treatments-xml/data/6A/5A/87/6A5A87CEFFD8B035FF7D781FFB38D909.xml
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<document id="8FC96995ABA0E1280905BF669B33240E" ID-DOI="10.11646/zootaxa.4995.3.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5057494" ID-ZooBank="8494CF5A-7C7A-42B7-8711-C87564D4B7E2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625174785369" checkinUser="plazi" docAuthor="Torii, Takaaki &amp; Kobayashi, Tohru" docDate="2021" docId="6A5A87CEFFD8B035FF7D781FFB38D909" docLanguage="en" docName="zootaxa.4995.3.8.pdf" docOrigin="Zootaxa 4995 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhyacophila yukii Tsuda 1942" docType="treatment" docVersion="3" lastPageNumber="540" masterDocId="9663FFB6FFD9B036FFEA7F23FFBBDC70" masterDocTitle="Redescription of Rhyacophila yukii Tsuda 1942 and description of three new Rhyacophila species from Japan (Trichoptera: Rhyacophilidae)" masterLastPageNumber="550" masterPageNumber="537" pageNumber="538" updateTime="1698982374593" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FF6F10E1EA7AABD916C8B0EC30909C9B">Redescription of Rhyacophila yukii Tsuda 1942 and description of three new Rhyacophila species from Japan (Trichoptera: Rhyacophilidae)</mods:title>
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<mods:namePart id="081285073E62564B040202258C334D8A">Torii, Takaaki</mods:namePart>
<mods:affiliation id="EA45710E1B65119E8D9FE01D493D7D17">Laboratory of Molecular Reproductive Biology, Graduate Division of Nutritional and Environmental Sciences, University of Shizuoka, 52 - 1, Yada, Suruga-Ku, Shizuoka, 422 - 8526, Japan &amp; Idea Consultant Inc., 1334 - 5 Riemon, Yaizu City, Shizuoka, 425 - 0084, Japan</mods:affiliation>
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<mods:namePart id="9F494D502DCA043B21CF72D13D82CB7B">Kobayashi, Tohru</mods:namePart>
<mods:affiliation id="252390245102C4ABE58197AF966C807B">Laboratory of Molecular Reproductive Biology, Graduate Division of Nutritional and Environmental Sciences, University of Shizuoka, 52 - 1, Yada, Suruga-Ku, Shizuoka, 422 - 8526, Japan</mods:affiliation>
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<paragraph id="E24C36D8FFD8B037FF7D781FFDB2DB26" blockId="1.[151,521,1852,1915]" box="[151,521,1852,1879]" pageId="1" pageNumber="538">
<heading id="B90481B4FFD8B037FF7D781FFDB2DB26" bold="true" box="[151,521,1852,1879]" fontSize="11" level="1" pageId="1" pageNumber="538" reason="1">
<taxonomicName id="25F34D5BFFD8B037FF7D781FFDB2DB26" ID-CoL="4SP8S" authority="Tsuda 1942" authorityName="Tsuda" authorityYear="1942" box="[151,521,1852,1879]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="538" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFD8B037FF7D781FFDB2DB26" bold="true" box="[151,521,1852,1879]" pageId="1" pageNumber="538">
<emphasis id="D087EACAFFD8B037FF7D781FFECFDB27" bold="true" box="[151,372,1852,1879]" italics="true" pageId="1" pageNumber="538">Rhyacophila yukii</emphasis>
<bibRefCitation id="86624B29FFD8B037FE91781EFDB2DB26" author="Tsuda, M." box="[379,521,1852,1879]" pageId="1" pageNumber="538" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda 1942</bibRefCitation>
</emphasis>
</taxonomicName>
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</paragraph>
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<paragraph id="E24C36D8FFD8B037FF7D7842FEADDB0B" blockId="1.[151,521,1852,1915]" box="[151,278,1889,1915]" pageId="1" pageNumber="538">
