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<document id="5B521115969AA724A540209C28986948" ID-CLB-Dataset="31530" ID-DOI="10.5281/zenodo.1050060" ID-GBIF-Dataset="2d28bebf-e3c5-457b-9152-8a6877ded281" ID-ISSN="1175-5326" ID-Zenodo-Dep="1050060" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510742923475" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2011" docId="C063786AFFA2FFF1FF0D13BD5ABA90CC" docLanguage="en" docName="zt03074p198.pdf.imf" docOrigin="Zootaxa 3074" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena" docType="key" docVersion="7" lastPageNumber="13" masterDocId="3C5A0012FFAEFFFDFF9A11335F219540" masterDocTitle="New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)" masterLastPageNumber="198" masterPageNumber="1" pageNumber="13" updateTime="1698485333395" updateUser="ExternalLinkService">
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<mods:title id="9A6645FAAA9C5EDAA291F071C62EFFB7">New records and description of fifty-four new species of aquatic beetles in the genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="5AF4A7592584A3BD2F267608B7B97CF9">Perkins, Philip D.</mods:namePart>
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<mods:date id="C58B84ACF41432C830526FB6E5DB6F89">2011</mods:date>
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<paragraph id="4875C97CFFA2FFF1FF0D13BD5DA597E8" blockId="12.[151,644,654,680]" box="[151,644,654,680]" pageId="12" pageNumber="13">
<heading id="133D7E10FFA2FFF1FF0D13BD5DA597E8" bold="true" box="[151,644,654,680]" fontSize="11" level="1" pageId="12" pageNumber="13" reason="1">
<emphasis id="7ABE156EFFA2FFF1FF0D13BD5DA597E8" bold="true" box="[151,644,654,680]" pageId="12" pageNumber="13">
Key to Groups of New World
<taxonomicName id="8FCAB2FFFFA2FFF1FD9113BD5DA597E8" ID-CoL="8VRR8" box="[523,644,654,680]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="7ABE156EFFA2FFF1FD9113BD5DA597E8" bold="true" box="[523,644,654,680]" italics="true" pageId="12" pageNumber="13">Hydraena</emphasis>
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<keyLead id="F33B6A49FFA2FFF1FF0D13E65ABD97AA" box="[151,1436,725,746]" pageId="12" pageNumber="13">
1 Posterior ridge of genae present, at least laterally (e.g.,
<figureCitation id="D0F1D5F9FFA2FFF1FD7413E65C1197AA" box="[750,816,725,746]" captionStart="FIGURE 2" captionStartId="71.[151,250,1650,1673]" captionTargetBox="[278,1308,193,1626]" captionTargetId="figure@71.[278,1308,193,1629]" captionTargetPageId="71" captionText="FIGURE 2. Pronotal sculpture and measurements, and metavental structures and measurements." httpUri="https://zenodo.org/record/1050064/files/figure.png" pageId="12" pageNumber="13">Figs. 2</figureCitation>
c, 2d); abdominal intercoxal ventrite various; scintilla absent ... 2
</keyLead>
</paragraph>
<paragraph id="4875C97CFFA2FFF1FF0D13C15ABD9664" blockId="12.[151,1436,725,804]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D13C15ABD9664" pageId="12" pageNumber="13">
1' Posterior ridge of genae absent (e.g.,
<figureCitation id="D0F1D5F9FFA2FFF1FDD013C15DBA9647" box="[586,667,754,775]" captionStart="FIGURE 44" captionStartId="102.[151,250,1650,1673]" captionTargetBox="[291,1310,212,1626]" captionTargetId="figure@102.[265,1322,193,1629]" captionTargetPageId="102" captionText="FIGURE 44. Hydraena scintillabella, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050110/files/figure.png" pageId="12" pageNumber="13">Figs. 44</figureCitation>
,
<figureCitation id="D0F1D5F9FFA2FFF1FD3F13C15DEB9647" box="[677,714,754,775]" captionStart="FIGURE 126" captionStartId="159.[151,250,1650,1673]" captionTargetBox="[267,1320,209,1615]" captionTargetId="figure@159.[248,1338,193,1629]" captionTargetPageId="159" captionText="FIGURE 126. Hydraena ampla, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050198/files/figure.png" pageId="12" pageNumber="13">126</figureCitation>
