0FD7DD87E" box="[258,710,1011,1038]" collectingDate="1991-06-06" collectionCode="CBM" county="Male" elevation="1350" latitude="35.8432" location="Kohshu-shi" longLatPrecision="6" longitude="138.8336" municipality="Hondani" pageId="5" pageNumber="542" specimenCount="16" specimenCount-male="16" stateProvince="Yamanashi" typeStatus="paratype">
<specimenCount id="F4F5FD51FFDCB033FEE87CD0FED2D87D" box="[258,361,1011,1037]" pageId="5" pageNumber="542" type="male">16 males</specimenCount>
, same data as holotype (
<collectionCode id="84E2AE1DFFDCB033FD907CD7FD05D87E" box="[634,702,1012,1038]" country="Japan" lsid="urn:lsid:biocol.org:col:13737" name="Natural History Museum and Institute" pageId="5" pageNumber="542" type="Herbarium">CBM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E24C36D8FFDCB033FF2D7B18FCBBDA73" blockId="5.[151,1437,1083,2043]" pageId="5" pageNumber="542">
<emphasis id="D087EACAFFDCB033FF2D7B18FE4DD825" bold="true" box="[199,502,1083,1109]" pageId="5" pageNumber="542">Other material examined</emphasis>
. 
<materialsCitation id="529B3C85FFDCB033FDEE7B18FA23D825" box="[516,1432,1083,1110]" collectingDate="1991-06-05" collectionCode="T" county="Nippara-gawa" location="Nishiokutama-gun" municipality="Okutama-machi" pageId="5" pageNumber="542" specimenCount="4" specimenCount-male="4" stateProvince="Tokyo">
<specimenCount id="F4F5FD51FFDCB033FDEE7B18FDE5D825" box="[516,606,1083,1109]" pageId="5" pageNumber="542" type="male">4 males</specimenCount>
, 
<collectingCounty id="0B2D4E54FFDCB033FD807B18FCB7D825" box="[618,780,1083,1109]" pageId="5" pageNumber="542">Nippara-gawa</collectingCounty>
, 
<collectingMunicipality id="0228ACA2FFDCB033FCFD7B18FC75D825" box="[791,974,1083,1109]" pageId="5" pageNumber="542">Okutama-machi</collectingMunicipality>
, 
<location id="E72C6003FFDCB033FC307B18FB16D825" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FC307B18FB16D825" box="[986,1197,1083,1109]" county="Nippara-gawa" municipality="Okutama-machi" name="Nishiokutama-gun" pageId="5" pageNumber="542" stateProvince="Tokyo">Nishiokutama-gun</location>
, 
<collectingRegion id="2037F83AFFDCB033FB537B18FAB8D825" box="[1209,1283,1083,1109]" country="Japan" name="Tokyo" pageId="5" pageNumber="542">Tokyo</collectingRegion>
, 
<date id="964D1018FFDCB033FAE47B18FAC0D826" box="[1294,1403,1083,1110]" pageId="5" pageNumber="542" value="1991-06-05">
<collectingDate id="8609E9F0FFDCB033FAE47B18FAC0D826" box="[1294,1403,1083,1110]" pageId="5" pageNumber="542" value="1991-06-05">5.vi.1991</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FA6C7B18FA23D825" box="[1414,1432,1083,1109]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
</materialsCitation>
. Nozaki; 
<materialsCitation id="529B3C85FFDCB033FF137B7CFC70D809" box="[249,971,1119,1145]" collectingDate="2007-07-19" collectionCode="T" location="Katsuyama-shi" municipality="Akausagi-yama" pageId="5" pageNumber="542" specimenCount="4" specimenCount-male="4" stateProvince="Fukui">
<specimenCount id="F4F5FD51FFDCB033FF137B7CFEEFD809" box="[249,340,1119,1145]" pageId="5" pageNumber="542" type="male">4 males</specimenCount>
, 
<collectingMunicipality id="0228ACA2FFDCB033FE8A7B7CFDA8D809" box="[352,531,1119,1145]" pageId="5" pageNumber="542">Akausagi-yama</collectingMunicipality>
, 
<location id="E72C6003FFDCB033FDCA7B7CFD70D809" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FDCA7B7CFD70D809" box="[544,715,1119,1145]" municipality="Akausagi-yama" name="Katsuyama-shi" pageId="5" pageNumber="542" stateProvince="Fukui">Katsuyama-shi</location>
