254 lines
39 KiB
XML
254 lines
39 KiB
XML
<document id="43247100A9A36E45AC205AFA7059EC20" ID-CLB-Dataset="27622" ID-DOI="10.11646/zootaxa.4546.1.1" ID-GBIF-Dataset="aa93de4b-c9ed-4b3e-a1c2-3f5d90faba57" ID-ISSN="1175-5326" ID-Zenodo-Dep="2619020" ID-ZooBank="D2C9F49A-9B76-40AE-9A60-CAE9B99BA547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553964486275" checkinUser="plazi" docAuthor="Huber, Bernhard A. & Carvalho, Leonardo S." docDate="2019" docId="E21587DBFF8DFFC3FF11F8924FFEF9A8" docLanguage="en" docName="zootaxa.4546.1.1.pdf" docOrigin="Zootaxa 4546 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Arnapa tolire Huber & Carvalho 2019, sp. n." docType="treatment" docVersion="16" lastPageNumber="24" masterDocId="1E2CFFA3FF98FFD4FF86FFCD4B03FFA0" masterDocTitle="Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="22" updateTime="1698716013903" updateUser="ExternalLinkService">
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<mods:mods id="CF14B0A753D33F24A2F204B3B9F3C392" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="198D0FFC1B71EDE7CDC9E639CF885AA0">
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<mods:title id="AF49E4D0E8757F9947C7EE1056A571EB">Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae)</mods:title>
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<mods:namePart id="196BF56E8955EE22E9448B1874B351A1">Huber, Bernhard A.</mods:namePart>
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<mods:namePart id="1ACAD90CB5851DF6D8D6DF99F4A68C07">Carvalho, Leonardo S.</mods:namePart>
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<mods:title id="5879096717ED47518BDE9056E1777FF9">Zootaxa</mods:title>
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<mods:part id="A0BEB790F134680173E09CE7F58ED0BA">
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<mods:date id="27C68F49C0586FF65254045AEF073CFA">2019</mods:date>
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<mods:number id="A4089165EA07715B1436D66DBCDE77B7">2019-01-22</mods:number>
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<mods:number id="6CCA1BFFF5219D55D8933228829801CA">4546</mods:number>
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<mods:identifier id="665A1C2F2A4F4F0C0FFE8CEAC75B0884" type="DOI">10.11646/zootaxa.4546.1.1</mods:identifier>
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<treatment id="E21587DBFF8DFFC3FF11F8924FFEF9A8" ID-DOI="http://doi.org/10.5281/zenodo.5449654" ID-GBIF-Taxon="154531226" ID-Zenodo-Dep="5449654" LSID="urn:lsid:plazi:treatment:E21587DBFF8DFFC3FF11F8924FFEF9A8" httpUri="http://treatment.plazi.org/id/E21587DBFF8DFFC3FF11F8924FFEF9A8" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
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<subSubSection id="22A66546FF8DFFC1FF11F8924AD8F8DA" box="[151,475,1887,1914]" pageId="21" pageNumber="22" type="nomenclature">
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<paragraph id="6A0336CDFF8DFFC1FF11F8924AD8F8DA" blockId="21.[151,570,1887,1947]" box="[151,475,1887,1914]" pageId="21" pageNumber="22">
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<heading id="314B81A1FF8DFFC1FF11F8924AD8F8DA" bold="true" box="[151,475,1887,1914]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
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<emphasis id="58C8EADFFF8DFFC1FF11F8924AD8F8DA" bold="true" box="[151,475,1887,1914]" pageId="21" pageNumber="22">
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<taxonomicName id="ADBC4D4EFF8DFFC1FF11F8924A91F8DA" ID-CoL="GSX7" authority="Huber" authorityName="Huber & Carvalho" authorityYear="2019" box="[151,402,1887,1914]" class="Arachnida" family="Pholcidae" genus="Arnapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="tolire" status="sp. nov.">
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<emphasis id="58C8EADFFF8DFFC1FF11F8924A3BF8D9" bold="true" box="[151,312,1887,1913]" italics="true" pageId="21" pageNumber="22">Arnapa tolire</emphasis>
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Huber
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</taxonomicName>
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<taxonomicNameLabel id="43FB57A4FF8DFFC1FE1FF8AD4AD8F8DA" box="[409,475,1888,1914]" pageId="21" pageNumber="22" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="22A66546FF8DFFC1FF11F84F489EF840" pageId="21" pageNumber="22" type="description">
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<paragraph id="6A0336CDFF8DFFC1FF11F84F4939F83B" blockId="21.[151,570,1887,1947]" box="[151,570,1922,1947]" pageId="21" pageNumber="22">
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<figureCitation id="F2872A48FF8DFFC1FF11F84F4A14F83B" box="[151,279,1922,1947]" captionStart="FIGURES 39–44" captionStartId="16.[151,264,1927,1949]" captionTargetBox="[151,1436,589,1905]" captionTargetId="figure@16.[151,1436,589,1905]" captionTargetPageId="16" captionText="FIGURES 39–44. Arnapa spp., live specimens. 39–40. A. arfak sp. n., male and female from Mokwam. 41–42. A. tolire sp. n., male and female with egg sac from Ternate. 43–44. A. tinoor sp. n., male and female from near Tomohon." httpUri="https://zenodo.org/record/2619036/files/figure.png" pageId="21" pageNumber="22">Figs 41–42</figureCitation>
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,
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<figureCitation id="F2872A48FF8DFFC1FEA4F84F4A68F83B" box="[290,363,1922,1947]" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="21" pageNumber="22">60–64</figureCitation>
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,