) or nearly imperceptible; abdominal intercoxal ventrite flat, with posterior margin straight or weakly arcuate; scintilla (
<figureCitation id="D0F1D5F9FFA2FFF1FD12123C5DE09664" box="[648,705,783,804]" captionStart="FIGURE 2" captionStartId="71.[151,250,1650,1673]" captionTargetBox="[278,1308,193,1626]" captionTargetId="figure@71.[278,1308,193,1629]" captionTargetPageId="71" captionText="FIGURE 2. Pronotal sculpture and measurements, and metavental structures and measurements." httpUri="https://zenodo.org/record/1050064/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
b) present or absent........................................... 5
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<keyStep id="F33ED1D9FFA2FFF1FF0D12795ABD96DA" pageId="12" pageNumber="13">
<paragraph id="4875C97CFFA2FFF1FF0D12795ABA963C" blockId="12.[151,1436,842,922]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D12795ABA963C" pageId="12" pageNumber="13">
2 Elytra and labrum with spines (
<figureCitation id="D0F1D5F9FFA2FFF1FD8E12795D44961F" box="[532,613,842,863]" captionStart="FIGURE 21" captionStartId="85.[151,250,1650,1673]" captionTargetBox="[253,1331,193,1626]" captionTargetId="figure@85.[253,1334,193,1629]" captionTargetPageId="85" captionText="FIGURE 21. Hydraena multispina, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050084/files/figure.png" pageId="12" pageNumber="13">Figs. 21</figureCitation>
,
<figureCitation id="D0F1D5F9FFA2FFF1FDEA12795DAA961F" box="[624,651,842,863]" captionStart="FIGURE 22" captionStartId="86.[151,250,1650,1673]" captionTargetBox="[305,1294,233,1606]" captionTargetId="figure@86.[270,1317,193,1629]" captionTargetPageId="86" captionText="FIGURE 22. Hydraena cordispina, holotype female, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050086/files/figure.png" pageId="12" pageNumber="13">22</figureCitation>
); elytron with five rows of punctures between suture and humerous; two species from South
<collectingCountry id="30DD89ECFFA2FFF1FEC912545E87963C" box="[339,422,871,892]" name="United States of America" pageId="12" pageNumber="13">America</collectingCountry>
......................................................................
<taxonomicName id="8FCAB2FFFFA2FFF1FB77125B5A78963D" ID-CoL="3N3YY" box="[1261,1369,872,893]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="86" phylum="Arthropoda" rank="species" species="multispina">
<emphasis id="7ABE156EFFA2FFF1FB77125B5A78963D" box="[1261,1369,872,893]" italics="true" pageId="12" pageNumber="13">Multispina</emphasis>
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Group
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</paragraph>
<paragraph id="4875C97CFFA2FFF1FF0D12B65ABD96DA" blockId="12.[151,1436,842,922]" box="[151,1436,901,922]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D12B65ABD96DA" box="[151,1436,901,922]" pageId="12" pageNumber="13">2' Elytra and labrum without spines; elytron with more than five rows, or with many random punctures................... 3</keyLead>
</paragraph>
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<keyStep id="F33ED1D9FFA2FFF1FF0D128C5ABD9127" pageId="12" pageNumber="13">
<paragraph id="4875C97CFFA2FFF1FF0D128C5ABA914F" blockId="12.[151,1437,959,1127]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D128C5ABA914F" pageId="12" pageNumber="13">
3 Abdominal intercoxal ventrite concave, with posterior margin markedly arcuate; elytral punctures serial or, in few adults, with few random punctures on disc; spermatheca C-shaped; female gonocoxite not divided;
<collectingCountry id="30DD89ECFFA2FFF1FBBE12EE5B4D96B2" box="[1060,1132,989,1010]" name="Canada" pageId="12" pageNumber="13">Canada</collectingCountry>
and the
<collectingCountry id="30DD89ECFFA2FFF1FB2412EE5A1F96B2" box="[1214,1342,989,1010]" name="United States of America" pageId="12" pageNumber="13">United States</collectingCountry>
...............................................................................................