<figureCitation id="7AC82A5DFFD8B037FF7D7842FEADDB0B" box="[151,278,1889,1915]" captionStart="FIGURES 1" captionStartId="2.[151,275,1802,1828]" captionTargetBox="[253,1334,194,1767]" captionTargetId="figure-20@2.[239,1347,179,1778]" captionTargetPageId="2" captionText="FIGURES 1A1I. Rhyacophila yukii Tsuda 1942, male genitalia (1A1I): 1A, left lateral; 1B, segment X, dorsal; 1C, segment X, caudal (variation from same locality); 1D, ventral lobe of aedeagus, dorsal; 1E, segment IX and inferior appendages, ventral; 1F, phallic apparatus, left lateral; 1G, caudal; 1H, left lateral, by Tsuda (1942); 1I, dorsal, by Tsuda (1942). Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a. = apical segment of an inferior appendage (paired); b.i.a. = basal segment of an inferior appendage (paired); d.b.a.= dorsal branch of aedeagus; pht. = phallotheca; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057496" httpUri="https://zenodo.org/record/5057496/files/figure.png" pageId="1" pageNumber="538">Figs 1A1I</figureCitation>
</paragraph>
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<paragraph id="E24C36D8FFD8B037FF7D7886FE2FDB8A" blockId="1.[151,1437,1957,2042]" pageId="1" pageNumber="538">
<taxonomicName id="25F34D5BFFD8B037FF7D7886FDDADBCD" ID-CoL="4SP8S" authority="Tsuda 1942, pp. 248 - 249" authorityName="Tsuda" authorityPageNumber="248 - 249" authorityYear="1942" box="[151,609,1957,1981]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="538" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFD8B037FF7D7886FEEBDBCD" box="[151,336,1957,1981]" italics="true" pageId="1" pageNumber="538">Rhyacophila yukii</emphasis>
<bibRefCitation id="86624B29FFD8B037FEB37886FE69DBCD" author="Tsuda, M." box="[345,466,1957,1981]" pageId="1" pageNumber="538" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda 1942</bibRefCitation>
, pp. 248249
</taxonomicName>
, fig. 7, male;
<bibRefCitation id="86624B29FFD8B037FD197886FCC3DBCD" author="Hattori, T." box="[755,888,1957,1981]" pageId="1" pageNumber="538" pagination="474 - 497" refId="ref7502" refString="Hattori, T. (2005) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations. Tokai University Press, Kanagawa, pp. 474 - 497. [in Japanese]" type="book chapter" year="2005">Hattori 2005</bibRefCitation>
, pp. 427, 435, fig. 6, male from original description;
<bibRefCitation id="86624B29FFD8B037FF2D78E7FE01DBAC" author="Inazu, K. &amp; Nishida, A." box="[199,442,1988,2012]" pageId="1" pageNumber="538" pagination="169 - 212" refId="ref7610" refString="Inazu, K. &amp; Nishida, A. (2011) Trichoptera. In: Editorial office of Hyogo Fresh Water Biology (Ed.), Hyogo no kawa no ikimono zukan. Hyogo Rikusui-seibutsu Kenkyu-kai, Himeji, Hyogo, pp. 169 - 212. [in Japanese]" type="book chapter" year="2011">Inazu and Nishida 2011</bibRefCitation>
, p. 176, fig. 20, male;
<bibRefCitation id="86624B29FFD8B037FD7478E7FCD9DBAC" author="Kawase, N. &amp; Morita, H. &amp; Takeda, S. &amp; Uenishi, M." box="[670,866,1988,2012]" pageId="1" pageNumber="538" pagination="21 - 44" refId="ref7826" refString="Kawase, N., Morita, H., Takeda, S., &amp; Uenishi, M. (2017) Caddisfly (Insecta: Trichoptera) fauna of the Lake Biwa Basin in Shiga Prefecture, central Japan. Biology of Inland Waters, 31, 21 - 44. [in Japanese with English abstract]" type="journal article" year="2017">Kawase et al. 2017</bibRefCitation>
, p. 25, list;
<bibRefCitation id="86624B29FFD8B037FC3278E7FBE0DBAC" author="Hattori, T." box="[984,1115,1988,2012]" pageId="1" pageNumber="538" pagination="414 - 437" refId="ref7554" refString="Hattori, T. (2018) Rhyacophilidae. In: Kawai, T. &amp; Tanida, K. (Eds.), Aquatic Insects of Japan: Manual with Keys and Illustrations, 2 nd Edition. Tokai University Press, Kanagawa, pp. 414 - 437. [in Japanese]" type="book chapter" year="2018">Hattori 2018</bibRefCitation>
, pp. 487, 495, fig. 5, male from original description.