, 
<collectingRegion id="2037F83AFFDCB033FD327B7CFCA7D809" box="[728,796,1119,1145]" country="Japan" name="Fukui" pageId="5" pageNumber="542">Fukui</collectingRegion>
, 
<date id="964D1018FFDCB033FCC07B7CFC17D809" box="[810,940,1119,1145]" pageId="5" pageNumber="542" value="2007-07-19">
<collectingDate id="8609E9F0FFDCB033FCC07B7CFC17D809" box="[810,940,1119,1145]" pageId="5" pageNumber="542" value="2007-07-19">19.vii.2007</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FC537B7CFC70D809" box="[953,971,1119,1145]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="529B3C85FFDCB033FC327B7CFBD8D809" box="[984,1123,1119,1145]" collectionCode="SPMN" location="Ito" pageId="5" pageNumber="542" specimenCount="1" stateProvince="Fukui">
<location id="E72C6003FFDCB033FC327B7CFC4CD809" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FC327B7CFC4CD809" box="[984,1015,1119,1145]" name="Ito" pageId="5" pageNumber="542" stateProvince="Fukui">Ito</location>
(
<collectionCode id="84E2AE1DFFDCB033FBE27B7CFBE0D809" box="[1032,1115,1119,1145]" pageId="5" pageNumber="542">SPMN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="529B3C85FFDCB033FB9A7B7CFD86D8ED" collectingDate="2008-07-19" collectionCode="T" location="Katsuyama-shi" municipality="Akausagi-yama" pageId="5" pageNumber="542" specimenCount="10" specimenCount-male="10" stateProvince="Fukui">
<specimenCount id="F4F5FD51FFDCB033FB9A7B7CFB61D809" box="[1136,1242,1119,1145]" pageId="5" pageNumber="542" type="male">10 males</specimenCount>
, 
<collectingMunicipality id="0228ACA2FFDCB033FB0C7B7CFA22D809" box="[1254,1433,1119,1145]" pageId="5" pageNumber="542">Akausagi-yama</collectingMunicipality>
, 
<location id="E72C6003FFDCB033FF7D7BA1FEF9D8EC" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FF7D7BA1FEF9D8EC" box="[151,322,1154,1180]" municipality="Akausagi-yama" name="Katsuyama-shi" pageId="5" pageNumber="542" stateProvince="Fukui">Katsuyama-shi</location>
, 
<collectingRegion id="2037F83AFFDCB033FEA47BA1FE29D8EC" box="[334,402,1154,1180]" country="Japan" name="Fukui" pageId="5" pageNumber="542">Fukui</collectingRegion>
, 
<date id="964D1018FFDCB033FE747BA1FD9BD8EC" box="[414,544,1154,1180]" pageId="5" pageNumber="542" value="2008-07-19">
<collectingDate id="8609E9F0FFDCB033FE747BA1FD9BD8EC" box="[414,544,1154,1180]" pageId="5" pageNumber="542" value="2008-07-19">19.vii.2008</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FDC17BA0FD86D8ED" box="[555,573,1155,1181]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="529B3C85FFDCB033FDA37BA0FD69D8EC" box="[585,722,1154,1181]" collectionCode="SPMN" location="Ito" pageId="5" pageNumber="542" specimenCount="1" stateProvince="Fukui">
<location id="E72C6003FFDCB033FDA37BA0FDD3D8ED" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FDA37BA0FDD3D8ED" box="[585,616,1155,1181]" name="Ito" pageId="5" pageNumber="542" stateProvince="Fukui">Ito</location>
(
<collectionCode id="84E2AE1DFFDCB033FD9D7BA1FD71D8EC" box="[631,714,1154,1180]" pageId="5" pageNumber="542">SPMN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="529B3C85FFDCB033FD347BA1FE3ED8B1" collectingDate="2001-07-14" collectionCode="T" county="Abe-toge" location="Aoi-ku" municipality="Umegashima" pageId="5" pageNumber="542" specimenCount="2" specimenCount-female="2" stateProvince="Shizuoka">
<specimenCount id="F4F5FD51FFDCB033FD347BA1FCF6D8EC" box="[734,845,1154,1180]" pageId="5" pageNumber="542" type="female">2 females</specimenCount>