<emphasis id="7ABE156EFFA2FFF1FB6712C95A79914F" box="[1277,1368,1018,1039]" italics="true" pageId="12" pageNumber="13">Circulata</emphasis>
Group
</keyLead>
</paragraph>
<paragraph id="4875C97CFFA2FFF1FF0D15245ABD9127" blockId="12.[151,1437,959,1127]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D15245ABD9127" pageId="12" pageNumber="13">
3' Abdominal intercoxal ventrite flat, with posterior margin straight or weakly arcuate; elytral punctures random, at least near base; spermatheca disc-shaped (e.g.
<figureCitation id="D0F1D5F9FFA2FFF1FDA715065DE9910A" box="[573,712,1077,1098]" captionStart="FIGURES 151 154" captionStartId="177.[151,264,1650,1673]" captionTargetBox="[236,1345,201,1619]" captionTargetId="figure@177.[229,1357,193,1629]" captionTargetPageId="177" captionText="FIGURES 151 154. Spermathecae of Hydraena species." httpUri="https://zenodo.org/record/1050225/files/figure.png" pageId="12" pageNumber="13">Figs. 151154</figureCitation>
) or (in
<taxonomicName id="8FCAB2FFFFA2FFF1FC9515065CB5910A" ID-CoL="3N2JH" box="[783,916,1077,1098]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="paeminosa">
<emphasis id="7ABE156EFFA2FFF1FC9515065CB5910A" box="[783,916,1077,1098]" italics="true" pageId="12" pageNumber="13">H. paeminosa</emphasis>
</taxonomicName>
) trilobed (
<figureCitation id="D0F1D5F9FFA2FFF1FC6015065B6F910A" box="[1018,1102,1077,1098]" captionStart="FIGURES 155 159" captionStartId="178.[151,264,1608,1631]" captionText="FIGURES 155 159. Spermathecae of Hydraena species." pageId="12" pageNumber="13">Fig. 155</figureCitation>
); female gonocoxite divided (
<figureCitation id="D0F1D5F9FFA2FFF1FAF615065E199127" captionStart="FIGURES 148 150" captionStartId="176.[151,264,1581,1604]" captionTargetBox="[237,1360,212,1541]" captionTargetId="figure@176.[226,1360,193,1560]" captionTargetPageId="176" captionText="FIGURES 148 150. Female terminal abdominal segments of Hydraena species." httpUri="https://zenodo.org/record/1050223/files/figure.png" pageId="12" pageNumber="13">Figs. 148150</figureCitation>
); South
<collectingCountry id="30DD89ECFFA2FFF1FE1D15615EFB9127" box="[391,474,1106,1127]" name="United States of America" pageId="12" pageNumber="13">America</collectingCountry>
.............................................................................. 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="F33ED1D9FFA2FFF1FF0D15BE5ABA9057" pageId="12" pageNumber="13">
<paragraph id="4875C97CFFA2FFF1FF0D15BE5ABA919C" blockId="12.[151,1436,1165,1303]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D15BE5ABA919C" pageId="12" pageNumber="13">
4. Elytral punctures coarse and very random (e.g.,
<figureCitation id="D0F1D5F9FFA2FFF1FD2615BE5DD691E2" box="[700,759,1165,1186]" captionStart="FIGURE 3" captionStartId="72.[151,250,1650,1673]" captionTargetBox="[227,1366,208,1616]" captionTargetId="figure@72.[208,1379,193,1629]" captionTargetPageId="72" captionText="FIGURE 3. Hydraena paeminosa, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050066/files/figure.png" pageId="12" pageNumber="13">Fig. 3</figureCitation>
); pronotum more elongate; mesoventral intercoxal process (P2) very narrow, tapering to sharp point that is contiguous with midlongitudinal carina of metaventrite (e.g.,
<figureCitation id="D0F1D5F9FFA2FFF1FB0115995BF591FF" box="[1179,1236,1194,1215]" captionStart="FIGURE 2" captionStartId="71.[151,250,1650,1673]" captionTargetBox="[278,1308,193,1626]" captionTargetId="figure@71.[278,1308,193,1629]" captionTargetPageId="71" captionText="FIGURE 2. Pronotal sculpture and measurements, and metavental structures and measurements." httpUri="https://zenodo.org/record/1050064/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
c)....................................................................................................