</paragraph>
</subSubSection>
<caption id="B68C6650FFDBB034FF7D7829FDF0DBA8" ID-DOI="http://doi.org/10.5281/zenodo.5057496" ID-Zenodo-Dep="5057496" httpUri="https://zenodo.org/record/5057496/files/figure.png" pageId="2" pageNumber="539" startId="2.[151,275,1802,1828]" targetBox="[253,1334,194,1767]" targetPageId="2">
<paragraph id="E24C36D8FFDBB034FF7D7829FDF0DBA8" blockId="2.[151,1437,1802,2008]" pageId="2" pageNumber="539">
<emphasis id="D087EACAFFDBB034FF7D7829FEDDDB54" bold="true" box="[151,358,1802,1828]" pageId="2" pageNumber="539">FIGURES 1A1I</emphasis>
.
<taxonomicName id="25F34D5BFFDBB034FE9E7828FD77DB55" authority="Tsuda 1942" authorityName="Tsuda" authorityYear="1942" box="[372,716,1803,1829]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="539" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDBB034FE9E7828FD84DB55" box="[372,575,1803,1829]" italics="true" pageId="2" pageNumber="539">Rhyacophila yukii</emphasis>
<bibRefCitation id="86624B29FFDBB034FDAD7828FD77DB55" author="Tsuda, M." box="[583,716,1803,1829]" pageId="2" pageNumber="539" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda 1942</bibRefCitation>
</taxonomicName>
, male genitalia (1A1I): 1A, left lateral; 1B, segment X, dorsal; 1C, segment X, caudal (variation from same locality); 1D, ventral lobe of aedeagus, dorsal; 1E, segment IX and inferior appendages, ventral; 1F, phallic apparatus, left lateral; 1G, caudal; 1H, left lateral, by
<bibRefCitation id="86624B29FFDBB034FC067877FBCDDB1D" author="Tsuda, M." box="[1004,1142,1876,1901]" pageId="2" pageNumber="539" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda (1942)</bibRefCitation>
; 1I, dorsal, by
<bibRefCitation id="86624B29FFDBB034FAE57877FA22DB1D" author="Tsuda, M." box="[1295,1433,1876,1901]" pageId="2" pageNumber="539" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda (1942)</bibRefCitation>
. Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a. = apical segment of an inferior appendage (paired); b.i.a. = basal segment of an inferior appendage (paired); d.b.a.= dorsal branch of aedeagus; pht. = phallotheca; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus.
</paragraph>
</caption>
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<materialsCitation id="529B3C85FFDAB035FF7D7FB4FEF3DC89" collectingDate="2008-05-11" collectingDateMax="2014-05-17" collectingDateMin="2008-05-11" collectionCode="SEHU, CBM, NKW" collectorName="H. Morita &amp; Shiratani &amp; Makino-cho &amp; Takashima-shi &amp; S. Takeda" county="Ojigahata" elevation="654" latitude="35.203" location="Inukami-gun" longLatPrecision="6" longitude="136.4108" municipality="Taga-cho" pageId="3" pageNumber="540" specimenCount="23" specimenCount-male="23" stateProvince="Shiga">
Material.
<specimenCount id="F4F5FD51FFDAB035FEED7FB4FED4DCC1" box="[263,367,151,177]" pageId="3" pageNumber="540" type="male">11 males</specimenCount>
,
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,
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,
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,
<collectingRegion id="2037F83AFFDAB035FCFE7FB4FCEFDCC1" box="[788,852,151,177]" country="Japan" name="Shiga" pageId="3" pageNumber="540">Shiga</collectingRegion>
(
<geoCoordinate id="87C7501FFFDAB035FC8F7FB5FC58DCC1" box="[869,995,150,177]" degrees="35.2030" direction="north" orientation="latitude" pageId="3" pageNumber="540" precision="5" value="35.203">35.2030°N</geoCoordinate>
,
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<quantity id="250B9B3DFFDAB035FB6D7FB5FB6ADCC1" box="[1159,1233,150,177]" metricMagnitude="2" metricUnit="m" metricValue="6.54" pageId="3" pageNumber="540" unit="m" value="654.0">654 m</quantity>
a.s.l.