, 
<collectingCounty id="0B2D4E54FFDCB033FCBD7BA1FC79D8EC" box="[855,962,1154,1180]" pageId="5" pageNumber="542">Abe-toge</collectingCounty>
, 
<collectingMunicipality id="0228ACA2FFDCB033FC247BA1FBDED8EC" box="[974,1125,1154,1180]" pageId="5" pageNumber="542">Umegashima</collectingMunicipality>
, 
<location id="E72C6003FFDCB033FB857BA1FB7AD8EC" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FB857BA1FB7AD8EC" box="[1135,1217,1154,1180]" county="Abe-toge" municipality="Umegashima" name="Aoi-ku" pageId="5" pageNumber="542" stateProvince="Shizuoka">Aoi-ku</location>
, 
<location id="E72C6003FFDCB033FB277BA1FAD9D8EC" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FB277BA1FAD9D8EC" box="[1229,1378,1154,1180]" county="Abe-toge" municipality="Umegashima" name="Shizuoka-shi" pageId="5" pageNumber="542" stateProvince="Shizuoka">Shizuoka-shi</location>
, 
<collectingRegion id="2037F83AFFDCB033FA847BA1FF66D8B0" country="Japan" name="Shizuoka" pageId="5" pageNumber="542">Shizuoka</collectingRegion>
, 
<date id="964D1018FFDCB033FF0D7B85FED3D8B0" box="[231,360,1190,1216]" pageId="5" pageNumber="542" value="2001-07-14">
<collectingDate id="8609E9F0FFDCB033FF0D7B85FED3D8B0" box="[231,360,1190,1216]" pageId="5" pageNumber="542" value="2001-07-14">14.vii.2001</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FE997B84FE3ED8B1" box="[371,389,1191,1217]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="529B3C85FFDCB033FE657B85FDFDD8B0" box="[399,582,1190,1216]" collectionCode="SPMN" location="Hattori" pageId="5" pageNumber="542" specimenCount="1" stateProvince="Shizuoka">
<location id="E72C6003FFDCB033FE657B85FE65D8B0" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FE657B85FE65D8B0" box="[399,478,1190,1216]" name="Hattori" pageId="5" pageNumber="542" stateProvince="Shizuoka">Hattori</location>
(
<collectionCode id="84E2AE1DFFDCB033FE067B85FD84D8B0" box="[492,575,1190,1216]" pageId="5" pageNumber="542">SPMN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="529B3C85FFDCB033FDBA7B85FE95D894" collectingDate="2001-06-18" collectionCode="T" county="Abe-toge" location="Aoi-ku" municipality="Umegashima" pageId="5" pageNumber="542" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Shizuoka">
<specimenCount id="F4F5FD51FFDCB033FDBA7B85FD26D8B0" box="[592,669,1190,1216]" pageId="5" pageNumber="542" type="male">1 male</specimenCount>
, 
<specimenCount id="F4F5FD51FFDCB033FD4D7B85FCAED8B0" box="[679,789,1190,1216]" pageId="5" pageNumber="542" type="female">3 females</specimenCount>
, 
<collectingCounty id="0B2D4E54FFDCB033FCF47B85FC32D8B0" box="[798,905,1190,1216]" pageId="5" pageNumber="542">Abe-toge</collectingCounty>
, 
<collectingMunicipality id="0228ACA2FFDCB033FC797B85FB91D8B0" box="[915,1066,1190,1216]" pageId="5" pageNumber="542">Umegashima</collectingMunicipality>
, 
<location id="E72C6003FFDCB033FBD87B85FB3FD8B0" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FBD87B85FB3FD8B0" box="[1074,1156,1190,1216]" county="Abe-toge" municipality="Umegashima" name="Aoi-ku" pageId="5" pageNumber="542" stateProvince="Shizuoka">Aoi-ku</location>
, 
<location id="E72C6003FFDCB033FB647B85FA98D8B0" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FB647B85FA98D8B0" box="[1166,1315,1190,1216]" county="Abe-toge" municipality="Umegashima" name="Shizuoka-shi" pageId="5" pageNumber="542" stateProvince="Shizuoka">Shizuoka-shi</location>
, 