<taxonomicName id="8FCAB2FFFFA2FFF1FB7415FB5A79919D" ID-CoL="3N2JH" box="[1262,1368,1224,1245]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="73" phylum="Arthropoda" rank="species" species="paeminosa">
<emphasis id="7ABE156EFFA2FFF1FB7415FB5A79919D" box="[1262,1368,1224,1245]" italics="true" pageId="12" pageNumber="13">Paeminosa</emphasis>
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Group
</keyLead>
</paragraph>
<paragraph id="4875C97CFFA2FFF1FF0D15D65ABA9057" blockId="12.[151,1436,1165,1303]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D15D65ABA9057" pageId="12" pageNumber="13">
4'. Elytral punctures serial except in small area at base (
<figureCitation id="D0F1D5F9FFA2FFF1FD7315D65C4091BA" box="[745,865,1253,1274]" captionStart-0="FIGURE 16" captionStart-1="FIGURE 17" captionStart-2="FIGURE 18" captionStartId-0="81.[151,250,1650,1673]" captionStartId-1="82.[151,250,1650,1673]" captionStartId-2="83.[151,250,1628,1651]" captionTargetBox-0="[228,1355,193,1626]" captionTargetBox-1="[238,1348,193,1626]" captionTargetBox-2="[246,1343,212,1586]" captionTargetId-0="figure@81.[228,1359,193,1629]" captionTargetId-1="figure@82.[238,1348,193,1629]" captionTargetId-2="figure@83.[226,1360,193,1607]" captionTargetPageId-0="81" captionTargetPageId-1="82" captionTargetPageId-2="83" captionText-0="FIGURE 16. Hydraena curvosa, holotype, dorsal and lateral habitus, and venter." captionText-1="FIGURE 17. Hydraena takutu, holotype, dorsal and lateral habitus, and venter." captionText-2="FIGURE 18. Hydraena hintoni, holotype, dorsal and lateral habitus, and venter." httpUri-0="https://zenodo.org/record/1050078/files/figure.png" httpUri-1="https://zenodo.org/record/1050080/files/figure.png" httpUri-2="https://zenodo.org/record/1050082/files/figure.png" pageId="12" pageNumber="13">Figs. 1618</figureCitation>
); pronotum less elongate, sides more sinuate; mesoventral intercoxal process (P2) wide, apex blunt (e.g.,
<figureCitation id="D0F1D5F9FFA2FFF1FD0514315DF69057" box="[671,727,1282,1303]" captionStart="FIGURE 2" captionStartId="71.[151,250,1650,1673]" captionTargetBox="[278,1308,193,1626]" captionTargetId="figure@71.[278,1308,193,1629]" captionTargetPageId="71" captionText="FIGURE 2. Pronotal sculpture and measurements, and metavental structures and measurements." httpUri="https://zenodo.org/record/1050064/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
d)............................................