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<figureCitation id="F2872A48FF8DFFC1FEF0F84E4A97F83B" box="[374,404,1923,1947]" captionStart="FIGURES 70–74" captionStartId="25.[151,264,1653,1675]" captionTargetBox="[154,1422,209,1620]" captionTargetId="figure@25.[151,1436,193,1632]" captionTargetPageId="25" captionText="FIGURES 70–74. Arnapa spp., cleared female genitalia, dorsal views. 70. A. arfak sp. n. (ZFMK, Ar 20605). 71. A. manokwari sp. n. (ZFMK, Ar 20608). 72. A. meja sp. n. (ZFMK, Ar 20610). 73. A. tolire sp. n. (ZFMK, Ar 20613). 74. A. tinoor sp. n. (ZFMK, Ar 20616). Scale lines: 0.3." httpUri="https://zenodo.org/record/2619048/files/figure.png" pageId="21" pageNumber="22">73</figureCitation>
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,
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<figureCitation id="F2872A48FF8DFFC1FE26F84F4AEAF83B" box="[416,489,1922,1947]" captionStart="FIGURES 81–89" captionStartId="27.[151,264,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@27.[151,1436,193,1705]" captionTargetPageId="27" captionText="FIGURES 81–89. Arnapa spp., epigyna, ventral views, and cleared female genitalia, ventral and dorsal views. 81–83. A. meja sp. n. (ZFMK, Ar 20610). 84–86. A. tolire sp. n. (ZFMK, Ar 20613). 87–89. A. tinoor sp. n. (ZFMK, Ar 20616). Scale lines: 0.5." httpUri="https://zenodo.org/record/2619052/files/figure.png" pageId="21" pageNumber="22">84–86</figureCitation>
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,
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<figureCitation id="F2872A48FF8DFFC1FE72F84E4939F83B" box="[500,570,1923,1947]" captionStart="FIGURES 90–97" captionStartId="28.[151,264,1665,1687]" captionTargetBox="[151,1436,193,1644]" captionTargetId="figure@28.[151,1436,193,1644]" captionTargetPageId="28" captionText="FIGURES 90–97. Arnapa tolire sp. n. (90–94, ZFMK, Ar 20613) and A. tinoor sp. n. (95–97, ZFMK, Ar 20615–16). 90–91. Female prosoma, frontal views. 92. Left female palp and chelicerae; note absence of stridulatory files. 93. Female palpal tarsal organ. 94–95. Epigyna, ventral views. 96–97. Female and male spinnerets. ALS, anterior lateral spinneret; ch, chelicera; PMS, posterior median spinneret; pto, palpal tarsal organ. Scale lines: 10 µm (93, 96–97), 100 µm (90–92, 94–95)." httpUri="https://zenodo.org/record/2619054/files/figure.png" pageId="21" pageNumber="22">90–94</figureCitation>
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</paragraph>
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<paragraph id="6A0336CDFF8DFFC1FF11F807489EF840" blockId="21.[151,925,1994,2016]" box="[151,925,1994,2016]" pageId="21" pageNumber="22">
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Gen.n. Ind117:
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<bibRefCitation id="0E2D4B3CFF8DFFC1FEB3F8074AEAF840" author="Eberle, J. & Dimitrov, D. & Valdez-Mondragon, A. & Huber, B. A." box="[309,489,1994,2016]" pageId="21" pageNumber="22" pagination="141" refId="ref37668" refString="Eberle, J., Dimitrov, D., Valdez-Mondragon, A. & Huber, B. A. (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology, 18, 141. https: // doi. org / 10.1186 / s 12862 - 018 - 1244 - 8" type="journal article" year="2018">
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||
Eberle
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<emphasis id="58C8EADFFF8DFFC1FEFBF8064AB3F840" box="[381,432,1994,2016]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
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2018
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</bibRefCitation>
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||
(molecular data); Huber
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||
<emphasis id="58C8EADFFF8DFFC1FD6EF806481FF840" box="[744,796,1994,2016]" italics="true" pageId="21" pageNumber="22">et al.</emphasis>
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2018: fig. 2.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="3EC36645FF8EFFC2FF11F91B4BC3F88A" httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,264,1750,1772]" targetBox="[151,1436,193,1729]" targetPageId="22">
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<paragraph id="6A0336CDFF8EFFC2FF11F91B4BC3F88A" blockId="22.[151,1436,1750,1834]" pageId="22" pageNumber="23">
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||
<emphasis id="58C8EADFFF8EFFC2FF11F91B4A56F94C" bold="true" box="[151,341,1750,1772]" pageId="22" pageNumber="23">FIGURES 60–64.</emphasis>
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<taxonomicName id="ADBC4D4EFF8EFFC2FEDAF91A4AE5F94C" authority="Huber & Carvalho, 2019" authorityName="Huber & Carvalho" authorityYear="2019" box="[348,486,1750,1772]" class="Arachnida" family="Pholcidae" genus="Arnapa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="tolire" status="sp. nov.">
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<emphasis id="58C8EADFFF8EFFC2FEDAF91A4AE5F94C" box="[348,486,1750,1772]" italics="true" pageId="22" pageNumber="23">Arnapa tolire</emphasis>
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</taxonomicName>
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||
<emphasis id="58C8EADFFF8EFFC2FE6BF91B492BF94C" bold="true" box="[493,552,1750,1772]" pageId="22" pageNumber="23">
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||
<taxonomicNameLabel id="43FB57A4FF8EFFC2FE6BF91B492BF94C" box="[493,552,1750,1772]" pageId="22" pageNumber="23" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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(ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–6 2, 64).