<collectingRegion id="2037F83AFFDCB033FAC77B85FA23D8B0" box="[1325,1432,1190,1216]" country="Japan" name="Shizuoka" pageId="5" pageNumber="542">Shizuoka</collectingRegion>
, 
<date id="964D1018FFDCB033FF7D7BE9FEA9D894" box="[151,274,1226,1252]" pageId="5" pageNumber="542" value="2001-06-18">
<collectingDate id="8609E9F0FFDCB033FF7D7BE9FEA9D894" box="[151,274,1226,1252]" pageId="5" pageNumber="542" value="2001-06-18">18.vi.2001</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FEF67BE9FE95D894" box="[284,302,1226,1252]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="529B3C85FFDCB033FED27BE9FE4BD894" box="[312,496,1226,1252]" collectionCode="SPMN" location="Hattori" pageId="5" pageNumber="542" specimenCount="1" stateProvince="Shizuoka">
<location id="E72C6003FFDCB033FED27BE9FE3CD894" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FED27BE9FE3CD894" box="[312,391,1226,1252]" name="Hattori" pageId="5" pageNumber="542" stateProvince="Shizuoka">Hattori</location>
(
<collectionCode id="84E2AE1DFFDCB033FE7C7BE9FE52D894" box="[406,489,1226,1252]" pageId="5" pageNumber="542">SPMN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="529B3C85FFDCB033FE117BE9FADBD894" box="[507,1376,1226,1252]" collectingDate="1999-06-06" collectionCode="T" county="Abe-toge" location="Aoi-ku" municipality="Umegashima" pageId="5" pageNumber="542" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka">
<specimenCount id="F4F5FD51FFDCB033FE117BE9FDF3D894" box="[507,584,1226,1252]" pageId="5" pageNumber="542" type="male">1 male</specimenCount>
, 
<collectingCounty id="0B2D4E54FFDCB033FDBB7BE9FD07D894" box="[593,700,1226,1252]" pageId="5" pageNumber="542">Abe-toge</collectingCounty>
, 
<collectingMunicipality id="0228ACA2FFDCB033FD2D7BE9FCE5D894" box="[711,862,1226,1252]" pageId="5" pageNumber="542">Umegashima</collectingMunicipality>
, 
<location id="E72C6003FFDCB033FC8C7BE9FC03D894" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FC8C7BE9FC03D894" box="[870,952,1226,1252]" county="Abe-toge" municipality="Umegashima" name="Aoi-ku" pageId="5" pageNumber="542" stateProvince="Shizuoka">Aoi-ku</location>
, 
<location id="E72C6003FFDCB033FC297BE9FBE3D894" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FC297BE9FBE3D894" box="[963,1112,1226,1252]" county="Abe-toge" municipality="Umegashima" name="Shizuoka-shi" pageId="5" pageNumber="542" stateProvince="Shizuoka">Shizuoka-shi</location>
, 
<collectingRegion id="2037F83AFFDCB033FB887BE9FB76D894" box="[1122,1229,1226,1252]" country="Japan" name="Shizuoka" pageId="5" pageNumber="542">Shizuoka</collectingRegion>
, 
<date id="964D1018FFDCB033FB327BE9FAFFD894" box="[1240,1348,1226,1252]" pageId="5" pageNumber="542" value="1999-06-06">
<collectingDate id="8609E9F0FFDCB033FB327BE9FAFFD894" box="[1240,1348,1226,1252]" pageId="5" pageNumber="542" value="1999-06-06">6.vi.1999</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FAA47BE9FADBD894" box="[1358,1376,1226,1252]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="529B3C85FFDCB033FA807BE9FE93D978" collectionCode="SPMN" location="Hattori" pageId="5" pageNumber="542" specimenCount="1" stateProvince="Shizuoka">
<location id="E72C6003FFDCB033FA807BE9FF05D977" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FA807BE9FF05D977" name="Hattori" pageId="5" pageNumber="542" stateProvince="Shizuoka">Hattori</location>
(