<taxonomicName id="8FCAB2FFFFA2FFF1FA9D14315A799057" ID-CoL="3N38K" box="[1287,1368,1282,1303]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="82" phylum="Arthropoda" rank="species" species="curvosa">
<emphasis id="7ABE156EFFA2FFF1FA9D14315A799057" box="[1287,1368,1282,1303]" italics="true" pageId="12" pageNumber="13">Curvosa</emphasis>
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Group
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</paragraph>
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<keyStep id="F33ED1D9FFA2FFF1FF0D140E5ABA90CC" pageId="12" pageNumber="13">
<paragraph id="4875C97CFFA2FFF1FF0D140E5ABA9012" blockId="12.[151,1436,1341,1421]" box="[151,1435,1341,1362]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D140E5ABA9012" box="[151,1435,1341,1362]" pageId="12" pageNumber="13">
5 Pronotal scintilla (
<figureCitation id="D0F1D5F9FFA2FFF1FE0B140E5EEB9012" box="[401,458,1341,1362]" captionStart="FIGURE 2" captionStartId="71.[151,250,1650,1673]" captionTargetBox="[278,1308,193,1626]" captionTargetId="figure@71.[278,1308,193,1629]" captionTargetPageId="71" captionText="FIGURE 2. Pronotal sculpture and measurements, and metavental structures and measurements." httpUri="https://zenodo.org/record/1050064/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
b) present or absent; pronotum always with posteromedial foveae (PF2;
<figureCitation id="D0F1D5F9FFA2FFF1FBE4140E5B979012" box="[1150,1206,1341,1362]" captionStart="FIGURE 2" captionStartId="71.[151,250,1650,1673]" captionTargetBox="[278,1308,193,1626]" captionTargetId="figure@71.[278,1308,193,1629]" captionTargetPageId="71" captionText="FIGURE 2. Pronotal sculpture and measurements, and metavental structures and measurements." httpUri="https://zenodo.org/record/1050064/files/figure.png" pageId="12" pageNumber="13">Fig. 2</figureCitation>
a)......
<emphasis id="7ABE156EFFA2FFF1FA83140E5A789012" box="[1305,1369,1341,1362]" italics="true" pageId="12" pageNumber="13">Leechi</emphasis>
Group
</keyLead>
</paragraph>
<paragraph id="4875C97CFFA2FFF1FF0D14695ABA90CC" blockId="12.[151,1436,1341,1421]" pageId="12" pageNumber="13">
<keyLead id="F33B6A49FFA2FFF1FF0D14695ABA90CC" pageId="12" pageNumber="13">
5' Pronotum never with scintilla; pronotum without posteromedial foveae, or if present very shallow and pronotum coarsely punctate (e.g.,
<figureCitation id="D0F1D5F9FFA2FFF1FEF414445EE890CC" box="[366,457,1399,1420]" captionStart="FIGURE 110" captionStartId="148.[151,250,1650,1673]" captionTargetBox="[255,1324,211,1618]" captionTargetId="figure@148.[245,1342,193,1629]" captionTargetPageId="148" captionText="FIGURE 110. Hydraena loripes, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050184/files/figure.png" pageId="12" pageNumber="13">Figs. 110</figureCitation>
,
<figureCitation id="D0F1D5F9FFA2FFF1FE4914445ED690CC" box="[467,503,1399,1420]" captionStart="FIGURE 118" captionStartId="153.[151,250,1650,1673]" captionTargetBox="[237,1344,203,1620]" captionTargetId="figure@153.[227,1359,193,1629]" captionTargetPageId="153" captionText="FIGURE 118. Hydraena steineri, holotype, dorsal and lateral habitus, and venter." httpUri="https://zenodo.org/record/1050190/files/figure.png" pageId="12" pageNumber="13">118</figureCitation>
)...........................................................
<emphasis id="7ABE156EFFA2FFF1FB4D144B5A7890CD" box="[1239,1369,1400,1421]" italics="true" pageId="12" pageNumber="13">Marginicollis</emphasis>
Group
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
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