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</paragraph>
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</caption>
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<subSubSection id="22A66546FF8EFFC2FF11F89C4906F876" pageId="22" pageNumber="23" type="materials_examined">
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<paragraph id="6A0336CDFF8EFFC2FF11F89C49F1F82E" blockId="22.[151,1436,1873,2006]" pageId="22" pageNumber="23">
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<emphasis id="58C8EADFFF8EFFC2FF11F89C4A46F8CA" bold="true" box="[151,325,1873,1898]" pageId="22" pageNumber="23">Type material</emphasis>
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.
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<materialsCitation id="DAD43C90FF8EFFC2FEDAF89F49EDF82E" ID-GBIF-Occurrence="2147633699" collectingDate="2009-11-12" collectorName="S. Sutono" country="Indonesia" county="Tolire" elevation="200" latitude="0.83" location="Ar" longLatPrecision="78" longitude="127.311" municipality="Lake" pageId="22" pageNumber="23" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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||
<collectingCountry id="12AB765DFF8EFFC2FEDAF89F4AFAF8CA" box="[348,505,1874,1898]" name="Indonesia" pageId="22" pageNumber="23">INDONESIA</collectingCountry>
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||
:
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<specimenCount id="7CBAFD44FF8EFFC2FD8DF89F4923F8C9" box="[523,544,1874,1897]" pageId="22" pageNumber="23" type="male">♂</specimenCount>
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||
<typeStatus id="B507886FFF8EFFC2FDA8F89C4990F8CA" box="[558,659,1873,1898]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
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||
, ZFMK (
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<location id="6F636016FF8EFFC2FC8DF89F4829F8CA" LSID="urn:lsid:plazi:treatment:E21587DBFF8DFFC3FF11F8924FFEF9A8:6F636016FF8EFFC2FC8DF89F4829F8CA" box="[779,810,1874,1898]" country="Indonesia" county="Tolire" latitude="0.83" longLatPrecision="78" longitude="127.311" municipality="Lake" name="Ar" pageId="22" pageNumber="23">Ar</location>
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||
20611),
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||
<location id="6F636016FF8EFFC2FC18F89F48F6F8C9" LSID="urn:lsid:plazi:treatment:E21587DBFF8DFFC3FF11F8924FFEF9A8:6F636016FF8EFFC2FC18F89F48F6F8C9" box="[926,1013,1874,1897]" country="Indonesia" county="Tolire" latitude="0.83" longLatPrecision="78" longitude="127.311" municipality="Lake" name="Ternate" pageId="22" pageNumber="23">Ternate</location>
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||
, at
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||
<location id="6F636016FF8EFFC2FBACF89F4F70F8C9" LSID="urn:lsid:plazi:treatment:E21587DBFF8DFFC3FF11F8924FFEF9A8:6F636016FF8EFFC2FBACF89F4F70F8C9" box="[1066,1139,1874,1897]" country="Indonesia" county="Tolire" latitude="0.83" longLatPrecision="78" longitude="127.311" municipality="Lake" name="Danau" pageId="22" pageNumber="23">Danau</location>
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||
(
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||
<collectingMunicipality id="8A67ACB7FF8EFFC2FB0CF89C4FC6F8CA" box="[1162,1221,1873,1898]" pageId="22" pageNumber="23">Lake</collectingMunicipality>
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||
)
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||
<collectingCounty id="83624E41FF8EFFC2FB5AF89C4E1DF8CA" box="[1244,1310,1873,1898]" pageId="22" pageNumber="23">Tolire</collectingCounty>
|
||
(
|
||
<geoCoordinate id="0F88500AFF8EFFC2FAB3F89F4E94F8CA" box="[1333,1431,1874,1898]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="55" value="0.83">0.830°N</geoCoordinate>
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||
,
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||
<geoCoordinate id="0F88500AFF8EFFC2FF11F8BB4A12F82E" box="[151,273,1910,1934]" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="55" value="127.311">127.311°E</geoCoordinate>
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||
),
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||
<elevation id="E191D1FEFF8EFFC2FEA2F8BB4AE3F82E" box="[292,480,1909,1934]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="22" pageNumber="23" unit="m" value="200.0" valueMax="220.0" valueMin="180.0">
|
||
<quantity id="AD449B28FF8EFFC2FEA2F8BB4AA0F82D" box="[292,419,1910,1934]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="22" pageNumber="23" unit="m" value="200.0" valueMax="220.0" valueMin="180.0">180–220 m</quantity>
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||
a.s.l.