</elevation>
),
<date id="964D1018FFDAB035FAC27FB4FA2CDCC1" box="[1320,1431,151,177]" pageId="3" pageNumber="540" value="2008-05-11">
<collectingDate id="8609E9F0FFDAB035FAC27FB4FA2CDCC1" box="[1320,1431,151,177]" pageId="3" pageNumber="540" value="2008-05-11">11.v.2008</collectingDate>
</date>
,
<collectorName id="4F06530EFFDAB035FF7D7F98FEBCDCA5" box="[151,263,187,213]" pageId="3" pageNumber="540">H. Morita</collectorName>
(
<specimenCount id="F4F5FD51FFDAB035FEF37F98FECDDCA5" box="[281,374,187,213]" pageId="3" pageNumber="540" type="male">2 males</specimenCount>
:
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,
<specimenCount id="F4F5FD51FFDAB035FE317F98FD83DCA5" box="[475,568,187,213]" pageId="3" pageNumber="540" type="male">9 males</specimenCount>
:
<collectionCode id="84E2AE1DFFDAB035FDAE7F98FD3CDCA5" box="[580,647,187,213]" country="Japan" lsid="urn:lsid:biocol.org:col:13737" name="Natural History Museum and Institute" pageId="3" pageNumber="540" type="Herbarium">CBM</collectionCode>
);
<specimenCount id="F4F5FD51FFDAB035FD767F98FD50DCA5" box="[668,747,187,213]" pageId="3" pageNumber="540" type="male">1 male</specimenCount>
,
<collectorName id="4F06530EFFDAB035FD137F98FCDADCA5" box="[761,865,187,213]" pageId="3" pageNumber="540">Shiratani</collectorName>
,
<collectorName id="4F06530EFFDAB035FC877F98FC42DCA5" box="[877,1017,187,213]" pageId="3" pageNumber="540">Makino-cho</collectorName>
,
<collectorName id="4F06530EFFDAB035FBEC7F98FB17DCA5" box="[1030,1196,187,213]" pageId="3" pageNumber="540">Takashima-shi</collectorName>
,
<collectingRegion id="2037F83AFFDAB035FB527F98FB40DCA5" box="[1208,1275,187,213]" country="Japan" name="Shiga" pageId="3" pageNumber="540">Shiga</collectingRegion>
,
<date id="964D1018FFDAB035FAE27F98FAC3DCA5" box="[1288,1400,187,213]" pageId="3" pageNumber="540" value="2014-05-17">
<collectingDate id="8609E9F0FFDAB035FAE27F98FAC3DCA5" box="[1288,1400,187,213]" pageId="3" pageNumber="540" value="2014-05-17">17.v.2014</collectingDate>
</date>
,
<collectorName id="4F06530EFFDAB035FA6C7F98FF5CDC89" pageId="3" pageNumber="540">S. Takeda</collectorName>
(
<collectionCode id="84E2AE1DFFDAB035FF1D7FFCFEFBDC89" box="[247,320,223,249]" pageId="3" pageNumber="540">NKW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
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<emphasis id="D087EACAFFDAB035FF2D7E20FEBFDD6D" bold="true" box="[199,260,259,285]" pageId="3" pageNumber="540">Male</emphasis>
. Length of each forewing 6.58.0 mm (n = 13); specimens in alcohol with head, thorax, and wings dark brown; legs yellowish brown, with spurs dark brown.
</paragraph>
<paragraph id="E24C36D8FFDAB035FF2D7E68FBDBDED9" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="540">
<emphasis id="D087EACAFFDAB035FF2D7E68FEC9DD14" bold="true" box="[199,370,330,357]" pageId="3" pageNumber="540">Male genitalia</emphasis>
(
<figureCitation id="7AC82A5DFFDAB035FE6A7E68FDBFDD15" box="[384,516,331,357]" captionStart="FIGURES 1" captionStartId="2.[151,275,1802,1828]" captionTargetBox="[253,1334,194,1767]" captionTargetId="figure-20@2.[239,1347,179,1778]" captionTargetPageId="2" captionText="FIGURES 1A1I. Rhyacophila yukii Tsuda 1942, male genitalia (1A1I): 1A, left lateral; 1B, segment X, dorsal; 1C, segment X, caudal (variation from same locality); 1D, ventral lobe of aedeagus, dorsal; 1E, segment IX and inferior appendages, ventral; 1F, phallic apparatus, left lateral; 1G, caudal; 1H, left lateral, by Tsuda (1942); 1I, dorsal, by Tsuda (1942). Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a. = apical segment of an inferior appendage (paired); b.i.a. = basal segment of an inferior appendage (paired); d.b.a.= dorsal branch of aedeagus; pht. = phallotheca; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057496" httpUri="https://zenodo.org/record/5057496/files/figure.png" pageId="3" pageNumber="540">Figs 1A1I</figureCitation>
). Segment IX in lateral view subrectangular, with upper portion longer than lower portion; rectangular in dorsal view, twice as wide as long. Segment X in lateral view rod-shaped, tapered and slightly hooked ventrad at tip, variably trapezoid to triangular in dorsal view (