<collectionCode id="84E2AE1DFFDCB033FF247BCDFE9AD978" box="[206,289,1262,1288]" pageId="5" pageNumber="542">SPMN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="529B3C85FFDCB033FEDF7BCDFB06D978" box="[309,1213,1261,1288]" collectingDate="1997-07-21" collectionCode="T" county="Abe-toge" location="Aoi-ku" municipality="Umegashima" pageId="5" pageNumber="542" specimenCount="1" specimenCount-male="1" stateProvince="Shizuoka">
<specimenCount id="F4F5FD51FFDCB033FEDF7BCDFE38D977" box="[309,387,1261,1288]" pageId="5" pageNumber="542" type="male">1 male</specimenCount>
, 
<collectingCounty id="0B2D4E54FFDCB033FE647BCEFE42D977" box="[398,505,1261,1287]" pageId="5" pageNumber="542">Abe-toge</collectingCounty>
, 
<collectingMunicipality id="0228ACA2FFDCB033FDEF7BCEFD27D977" box="[517,668,1261,1287]" pageId="5" pageNumber="542">Umegashima</collectingMunicipality>
, 
<location id="E72C6003FFDCB033FD4C7BCEFD43D977" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FD4C7BCEFD43D977" box="[678,760,1261,1287]" county="Abe-toge" municipality="Umegashima" name="Aoi-ku" pageId="5" pageNumber="542" stateProvince="Shizuoka">Aoi-ku</location>
, 
<location id="E72C6003FFDCB033FCEE7BCEFC22D977" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FCEE7BCEFC22D977" box="[772,921,1261,1287]" county="Abe-toge" municipality="Umegashima" name="Shizuoka-shi" pageId="5" pageNumber="542" stateProvince="Shizuoka">Shizuoka-shi</location>
, 
<collectingRegion id="2037F83AFFDCB033FC4C7BCEFBAAD977" box="[934,1041,1261,1287]" country="Japan" name="Shizuoka" pageId="5" pageNumber="542">Shizuoka</collectingRegion>
, 
<date id="964D1018FFDCB033FBF77BCEFB24D977" box="[1053,1183,1261,1287]" pageId="5" pageNumber="542" value="1997-07-21">
<collectingDate id="8609E9F0FFDCB033FBF77BCEFB24D977" box="[1053,1183,1261,1287]" pageId="5" pageNumber="542" value="1997-07-21">21.vii.1997</collectingDate>
</date>
, 
<collectionCode id="84E2AE1DFFDCB033FB417BCDFB06D978" box="[1195,1213,1262,1288]" name="Tavera, Department of Geology and Geophysics" pageId="5" pageNumber="542">T</collectionCode>
</materialsCitation>
. 
<materialsCitation id="529B3C85FFDCB033FB227BCEFA39D978" box="[1224,1410,1261,1288]" collectionCode="SPMN" location="Hattori" pageId="5" pageNumber="542" specimenCount="1" stateProvince="Shizuoka">
<location id="E72C6003FFDCB033FB227BCEFAACD977" LSID="urn:lsid:plazi:treatment:6A5A87CEFFDAB030FF7D7A9CFD20DCC0:E72C6003FFDCB033FB227BCEFAACD977" box="[1224,1303,1261,1287]" name="Hattori" pageId="5" pageNumber="542" stateProvince="Shizuoka">Hattori</location>
(
<collectionCode id="84E2AE1DFFDCB033FAC27BCDFAC0D978" box="[1320,1403,1262,1288]" pageId="5" pageNumber="542">SPMN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="529B3C85FFDCB033FA647BCDFBB4D95C" collectingDate="1994-06-11" collectionCode="T" county="Kawamata-gawa" location="Hokuto-shi" municipality="Ohizumi-cho" pageId="5" pageNumber="542" specimenCount="2" specimenCount-female="2" stateProvince="Yamanashi">
<specimenCount id="F4F5FD51FFDCB033FA647BCDFF4AD95B" pageId="5" pageNumber="542" type="female">2 females</specimenCount>
, 
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</paragraph>
</subSubSection>
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<emphasis id="D087EACAFFDCB033FF2D792EFEF3DA57" bold="true" box="[199,328,1549,1575]" pageId="5" pageNumber="542">Etymology</emphasis>
. The new species is named in memory and honor of the late Mr. Toshio Hattori, a specialist of 
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taxonomy, especially 
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in 
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.