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||
</elevation>
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||
,
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||
<date id="1E02100DFF8EFFC2FE6AF8B84967F82E" box="[492,612,1909,1934]" pageId="22" pageNumber="23" value="2009-11-12">
|
||
<collectingDate id="0E46E9E5FF8EFFC2FE6AF8B84967F82E" box="[492,612,1909,1934]" pageId="22" pageNumber="23" value="2009-11-12">12.xi.2009</collectingDate>
|
||
</date>
|
||
(
|
||
<collectorName id="C749531BFF8EFFC2FDF3F8BB49E5F82E" box="[629,742,1910,1934]" pageId="22" pageNumber="23">S. Sutono</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF8EFFC2FF41F8544906F876" blockId="22.[151,1436,1873,2006]" pageId="22" pageNumber="23">
|
||
<emphasis id="58C8EADFFF8EFFC2FF41F8544AF4F812" bold="true" box="[199,503,1945,1970]" pageId="22" pageNumber="23">Other material examined</emphasis>
|
||
.
|
||
<materialsCitation id="DAD43C90FF8EFFC2FD83F8574F45F812" ID-GBIF-Occurrence="2147633785" box="[517,1094,1945,1970]" country="Indonesia" location="INDONESIA" pageId="22" pageNumber="23" specimenCount="16" specimenCount-female="11" specimenCount-male="5">
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||
<collectingCountry id="12AB765DFF8EFFC2FD83F85749A1F812" box="[517,674,1946,1970]" name="Indonesia" pageId="22" pageNumber="23">INDONESIA</collectingCountry>
|
||
:
|
||
<specimenCount id="7CBAFD44FF8EFFC2FD28F85449D2F812" box="[686,721,1945,1970]" pageId="22" pageNumber="23" type="male">5♂</specimenCount>
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||
<specimenCount id="7CBAFD44FF8EFFC2FD5FF8574808F812" box="[729,779,1946,1970]" pageId="22" pageNumber="23" type="female">11♀</specimenCount>
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, ZFMK (Ar 20612–13) and
|
||
</materialsCitation>
|
||
<materialsCitation id="DAD43C90FF8EFFC2FBC8F8574902F876" ID-GBIF-Occurrence="2147633774" collectingDate="2009-11-12" collectorName="S. Sutono" country="Indonesia" county="Tolire" elevation="200" latitude="0.83" location="Ar" longLatPrecision="78" longitude="127.311" municipality="Lake" pageId="22" pageNumber="23" specimenCount="1">
|
||
<specimenCount id="7CBAFD44FF8EFFC2FBC8F8574F8DF812" box="[1102,1166,1945,1970]" pageId="22" pageNumber="23" type="generic">1 juv.</specimenCount>
|
||
in pure ethanol, ZFMK (Ind209), same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22A66546FF8FFFC3FF41FF5A4F73FF10" box="[199,1136,151,176]" pageId="23" pageNumber="24" type="etymology">
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FF5A4F73FF10" blockId="23.[151,1437,151,1544]" box="[199,1136,151,176]" pageId="23" pageNumber="24">
|
||
<emphasis id="58C8EADFFF8FFFC3FF41FF5A4A45FF10" bold="true" box="[199,326,151,176]" pageId="23" pageNumber="24">Etymology</emphasis>
|
||
. The species name is derived from the
|
||
<typeStatus id="B507886FFF8FFFC3FD78FF55482DFF10" box="[766,814,152,176]" pageId="23" pageNumber="24">type</typeStatus>
|
||
locality; noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22A66546FF8FFFC3FF41FF714E17FEC5" pageId="23" pageNumber="24" type="diagnosis">
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FF714E17FEC5" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">
|
||
<emphasis id="58C8EADFFF8FFFC3FF41FF714A39FF75" bold="true" box="[199,314,188,213]" pageId="23" pageNumber="24">Diagnosis</emphasis>
|
||
. Distinguished from known congeners by armature of male chelicerae (
|
||
<figureCitation id="F2872A48FF8FFFC3FBF7FF714FC5FF75" box="[1137,1222,188,213]" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="23" pageNumber="24">Fig. 63</figureCitation>
|
||
; ~3–4 large lateral apophyses plus proximal patch of 3–4 smaller apophyses), by more slender palpal segments in general (
|
||
<figureCitation id="F2872A48FF8FFFC3FABEFF124BB7FEBD" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="23" pageNumber="24">Figs 60– 61</figureCitation>
|
||
) and procursus in particular, the latter with distinctive prolateral sclerite distally (
|
||
<figureCitation id="F2872A48FF8FFFC3FBE8FEC94FC6FEBD" box="[1134,1221,260,285]" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="23" pageNumber="24">Fig. 62</figureCitation>
|
||
; with pointed side branch), by shape of bulbal process (
|
||