<figureCitation id="7AC82A5DFFDAB035FBFB7EB0FB26DDDD" box="[1041,1181,403,429]" captionStart="FIGURES 1" captionStartId="2.[151,275,1802,1828]" captionTargetBox="[253,1334,194,1767]" captionTargetId="figure-20@2.[239,1347,179,1778]" captionTargetPageId="2" captionText="FIGURES 1A1I. Rhyacophila yukii Tsuda 1942, male genitalia (1A1I): 1A, left lateral; 1B, segment X, dorsal; 1C, segment X, caudal (variation from same locality); 1D, ventral lobe of aedeagus, dorsal; 1E, segment IX and inferior appendages, ventral; 1F, phallic apparatus, left lateral; 1G, caudal; 1H, left lateral, by Tsuda (1942); 1I, dorsal, by Tsuda (1942). Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a. = apical segment of an inferior appendage (paired); b.i.a. = basal segment of an inferior appendage (paired); d.b.a.= dorsal branch of aedeagus; pht. = phallotheca; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057496" httpUri="https://zenodo.org/record/5057496/files/figure.png" pageId="3" pageNumber="540">Figs 1B, 1C</figureCitation>
), approximately triangular in diagonally above of caudal view, slightly bends downward at apex, 1.21.3 times as wide as long. Apical band (a.b.) in lateral view long and narrow, bar shaped. Dorsal branch of aedeagus (d.b.a.) in lateral view slightly protruding. Ventral branch of aedeagus (v.b.a.) in lateral view cylindrical and protruding. Aedeagus (aed.) in lateral view broadly curved 90° like long and narrow spine. Ventral lobe of aedeagus (v.l.a.) elongate subelliptical in dorsal view. Basal segment of each inferior appendage (b.i.a.) nearly rectangular in lateral view, in ventral view with its mesal margin concave and lateral margin straight; apical segment of each inferior appendage (a.i.a.) in lateral view with apex depressed obliquely, in ventral view with mesal and lateral margins straight.
</paragraph>
<paragraph id="E24C36D8FFDAB035FF2D7D90FE19DEBD" blockId="3.[151,1437,150,1401]" box="[199,418,691,717]" pageId="3" pageNumber="540">
<emphasis id="D087EACAFFDAB035FF2D7D90FE9EDEBD" bold="true" box="[199,293,691,717]" pageId="3" pageNumber="540">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="E24C36D8FFDAB035FF2D7DF4FDA8DE81" blockId="3.[151,1437,150,1401]" box="[199,531,727,753]" pageId="3" pageNumber="540">
<emphasis id="D087EACAFFDAB035FF2D7DF4FE34DE81" bold="true" box="[199,399,727,753]" pageId="3" pageNumber="540">Immature stages</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="AAE96553FFDAB035FF2D7DD9FD9BDF66" box="[199,544,762,790]" pageId="3" pageNumber="540" type="distribution">
<paragraph id="E24C36D8FFDAB035FF2D7DD9FD9BDF66" blockId="3.[151,1437,150,1401]" box="[199,544,762,790]" pageId="3" pageNumber="540">
<emphasis id="D087EACAFFDAB035FF2D7DD9FEE2DF64" bold="true" box="[199,345,762,788]" pageId="3" pageNumber="540">Distribution</emphasis>
.
<collectingCountry id="9AE47648FFDAB035FE8D7DD8FE1CDF65" box="[359,423,763,789]" name="Japan" pageId="3" pageNumber="540">Japan</collectingCountry>
(Honshu).
</paragraph>
</subSubSection>
<subSubSection id="AAE96553FFDAB035FF2D7C3CFB38D909" pageId="3" pageNumber="540" type="discussion">
<paragraph id="E24C36D8FFDAB035FF2D7C3CFD28D829" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="540">
<emphasis id="D087EACAFFDAB035FF2D7C3CFE80DF49" bold="true" box="[199,315,799,825]" pageId="3" pageNumber="540">Remarks.</emphasis>
This species belongs in the
<taxonomicName id="25F34D5BFFDAB035FD967C3CFCDCDF49" authorityName="R McLachlan" authorityYear="1879" box="[636,871,799,825]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="sibirica">
<emphasis id="D087EACAFFDAB035FD967C3CFCDCDF49" box="[636,871,799,825]" italics="true" pageId="3" pageNumber="540">Rhyacophila sibirica</emphasis>
</taxonomicName>
Group of
<bibRefCitation id="86624B29FFDAB035FC347C3CFB32DF49" author="Schmid, F." box="[990,1161,799,825]" pageId="3" pageNumber="540" pagination="5 - 230" refId="ref8745" refString="Schmid, F. (1970) Le genre Rhyacophila et la famille des Rhyacophilidae (Trichoptera). Memoires de la Societe Entomologique du Canada, 66, 5 - 230, pls. 1 - 52. https: // doi. org / 10.4039 / entm 10266 fv" type="journal article" year="1970">Schmid (1970)</bibRefCitation>
.