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<emphasis id="D087EACAFFDCB033FF2D7977FEE2DA1E" bold="true" box="[199,345,1620,1646]" pageId="5" pageNumber="542">Distribution</emphasis>
. 
<collectingCountry id="9AE47648FFDCB033FE8D7976FE1CDA1F" box="[359,423,1621,1647]" name="Japan" pageId="5" pageNumber="542">Japan</collectingCountry>
(Honshu).
</paragraph>
<paragraph id="E24C36D8FFDCB033FF2D795BFC8EDAAB" blockId="5.[151,1437,1083,2043]" pageId="5" pageNumber="542">
<emphasis id="D087EACAFFDCB033FF2D795BFE98DAE2" bold="true" box="[199,291,1656,1682]" pageId="5" pageNumber="542">Habitat</emphasis>
. The 
<typeStatus id="3D48887AFFDCB033FE8C795AFE2DDAE3" box="[358,406,1657,1683]" pageId="5" pageNumber="542">type</typeStatus>
locality is a small, narrow, mountain stream. The rivers at the other collecting sites varied from 
<quantity id="250B9B3DFFDCB033FF3C79BEFE96DAC7" box="[214,301,1693,1719]" metricMagnitude="0" metricUnit="m" metricValue="6.0" metricValueMax="10.0" metricValueMin="2.0" pageId="5" pageNumber="542" unit="m" value="6.0" valueMax="10.0" valueMin="2.0">2–10 m</quantity>
wide, with bottoms consisting of boulders of different sizes, pebbles, and leaf litter, and with steep banks. Water was clear, colorless, transparent, and odorless.
</paragraph>
</subSubSection>
<subSubSection id="AAE96553FFDCB030FF2D79C6FD20DCC0" lastPageId="6" lastPageNumber="543" pageId="5" pageNumber="542" type="discussion">
<paragraph id="E24C36D8FFDCB033FF2D79C6FBDCDB8B" blockId="5.[151,1437,1083,2043]" pageId="5" pageNumber="542">
<emphasis id="D087EACAFFDCB033FF2D79C6FE8FDA8F" bold="true" box="[199,308,1765,1791]" pageId="5" pageNumber="542">Remarks</emphasis>
. This species is determined to be the same species as “ 
<taxonomicName id="25F34D5BFFDCB033FC7479C5FC4FDA8F" authorityName="Tsuda" authorityYear="1942" box="[926,1012,1765,1791]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDCB033FC7479C5FC14DA8F" box="[926,943,1766,1791]" italics="true" pageId="5" pageNumber="542">R</emphasis>
. 
<emphasis id="D087EACAFFDCB033FC5479C6FC4FDA8F" box="[958,1012,1765,1791]" italics="true" pageId="5" pageNumber="542">yukii</emphasis>
</taxonomicName>
” figured by 
<bibRefCitation id="86624B29FFDCB033FB6979C6FA90DA8F" author="Schmid, F." box="[1155,1323,1765,1791]" pageId="5" pageNumber="542" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid (1970)</bibRefCitation>
(
<figureCitation id="7AC82A5DFFDCB033FAD079C6FF0CDB53" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Figs 2K, 2L</figureCitation>
) because the morphology of the male genitalia (
<figureCitation id="7AC82A5DFFDCB033FD09782AFCE4DB53" box="[739,863,1801,1827]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Figs 2A–F</figureCitation>
) is identical to that of the species. 
<bibRefCitation id="86624B29FFDCB033FB19782AFA27DB53" author="Schmid, F." box="[1267,1436,1801,1827]" pageId="5" pageNumber="542" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid (1970)</bibRefCitation>
misidentified and figured this species as 
<taxonomicName id="25F34D5BFFDCB033FD82780DFD04DB37" authorityName="Tsuda" authorityYear="1942" box="[616,703,1837,1863]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDCB033FD82780DFDC2DB37" box="[616,633,1838,1863]" italics="true" pageId="5" pageNumber="542">R</emphasis>
. 