<figureCitation id="F2872A48FF8FFFC3FDB4FEEA4985FEE0" box="[562,646,295,320]" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="23" pageNumber="24">Fig. 64</figureCitation>
|
||
), by shape of epigynum (
|
||
<figureCitation id="F2872A48FF8FFFC3FC27FEEA48F4FEE0" box="[929,1015,295,320]" captionStart="FIGURES 81–89" captionStartId="27.[151,264,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@27.[151,1436,193,1705]" captionTargetPageId="27" captionText="FIGURES 81–89. Arnapa spp., epigyna, ventral views, and cleared female genitalia, ventral and dorsal views. 81–83. A. meja sp. n. (ZFMK, Ar 20610). 84–86. A. tolire sp. n. (ZFMK, Ar 20613). 87–89. A. tinoor sp. n. (ZFMK, Ar 20616). Scale lines: 0.5." httpUri="https://zenodo.org/record/2619052/files/figure.png" pageId="23" pageNumber="24">Figs 84</figureCitation>
|
||
,
|
||
<figureCitation id="F2872A48FF8FFFC3FB85FEE54F21FEE0" box="[1027,1058,296,320]" captionStart="FIGURES 90–97" captionStartId="28.[151,264,1665,1687]" captionTargetBox="[151,1436,193,1644]" captionTargetId="figure@28.[151,1436,193,1644]" captionTargetPageId="28" captionText="FIGURES 90–97. Arnapa tolire sp. n. (90–94, ZFMK, Ar 20613) and A. tinoor sp. n. (95–97, ZFMK, Ar 20615–16). 90–91. Female prosoma, frontal views. 92. Left female palp and chelicerae; note absence of stridulatory files. 93. Female palpal tarsal organ. 94–95. Epigyna, ventral views. 96–97. Female and male spinnerets. ALS, anterior lateral spinneret; ch, chelicera; PMS, posterior median spinneret; pto, palpal tarsal organ. Scale lines: 10 µm (93, 96–97), 100 µm (90–92, 94–95)." httpUri="https://zenodo.org/record/2619054/files/figure.png" pageId="23" pageNumber="24">94</figureCitation>
|
||
; pair of anterior areas with strong transversal ridges and pair of rounded posterior projections), and by elongate pore plates (
|
||
<figureCitation id="F2872A48FF8FFFC3FB03FE814FDFFEC4" box="[1157,1244,332,357]" captionStart="FIGURES 70–74" captionStartId="25.[151,264,1653,1675]" captionTargetBox="[154,1422,209,1620]" captionTargetId="figure@25.[151,1436,193,1632]" captionTargetPageId="25" captionText="FIGURES 70–74. Arnapa spp., cleared female genitalia, dorsal views. 70. A. arfak sp. n. (ZFMK, Ar 20605). 71. A. manokwari sp. n. (ZFMK, Ar 20608). 72. A. meja sp. n. (ZFMK, Ar 20610). 73. A. tolire sp. n. (ZFMK, Ar 20613). 74. A. tinoor sp. n. (ZFMK, Ar 20616). Scale lines: 0.3." httpUri="https://zenodo.org/record/2619048/files/figure.png" pageId="23" pageNumber="24">Figs 73</figureCitation>
|
||
,
|
||
<figureCitation id="F2872A48FF8FFFC3FB6EFE814E0BFEC5" box="[1256,1288,332,357]" captionStart="FIGURES 81–89" captionStartId="27.[151,264,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@27.[151,1436,193,1705]" captionTargetPageId="27" captionText="FIGURES 81–89. Arnapa spp., epigyna, ventral views, and cleared female genitalia, ventral and dorsal views. 81–83. A. meja sp. n. (ZFMK, Ar 20610). 84–86. A. tolire sp. n. (ZFMK, Ar 20613). 87–89. A. tinoor sp. n. (ZFMK, Ar 20616). Scale lines: 0.5." httpUri="https://zenodo.org/record/2619052/files/figure.png" pageId="23" pageNumber="24">86</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22A66546FF8FFFC3FF41FEA24896FA3C" pageId="23" pageNumber="24" type="description">
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FEA24F78FE70" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">
|
||
<emphasis id="58C8EADFFF8FFFC3FF41FEA24A9CFE28" bold="true" box="[199,415,367,392]" pageId="23" pageNumber="24">Description. Male</emphasis>
|
||
(
|
||
<typeStatus id="B507886FFF8FFFC3FE36FEA2491BFE28" box="[432,536,367,392]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
|
||
). MEASUREMENTS. Total length 2.1, carapace width 0.95. Distance PME-PME 75 µm; diameter PME 85 µm; distance PME-ALE 60 µm; distance AME-AME 15 µm, diameter AME 40 µm. Leg 1: 24.6 (5.9 + 0.4 + 6.1 + 10.2 + 2.0), tibia 2: 3.4, tibia 3: 2.4, tibia 4: 3.3; tibia 1 L/d: 81.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FE1148E1FD08" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">COLOR (in ethanol). Carapace ochre-yellow with dark brown bands laterally and wide band medially including ocular area; clypeus with pair of dark bands below eye triads; sternum ochre-yellow with pair of light brown bands on both sides of median line; legs brown, with indistinct darker rings on femora (subdistally) and tibiae (proximally and subdistally), tips of femora and tibiae whitish. Abdomen gray, dorsally densely covered with dark marks, ventrally with large brown mark in front of gonopore, dark median band behind gonopore, pair of brown external marks and pair of larger internal ventro-lateral marks in front of spinnerets.</paragraph>