<bibRefCitation id="86624B29FFDAB035FB7E7C3CFA92DF49" author="Tsuda, M." box="[1172,1321,799,825]" pageId="3" pageNumber="540" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda (1942)</bibRefCitation>
described it based on
<specimenCount id="F4F5FD51FFDAB035FEF57C60FE3CDF2D" box="[287,391,835,861]" pageId="3" pageNumber="540" type="male">one male</specimenCount>
collected from Ohmiya-gawa, Hiei-zan,
<collectingRegion id="2037F83AFFDAB035FC8A7C60FB9BDF2D" box="[864,1056,835,861]" country="Japan" name="Shiga" pageId="3" pageNumber="540">Shiga Prefecture</collectingRegion>
, central
<collectingCountry id="9AE47648FFDAB035FB697C60FB7DDF2D" box="[1155,1222,835,861]" name="Japan" pageId="3" pageNumber="540">Japan</collectingCountry>
. The specimens in this study are from “Ojigahata, Taga-cho, Inukami-gun,
<collectingRegion id="2037F83AFFDAB035FCE07C44FCF4DFF1" box="[778,847,871,897]" country="Japan" name="Shiga" pageId="3" pageNumber="540">Shiga</collectingRegion>
,” which is
<quantity id="250B9B3DFFDAB035FC227C45FBB5DFF1" box="[968,1038,870,897]" metricMagnitude="4" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="540" unit="km" value="56.0">56 km</quantity>
northeast in a straight line from the
<typeStatus id="3D48887AFFDAB035FF7D7CA8FF7CDFD5" box="[151,199,907,933]" pageId="3" pageNumber="540">type</typeStatus>
locality. His description and illustrations of segment X and other characters of the male genitalia (
<figureCitation id="7AC82A5DFFDAB035FAF97CA8FA28DFD5" box="[1299,1427,907,933]" captionStart="FIGURES 1" captionStartId="2.[151,275,1802,1828]" captionTargetBox="[253,1334,194,1767]" captionTargetId="figure-20@2.[239,1347,179,1778]" captionTargetPageId="2" captionText="FIGURES 1A1I. Rhyacophila yukii Tsuda 1942, male genitalia (1A1I): 1A, left lateral; 1B, segment X, dorsal; 1C, segment X, caudal (variation from same locality); 1D, ventral lobe of aedeagus, dorsal; 1E, segment IX and inferior appendages, ventral; 1F, phallic apparatus, left lateral; 1G, caudal; 1H, left lateral, by Tsuda (1942); 1I, dorsal, by Tsuda (1942). Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a. = apical segment of an inferior appendage (paired); b.i.a. = basal segment of an inferior appendage (paired); d.b.a.= dorsal branch of aedeagus; pht. = phallotheca; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057496" httpUri="https://zenodo.org/record/5057496/files/figure.png" pageId="3" pageNumber="540">Figs 1H, 1I</figureCitation>
) agree with those of males examined in this study (
<figureCitation id="7AC82A5DFFDAB035FD2C7C8CFC3BDFB9" box="[710,896,943,969]" captionStart="FIGURES 1" captionStartId="2.[151,275,1802,1828]" captionTargetBox="[253,1334,194,1767]" captionTargetId="figure-20@2.[239,1347,179,1778]" captionTargetPageId="2" captionText="FIGURES 1A1I. Rhyacophila yukii Tsuda 1942, male genitalia (1A1I): 1A, left lateral; 1B, segment X, dorsal; 1C, segment X, caudal (variation from same locality); 1D, ventral lobe of aedeagus, dorsal; 1E, segment IX and inferior appendages, ventral; 1F, phallic apparatus, left lateral; 1G, caudal; 1H, left lateral, by Tsuda (1942); 1I, dorsal, by Tsuda (1942). Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a. = apical segment of an inferior appendage (paired); b.i.a. = basal segment of an inferior appendage (paired); d.b.a.= dorsal branch of aedeagus; pht. = phallotheca; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057496" httpUri="https://zenodo.org/record/5057496/files/figure.png" pageId="3" pageNumber="540">Figs 1A, 1B, 1C</figureCitation>
).