<emphasis id="D087EACAFFDCB033FD63780EFD04DB37" box="[649,703,1837,1863]" italics="true" pageId="5" pageNumber="542">yukii</emphasis>
</taxonomicName>
and did not describe any features. From the original description of 
<taxonomicName id="25F34D5BFFDCB033FF5C7871FEB7DB1B" authorityName="Tsuda" authorityYear="1942" box="[182,268,1873,1899]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDCB033FF5C7871FF7CDB1B" box="[182,199,1874,1899]" italics="true" pageId="5" pageNumber="542">R</emphasis>
. 
<emphasis id="D087EACAFFDCB033FF3C7872FEB7DB1B" box="[214,268,1873,1899]" italics="true" pageId="5" pageNumber="542">yukii</emphasis>
</taxonomicName>
by 
<bibRefCitation id="86624B29FFDCB033FEDD7872FE75DB1B" author="Tsuda, M." box="[311,462,1873,1899]" pageId="5" pageNumber="542" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda (1942)</bibRefCitation>
, segment X is “approximately triangular in dorsal view, slightly advanced at the apex, rod-shaped in lateral view, tapered at the apex.” His figures (
<figureCitation id="7AC82A5DFFDCB033FCAE7856FC7EDBFF" box="[836,965,1909,1935]" captionStart="FIGURES 1" captionStartId="2.[151,275,1802,1828]" captionTargetBox="[253,1334,194,1767]" captionTargetId="figure-20@2.[239,1347,179,1778]" captionTargetPageId="2" captionText="FIGURES 1A−1I. Rhyacophila yukii Tsuda 1942, male genitalia (1A−1I): 1A, left lateral; 1B, segment X, dorsal; 1C, segment X, caudal (variation from same locality); 1D, ventral lobe of aedeagus, dorsal; 1E, segment IX and inferior appendages, ventral; 1F, phallic apparatus, left lateral; 1G, caudal; 1H, left lateral, by Tsuda (1942); 1I, dorsal, by Tsuda (1942). Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a. = apical segment of an inferior appendage (paired); b.i.a. = basal segment of an inferior appendage (paired); d.b.a.= dorsal branch of aedeagus; pht. = phallotheca; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057496" httpUri="https://zenodo.org/record/5057496/files/figure.png" pageId="5" pageNumber="542">Figs 1H, 1I</figureCitation>
) show that the morphology of segment X is clearly different from that of 
<taxonomicName id="25F34D5BFFDCB033FE1C78B9FDF0DBC3" authorityName="Tsuda" authorityYear="1942" box="[502,587,1945,1971]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDCB033FE1C78B9FDBCDBC3" box="[502,519,1946,1971]" italics="true" pageId="5" pageNumber="542">R</emphasis>
. 
<emphasis id="D087EACAFFDCB033FDFF78BAFDF0DBC3" box="[533,587,1945,1971]" italics="true" pageId="5" pageNumber="542">yukii</emphasis>
</taxonomicName>
figured by 
<bibRefCitation id="86624B29FFDCB033FD2778BAFCCFDBC3" author="Schmid, F." box="[717,884,1945,1971]" pageId="5" pageNumber="542" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid (1970)</bibRefCitation>
(
<figureCitation id="7AC82A5DFFDCB033FC6878BAFBB5DBC3" box="[898,1038,1945,1971]" captionStart="FIGURES 2" captionStartId="4.[151,264,1727,1751]" captionTargetBox="[183,1401,196,1691]" captionTargetId="figure-20@4.[170,1416,181,1703]" captionTargetPageId="4" captionText="FIGURES 2A−2L. Rhyacophila hattorii new species. 2A−2F, male genitalia: 2A, left lateral; 2B, dorsal; 2C, caudal; 2D, phal- lic apparatus, left lateral; 2E, ventral lobe of aedeagus, dorsal; 2F, inferior appendages, ventral. 2G−2J, female genitalia: 2G, segments VII and VIII, left lateral, with segments IX, X, and XI retracted into segments VII and VIII; 2H, segments VII−XI, left lateral; 2I, vaginal apparatus, ventral; 2J, vaginal apparatus, left lateral. 2K−2L, male genitalia by Schmid: 2K, left lateral, by Schmid (1970) and labelled as “R. yukii”; 2L, dorsal, by Schmid (1970) and labelled as “R. yukii”. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a = apical segment of an inferior appendage (paired); ap.r.VIII = apodemal rod from segment VIII (paired); ap.r.XI = apodemal rod from segment XI (paired); b.i.a. = basal segment of an inferior appendage (paired); c. = cercus (paired); pht. = phallotheca; p.pr = posterior process of vaginal apparatus; pr.sp. = processus spermathecae of vaginal apparatus; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057500" httpUri="https://zenodo.org/record/5057500/files/figure.png" pageId="5" pageNumber="542">Figs 2K, 2L</figureCitation>
). 