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FD784919FD50" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">
|
||
BODY. Habitus as in
|
||
<figureCitation id="F2872A48FF8FFFC3FE2BFD794AFDFD6C" box="[429,510,692,717]" captionStart="FIGURES 39–44" captionStartId="16.[151,264,1927,1949]" captionTargetBox="[151,1436,589,1905]" captionTargetId="figure@16.[151,1436,589,1905]" captionTargetPageId="16" captionText="FIGURES 39–44. Arnapa spp., live specimens. 39–40. A. arfak sp. n., male and female from Mokwam. 41–42. A. tolire sp. n., male and female with egg sac from Ternate. 43–44. A. tinoor sp. n., male and female from near Tomohon." httpUri="https://zenodo.org/record/2619036/files/figure.png" pageId="23" pageNumber="24">Fig. 41</figureCitation>
|
||
. Ocular area elevated, thoracic furrow present; clypeus unmodified. Sternum wider than long (0.58/0.36), unmodified.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FD3148EAFC98" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">
|
||
CHELICERAE. As in
|
||
<figureCitation id="F2872A48FF8FFFC3FE2AFD314903FCB5" box="[428,512,764,789]" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="23" pageNumber="24">Fig. 63</figureCitation>
|
||
, each side with 3–4 large sclerotized cones along lateral margin and 3–4 smaller cones frontally proximally; without modified hairs; with stridulatory ridges.
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FC8849CEFBB0" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">
|
||
PALPS. As in
|
||
<figureCitation id="F2872A48FF8FFFC3FED9FC894ADCFCFD" box="[351,479,836,861]" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="23" pageNumber="24">Figs 60–61</figureCitation>
|
||
; coxa unmodified, trochanter ventrally only slightly protruding, femur widened, with rounded retrolateral process proximally, with prolateral stridulatory pick (modified hair) proximally and stronger than usual hairs distally on ventral-retrolateral side, patella ventrally reduced (not closed), tibia longer than in known relatives, retrolateral trichobothrium in very proximal position, tarsus small, procursus with dorsal process proximally, ventrally without pocket, distally with distinctive sclerotized and membranous elements (
|
||
<figureCitation id="F2872A48FF8FFFC3FAB0FC194E8CFC4D" box="[1334,1423,980,1005]" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="23" pageNumber="24">Fig. 62</figureCitation>
|
||
); genital bulb large, with complex process (
|
||
<figureCitation id="F2872A48FF8FFFC3FDEAFC3A49C3FBB0" box="[620,704,1015,1040]" captionStart="FIGURES 60–64" captionStartId="22.[151,264,1750,1772]" captionTargetBox="[151,1436,193,1729]" captionTargetId="figure@22.[151,1436,193,1729]" captionTargetPageId="22" captionText="FIGURES 60–64. Arnapa tolire sp. n. (ZFMK, Ar 20612). 60–61. Left male palp, prolateral and retrolateral views. 62. Left procursus, dorsal view. 63. Male chelicerae, frontal view. 64. Left genital bulb, dorsal view. Scale lines: 0.3 (63), 0.5 (60–62, 64)." httpUri="https://zenodo.org/record/2619044/files/figure.png" pageId="23" pageNumber="24">Fig. 64</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FBD04913FBDD" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">LEGS. Without spines, few vertical hairs; with curved hairs on tibiae and metatarsi 1–2 (few on legs 2); retrolateral trichobothrium of tibia 1 at 11%; prolateral trichobothrium present on all tibiae; tarsus 1 with ~30 pseudosegments, distally distinct.</paragraph>
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FB4A4988FB65" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">
|
||
<emphasis id="58C8EADFFF8FFFC3FF41FB4A4A07FB00" bold="true" box="[199,260,1159,1184]" pageId="23" pageNumber="24">Male</emphasis>
|
||
(variation). Size and number of sclerotized cones on chelicerae slightly variable but pattern consistent. Tibia
|
||
<quantity id="AD449B28FF8FFFC3FF5EFB614A00FB65" box="[216,259,1196,1221]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="23" pageNumber="24" unit="in" value="1.0">1 in</quantity>
|
||
5 other males: 5.2–6.1 (mean 5.7).