<bibRefCitation id="86624B29FFDAB035FC7E7C8CFB93DFB9" author="Tsuda, M." box="[916,1064,943,969]" pageId="3" pageNumber="540" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda (1942)</bibRefCitation>
described segment X of his male as approximately triangular and tapering to apex in dorsal view. Segment X of our specimens are triangular to trapezoidal in dorsal view but appears to be triangular and directed diagonally upward in caudal view; the morphology of segment X in dorsal view is slightly variable (
<figureCitation id="7AC82A5DFFDAB035FD5A7B38FCBDD845" box="[688,774,1051,1077]" captionStart="FIGURES 1" captionStartId="2.[151,275,1802,1828]" captionTargetBox="[253,1334,194,1767]" captionTargetId="figure-20@2.[239,1347,179,1778]" captionTargetPageId="2" captionText="FIGURES 1A1I. Rhyacophila yukii Tsuda 1942, male genitalia (1A1I): 1A, left lateral; 1B, segment X, dorsal; 1C, segment X, caudal (variation from same locality); 1D, ventral lobe of aedeagus, dorsal; 1E, segment IX and inferior appendages, ventral; 1F, phallic apparatus, left lateral; 1G, caudal; 1H, left lateral, by Tsuda (1942); 1I, dorsal, by Tsuda (1942). Abbreviations: a.b. = apical band; aed. = aedeagus; a.i.a. = apical segment of an inferior appendage (paired); b.i.a. = basal segment of an inferior appendage (paired); d.b.a.= dorsal branch of aedeagus; pht. = phallotheca; v.b.a. = ventral branch of aedeagus; v.l.a. = ventral lobe of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.5057496" httpUri="https://zenodo.org/record/5057496/files/figure.png" pageId="3" pageNumber="540">Fig. 1C</figureCitation>
). It is necessary to be careful about the angle of observation to observe the variety of shape of segment X.
</paragraph>
<paragraph id="E24C36D8FFDAB035FF2D7B40FC00D899" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="540">
<materialsCitation id="529B3C85FFDAB035FF2D7B40FC0CD899" county="Imperial University" location="The Tsuda" municipality="University" pageId="3" pageNumber="540" specimenCount="1" stateProvince="Kyoto" typeStatus="neotype">
<bibRefCitation id="86624B29FFDAB035FF2D7B40FEE7D80D" author="Tsuda, M." box="[199,348,1123,1149]" pageId="3" pageNumber="540" pagination="239 - 339" refId="ref9115" refString="Tsuda, M. (1942) Japanische Trichopteren I. Systematik. Memoirs of the College of Science, Kyoto Imperial University, Series B, 17, 239 - 339. [in German]" type="journal article" year="1942">Tsuda (1942)</bibRefCitation>
reported that the type specimen of
<taxonomicName id="25F34D5BFFDAB035FD007B40FC84D80D" authorityName="Tsuda" authorityYear="1942" box="[746,831,1123,1149]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDAB035FD007B40FD40D80C" box="[746,763,1123,1148]" italics="true" pageId="3" pageNumber="540">R</emphasis>
.
<emphasis id="D087EACAFFDAB035FCE37B40FC84D80D" box="[777,831,1123,1149]" italics="true" pageId="3" pageNumber="540">yukii</emphasis>
</taxonomicName>
was stored at
<collectingRegion id="2037F83AFFDAB035FC0B7B40FB9CD80D" box="[993,1063,1123,1149]" country="Japan" name="Kyoto" pageId="3" pageNumber="540">Kyoto</collectingRegion>
<collectingCounty id="0B2D4E54FFDAB035FBC47B40FAB0D80D" box="[1070,1291,1123,1149]" pageId="3" pageNumber="540">Imperial University</collectingCounty>
(
<collectingRegion id="2037F83AFFDAB035FAF07B40FAD9D80D" box="[1306,1378,1123,1149]" country="Japan" name="Kyoto" pageId="3" pageNumber="540">Kyoto</collectingRegion>
<collectingMunicipality id="0228ACA2FFDAB035FA837B40FF53D8D2" pageId="3" pageNumber="540">University</collectingMunicipality>
).