<bibRefCitation id="86624B29FFDCB033FBCB78BAFB00DBC4" author="Hattori, T." box="[1057,1211,1945,1972]" pageId="5" pageNumber="542" pagination="474 - 497" refId="ref7502" refString="Hattori, T. (2005) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations. Tokai University Press, Kanagawa, pp. 474 - 497. [in Japanese]" type="book chapter" year="2005">Hattori (2005</bibRefCitation>
, 
<bibRefCitation id="86624B29FFDCB033FB2C78BAFABBDBC3" author="Hattori, T." box="[1222,1280,1945,1971]" pageId="5" pageNumber="542" pagination="414 - 437" refId="ref7554" refString="Hattori, T. (2018) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations, 2 nd Edition. Tokai University Press, Kanagawa, pp. 414 - 437. [in Japanese]" type="book chapter" year="2018">2018</bibRefCitation>
) reported that the figures by 
<bibRefCitation id="86624B29FFDCB033FED3789EFE5ADBA7" author="Schmid, F." box="[313,481,1981,2007]" pageId="5" pageNumber="542" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid (1970)</bibRefCitation>
depict a species that is different from Tsuda’s 
<taxonomicName id="25F34D5BFFDCB033FC07789DFBF9DBA7" authorityName="Tsuda" authorityYear="1942" box="[1005,1090,1981,2007]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDCB033FC07789DFC45DBA7" box="[1005,1022,1982,2007]" italics="true" pageId="5" pageNumber="542">R</emphasis>
. 
<emphasis id="D087EACAFFDCB033FBE6789EFBF9DBA7" box="[1036,1090,1981,2007]" italics="true" pageId="5" pageNumber="542">yukii</emphasis>
</taxonomicName>
and that should be named as a new species. The new species name 
<taxonomicName id="25F34D5BFFDCB033FDC578C1FD19DB8B" authorityName="Torii &amp; Kobayashi" authorityYear="2021" box="[559,674,2017,2043]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="species" species="hattorii">
<emphasis id="D087EACAFFDCB033FDC578C1FDFBDB8B" box="[559,576,2018,2043]" italics="true" pageId="5" pageNumber="542">R</emphasis>
. 
<emphasis id="D087EACAFFDCB033FDA478C2FD19DB8B" box="[590,674,2017,2043]" italics="true" pageId="5" pageNumber="542">hattorii</emphasis>
</taxonomicName>
is given to “ 
<taxonomicName id="25F34D5BFFDCB033FCDB78C1FC3DDB8B" authorityName="Tsuda" authorityYear="1942" box="[817,902,2017,2043]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="542" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDCB033FCDB78C1FCF9DB8B" box="[817,834,2018,2043]" italics="true" pageId="5" pageNumber="542">R</emphasis>
. 
<emphasis id="D087EACAFFDCB033FCBA78C2FC3DDB8B" box="[848,902,2017,2043]" italics="true" pageId="5" pageNumber="542">yukii</emphasis>
</taxonomicName>
” of 
<bibRefCitation id="86624B29FFDCB033FC5278C2FBD8DB8B" author="Schmid, F." box="[952,1123,2017,2043]" pageId="5" pageNumber="542" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid (1970)</bibRefCitation>
.
</paragraph>
<paragraph id="E24C36D8FFDFB030FF2D7FB5FD20DCC0" blockId="6.[199,667,150,177]" box="[199,667,150,177]" pageId="6" pageNumber="543">
<emphasis id="D087EACAFFDFB030FF2D7FB5FEC7DCC0" bold="true" box="[199,380,150,176]" pageId="6" pageNumber="543">Japanese name</emphasis>
. Hattori-nagare-tobikera.
</paragraph>
</subSubSection>
</treatment>
</document>