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FB024896FA3C" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">
|
||
<emphasis id="58C8EADFFF8FFFC3FF41FB024A1DFB48" bold="true" box="[199,286,1231,1256]" pageId="23" pageNumber="24">Female</emphasis>
|
||
. In general similar to male (
|
||
<figureCitation id="F2872A48FF8FFFC3FDE1FB0249BDFB48" box="[615,702,1231,1256]" captionStart="FIGURES 39–44" captionStartId="16.[151,264,1927,1949]" captionTargetBox="[151,1436,589,1905]" captionTargetId="figure@16.[151,1436,589,1905]" captionTargetPageId="16" captionText="FIGURES 39–44. Arnapa spp., live specimens. 39–40. A. arfak sp. n., male and female from Mokwam. 41–42. A. tolire sp. n., male and female with egg sac from Ternate. 43–44. A. tinoor sp. n., male and female from near Tomohon." httpUri="https://zenodo.org/record/2619036/files/figure.png" pageId="23" pageNumber="24">Fig. 42</figureCitation>
|
||
), also with curved hairs on legs. Chelicerae without stridulatory ridges (
|
||
<figureCitation id="F2872A48FF8FFFC3FF6CFB394A3CFAAC" box="[234,319,1268,1293]" captionStart="FIGURES 90–97" captionStartId="28.[151,264,1665,1687]" captionTargetBox="[151,1436,193,1644]" captionTargetId="figure@28.[151,1436,193,1644]" captionTargetPageId="28" captionText="FIGURES 90–97. Arnapa tolire sp. n. (90–94, ZFMK, Ar 20613) and A. tinoor sp. n. (95–97, ZFMK, Ar 20615–16). 90–91. Female prosoma, frontal views. 92. Left female palp and chelicerae; note absence of stridulatory files. 93. Female palpal tarsal organ. 94–95. Epigyna, ventral views. 96–97. Female and male spinnerets. ALS, anterior lateral spinneret; ch, chelicera; PMS, posterior median spinneret; pto, palpal tarsal organ. Scale lines: 10 µm (93, 96–97), 100 µm (90–92, 94–95)." httpUri="https://zenodo.org/record/2619054/files/figure.png" pageId="23" pageNumber="24">Fig. 92</figureCitation>
|
||
). Tibia
|
||
<quantity id="AD449B28FF8FFFC3FE15FB394ABDFAAD" box="[403,446,1268,1293]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="23" pageNumber="24" unit="in" value="1.0">1 in</quantity>
|
||
<specimenCount id="7CBAFD44FF8FFFC3FE42FB394936FAAD" box="[452,565,1268,1293]" pageId="23" pageNumber="24" type="female">9 females</specimenCount>
|
||
: 3.8–4.6 (mean 4.1). Palpal tarsal organ exposed (
|
||
<figureCitation id="F2872A48FF8FFFC3FBE4FB394FB5FAAC" box="[1122,1206,1268,1293]" captionStart="FIGURES 90–97" captionStartId="28.[151,264,1665,1687]" captionTargetBox="[151,1436,193,1644]" captionTargetId="figure@28.[151,1436,193,1644]" captionTargetPageId="28" captionText="FIGURES 90–97. Arnapa tolire sp. n. (90–94, ZFMK, Ar 20613) and A. tinoor sp. n. (95–97, ZFMK, Ar 20615–16). 90–91. Female prosoma, frontal views. 92. Left female palp and chelicerae; note absence of stridulatory files. 93. Female palpal tarsal organ. 94–95. Epigyna, ventral views. 96–97. Female and male spinnerets. ALS, anterior lateral spinneret; ch, chelicera; PMS, posterior median spinneret; pto, palpal tarsal organ. Scale lines: 10 µm (93, 96–97), 100 µm (90–92, 94–95)." httpUri="https://zenodo.org/record/2619054/files/figure.png" pageId="23" pageNumber="24">Fig. 93</figureCitation>
|
||
). Epigynum anterior plate square-shaped (
|
||
<figureCitation id="F2872A48FF8FFFC3FE00FADA4ADDFA90" box="[390,478,1303,1328]" captionStart="FIGURES 81–89" captionStartId="27.[151,264,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@27.[151,1436,193,1705]" captionTargetPageId="27" captionText="FIGURES 81–89. Arnapa spp., epigyna, ventral views, and cleared female genitalia, ventral and dorsal views. 81–83. A. meja sp. n. (ZFMK, Ar 20610). 84–86. A. tolire sp. n. (ZFMK, Ar 20613). 87–89. A. tinoor sp. n. (ZFMK, Ar 20616). Scale lines: 0.5." httpUri="https://zenodo.org/record/2619052/files/figure.png" pageId="23" pageNumber="24">Figs 84</figureCitation>
|
||
,
|
||