<location id="E72C6003FFDAB035FF107BA4FED5D8D1" LSID="urn:lsid:plazi:treatment:6A5A87CEFFD8B035FF7D781FFB38D909:E72C6003FFDAB035FF107BA4FED5D8D1" box="[250,366,1159,1185]" county="Imperial University" municipality="University" name="The Tsuda" pageId="3" pageNumber="540" stateProvince="Kyoto">The Tsuda</location>
collection was acquired later by the
<collectingRegion id="2037F83AFFDAB035FCE37BA4FCF4D8D1" box="[777,847,1159,1185]" country="Japan" name="Osaka" pageId="3" pageNumber="540">Osaka</collectingRegion>
<location id="E72C6003FFDAB035FCBC7BA4FB37D8D1" LSID="urn:lsid:plazi:treatment:6A5A87CEFFD8B035FF7D781FFB38D909:E72C6003FFDAB035FCBC7BA4FB37D8D1" box="[854,1164,1159,1185]" county="Imperial University" municipality="University" name="Museum of Natural History" pageId="3" pageNumber="540" stateProvince="Kyoto">Museum of Natural History</location>
, but now the type specimen of this species cannot be found (
<location id="E72C6003FFDAB035FDDC7B88FD3FD8B5" LSID="urn:lsid:plazi:treatment:6A5A87CEFFD8B035FF7D781FFB38D909:E72C6003FFDAB035FDDC7B88FD3FD8B5" box="[566,644,1195,1221]" county="Imperial University" municipality="University" name="Tanida" pageId="3" pageNumber="540" stateProvince="Kyoto">Tanida</location>
per. com.).
<location id="E72C6003FFDAB035FCE07B88FCF3D8B5" LSID="urn:lsid:plazi:treatment:6A5A87CEFFD8B035FF7D781FFB38D909:E72C6003FFDAB035FCE07B88FCF3D8B5" box="[778,840,1195,1221]" county="Imperial University" municipality="University" name="Since" pageId="3" pageNumber="540" stateProvince="Kyoto">Since</location>
it could not be confirmed whether the type specimen has completely disappeared, we do not specify
<typeStatus id="3D48887AFFDAB035FD427BECFCBAD899" box="[680,769,1231,1257]" pageId="3" pageNumber="540" type="neotype">neotype</typeStatus>
specimens here
</materialsCitation>
.
</paragraph>
<paragraph id="E24C36D8FFDAB035FF2D7BD0FB38D909" blockId="3.[151,1437,150,1401]" pageId="3" pageNumber="540">
In addition, examination of the specimen of
<taxonomicName id="25F34D5BFFDAB035FD557BD0FCAED97D" authorityName="Tsuda" authorityYear="1942" box="[703,789,1267,1293]" class="Insecta" family="Rhyacophilidae" genus="Rhyacophila" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="540" phylum="Arthropoda" rank="species" species="yukii">
<emphasis id="D087EACAFFDAB035FD557BD0FD6BD97C" box="[703,720,1267,1292]" italics="true" pageId="3" pageNumber="540">R</emphasis>
.
<emphasis id="D087EACAFFDAB035FD357BD0FCAED97D" box="[735,789,1267,1293]" italics="true" pageId="3" pageNumber="540">yukii</emphasis>
</taxonomicName>
reported from Mt. Sentsu-zan, Okuizumio-cho,
<collectingRegion id="2037F83AFFDAB035FAD07BD0FEB6D941" country="Japan" name="Shimane" pageId="3" pageNumber="540">Shimane Prefecture</collectingRegion>
, by
<bibRefCitation id="86624B29FFDAB035FED77A34FDC0D941" author="Kawase, N. &amp; Hayashi, M." box="[317,635,1303,1329]" pageId="3" pageNumber="540" pagination="83 - 86" refId="ref7780" refString="Kawase, N. &amp; Hayashi, M. (2010) Records of caddisflies from Mt. Sentsu-zan, Okuizumio-cho, Shimane Prefecture, Japan. Bulletin of Hoshizaki Green Foundation, 13, 83 - 86. [in Japanese with English abstract]" type="journal article" year="2010">Kawase and Hayashi (2010)</bibRefCitation>
showed that the shape of the tip of male segment X did not agree with the original description and their specimen possibly actually is an example of an undescribed species. Whether this specimen is an intraspecific variant or a different species still needs further consideration.
</paragraph>
</subSubSection>
</treatment>
</document>