<figureCitation id="F2872A48FF8FFFC3FE6DFAD54909FA90" box="[491,522,1304,1328]" captionStart="FIGURES 90–97" captionStartId="28.[151,264,1665,1687]" captionTargetBox="[151,1436,193,1644]" captionTargetId="figure@28.[151,1436,193,1644]" captionTargetPageId="28" captionText="FIGURES 90–97. Arnapa tolire sp. n. (90–94, ZFMK, Ar 20613) and A. tinoor sp. n. (95–97, ZFMK, Ar 20615–16). 90–91. Female prosoma, frontal views. 92. Left female palp and chelicerae; note absence of stridulatory files. 93. Female palpal tarsal organ. 94–95. Epigyna, ventral views. 96–97. Female and male spinnerets. ALS, anterior lateral spinneret; ch, chelicera; PMS, posterior median spinneret; pto, palpal tarsal organ. Scale lines: 10 µm (93, 96–97), 100 µm (90–92, 94–95)." httpUri="https://zenodo.org/record/2619054/files/figure.png" pageId="23" pageNumber="24">94</figureCitation>
|
||
), with two areas of deep transversal ridges anteriorly laterally, medially whitish, posterior rim with pair of rounded projections; with separate brown plate in front of epigynum; posterior plate very short but wide. Internal genitalia with distinctive lateral folds, narrow anterior ‘valve’, and elongated pore plates (
|
||
<figureCitation id="F2872A48FF8FFFC3FF19FA494BF6FA3C" box="[159,245,1412,1437]" captionStart="FIGURES 70–74" captionStartId="25.[151,264,1653,1675]" captionTargetBox="[154,1422,209,1620]" captionTargetId="figure@25.[151,1436,193,1632]" captionTargetPageId="25" captionText="FIGURES 70–74. Arnapa spp., cleared female genitalia, dorsal views. 70. A. arfak sp. n. (ZFMK, Ar 20605). 71. A. manokwari sp. n. (ZFMK, Ar 20608). 72. A. meja sp. n. (ZFMK, Ar 20610). 73. A. tolire sp. n. (ZFMK, Ar 20613). 74. A. tinoor sp. n. (ZFMK, Ar 20616). Scale lines: 0.3." httpUri="https://zenodo.org/record/2619048/files/figure.png" pageId="23" pageNumber="24">Figs 73</figureCitation>
|
||
,
|
||
<figureCitation id="F2872A48FF8FFFC3FE87FA494A22FA3D" box="[257,289,1412,1437]" captionStart="FIGURES 81–89" captionStartId="27.[151,264,1726,1748]" captionTargetBox="[151,1436,193,1705]" captionTargetId="figure@27.[151,1436,193,1705]" captionTargetPageId="27" captionText="FIGURES 81–89. Arnapa spp., epigyna, ventral views, and cleared female genitalia, ventral and dorsal views. 81–83. A. meja sp. n. (ZFMK, Ar 20610). 84–86. A. tolire sp. n. (ZFMK, Ar 20613). 87–89. A. tinoor sp. n. (ZFMK, Ar 20616). Scale lines: 0.5." httpUri="https://zenodo.org/record/2619052/files/figure.png" pageId="23" pageNumber="24">86</figureCitation>
|
||
). ALS with only two spigots each (confirmed by SEM).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="22A66546FF8FFFC3FF41FA6A4FFEF9A8" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FA6A4842FA60" blockId="23.[151,1437,151,1544]" box="[199,833,1447,1472]" pageId="23" pageNumber="24">
|
||
<emphasis id="58C8EADFFF8FFFC3FF41FA6A4A5AFA60" bold="true" box="[199,345,1447,1472]" pageId="23" pageNumber="24">Distribution</emphasis>
|
||
. Known from
|
||
<typeStatus id="B507886FFF8FFFC3FE78FA65492DFA60" box="[510,558,1448,1472]" pageId="23" pageNumber="24">type</typeStatus>
|
||
locality only (
|
||
<figureCitation id="F2872A48FF8FFFC3FD57FA6A4836FA60" box="[721,821,1447,1472]" captionStart="FIGURE 343" captionStartId="89.[151,250,1861,1883]" captionTargetBox="[151,1436,1172,1840]" captionTargetId="figure@89.[151,1436,1172,1840]" captionTargetPageId="89" captionText="FIGURE 343. Known distribution of Arnapa gen. n.; red marks: undescribed species. Further undescribed species that might belong in this genus are known from the Caroline Islands and from Papua New Guinea east to the Louisiade Archipelago." httpUri="https://zenodo.org/record/2619156/files/figure.png" pageId="23" pageNumber="24">Fig. 343</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6A0336CDFF8FFFC3FF41FA014FFEF9A8" blockId="23.[151,1437,151,1544]" pageId="23" pageNumber="24">
|
||
<emphasis id="58C8EADFFF8FFFC3FF41FA014A7CFA45" bold="true" box="[199,383,1484,1509]" pageId="23" pageNumber="24">Natural history</emphasis>
|
||
. The spiders were found in a degraded forest in a ravine leading into the lake; their domed webs were built among rocks near the ground and under large dead palm leaves lying on the ground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |