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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1019.59881" ID-GBIF-Dataset="c81147b7-467a-4477-a36e-76f3fd8358c7" ID-PMC="PMC7921074" ID-Pensoft-Pub="1313-2970-1019-93" ID-Pensoft-UUID="68E025484FB95656A3666768A4D97EF1" ID-PubMed="33688849" ID-ZooBank="740EFFB93ADA4B2ABD23A839AAE71FB2" ModsDocID="1313-2970-1019-93" checkinTime="1614092297092" checkinUser="pensoft" docAuthor="Giron, Jennifer C. &amp; Short, Andrew Edward Z." docDate="2021" docId="C005666E376B50FC8C4C154284A560E4" docLanguage="en" docName="ZooKeys 1019: 93-140" docOrigin="ZooKeys 1019" docPubDate="2021-02-22" docSource="http://dx.doi.org/10.3897/zookeys.1019.59881" docTitle="Tobochares sulcatus species group" docType="treatment" docVersion="5" id="68E025484FB95656A3666768A4D97EF1" lastPageNumber="93" masterDocId="68E025484FB95656A3666768A4D97EF1" masterDocTitle="Review of the Neotropical water scavenger beetle genus Tobochares Short &amp; Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records" masterLastPageNumber="140" masterPageNumber="93" pageNumber="93" updateTime="1668149822001" updateUser="ExternalLinkService">
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<mods:title>Review of the Neotropical water scavenger beetle genus Tobochares Short &amp; Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae): new lineages, new species, and new records</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Giron, Jennifer C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0851-6883</mods:nameIdentifier>
<mods:affiliation>Department of Entomology, Purdue University, West Lafayette, IN 47907, USA &amp; Natural Science Research Laboratory, Museum of Texas Tech University, Lubbock, TX 79409, USA</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Short, Andrew Edward Z.</mods:namePart>
<mods:affiliation>Department of Ecology and Evolutionary Biology, and Division of Entomology, Biodiversity Institute, University of Kansas, Lawrence, KS 66045, USA</mods:affiliation>
<mods:nameIdentifier type="email">aezshort@ku.edu</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2021-02-22</mods:number>
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<mods:number>1019</mods:number>
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<mods:start>93</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1019.59881</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1019.59881</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1019-93</mods:identifier>
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<taxonomicName LSID="C005666E-376B-50FC-8C4C-154284A560E4" authority="species group" authorityName="species group" class="Insecta" family="Hydrophilidae" genus="Tobochares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tobochares sulcatus" order="Coleoptera" pageId="0" pageNumber="93" phylum="Arthropoda" rank="species" species="sulcatus">Tobochares sulcatus species group</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="93" type="diagnosis">
<paragraph pageId="0" pageNumber="93">Diagnosis.</paragraph>
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This species group can be recognized by the oblique and posteriorly directed anterior margin of the eye in lateral view, which emarginates the eye in lateral view (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Heads of Tobochares spp. A, B T. goias, black mark pointing to Tobochares canthus emarginating the eye A dorsal view B anterolateral view C T. fusus D, E T. luteomargo, black mark pointing to straight anterior margin of the eye D dorsal view E anterolateral view F T. romanoae G T. pemon H T. communis I T. microps." figureDoi="10.3897/zookeys.1019.59881.figure2" httpUri="https://binary.pensoft.net/fig/512600" pageId="0" pageNumber="93">2A-C</figureCitation>
), the broad, ovoid to diamond-shaped glabrous patch on the metaventrite (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.669.11773" author="Kohlenberg, AT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="93" pagination="113 - 146" refId="B3" refString="Kohlenberg, AT, Short, AEZ, 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short and Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113 - 146, DOI: https://doi.org/10.3897/zookeys.669.11773" title="Revision of the Neotropical water scavenger beetle genus Tobochares Short and Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)." url="https://doi.org/10.3897/zookeys.669.11773" volume="669" year="2017">Kohlenberg and Short 2017</bibRefCitation>
: fig. 10), the dense and uniform distribution of the hydrofuge pubescence on the abdominal ventrites (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.669.11773" author="Kohlenberg, AT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="93" pagination="113 - 146" refId="B3" refString="Kohlenberg, AT, Short, AEZ, 2017. Revision of the Neotropical water scavenger beetle genus Tobochares Short and Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae). ZooKeys 669: 113 - 146, DOI: https://doi.org/10.3897/zookeys.669.11773" title="Revision of the Neotropical water scavenger beetle genus Tobochares Short and Garcia, 2007 (Coleoptera, Hydrophilidae, Acidocerinae)." url="https://doi.org/10.3897/zookeys.669.11773" volume="669" year="2017">Kohlenberg and Short 2017</bibRefCitation>
: fig. 13A), and the moderate (in number and size) metatibial spines.
</paragraph>
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<subSubSection pageId="0" pageNumber="93" type="composition">
<paragraph pageId="0" pageNumber="93">Composition.</paragraph>
<paragraph pageId="0" pageNumber="93">
This species group includes the following species: 
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<emphasis italics="true" pageId="0" pageNumber="93">Tobochares benettii</emphasis>
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sp. nov., 
<taxonomicName lsidName="T. canaliculatus" pageId="0" pageNumber="93" rank="species" species="canaliculatus">
<emphasis italics="true" pageId="0" pageNumber="93">T. canaliculatus</emphasis>
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Kohlenberg &amp; Short, 2017, 
<taxonomicName lsidName="T. goias" pageId="0" pageNumber="93" rank="species" species="goias">
<emphasis italics="true" pageId="0" pageNumber="93">T. goias</emphasis>
</taxonomicName>
sp. nov., 
<taxonomicName lsidName="T. kasikasima" pageId="0" pageNumber="93" rank="species" species="kasikasima">
<emphasis italics="true" pageId="0" pageNumber="93">T. kasikasima</emphasis>
</taxonomicName>
Short, 2013, 
<taxonomicName lsidName="T. kusad" pageId="0" pageNumber="93" rank="species" species="kusad">
<emphasis italics="true" pageId="0" pageNumber="93">T. kusad</emphasis>
</taxonomicName>
Kohlenberg &amp; Short, 2017, 
<taxonomicName lsidName="T. pallidus" pageId="0" pageNumber="93" rank="species" species="pallidus">
<emphasis italics="true" pageId="0" pageNumber="93">T. pallidus</emphasis>
</taxonomicName>
Kohlenberg &amp; Short, 2017, 
<taxonomicName lsidName="T. sipaliwini" pageId="0" pageNumber="93" rank="species" species="sipaliwini">
<emphasis italics="true" pageId="0" pageNumber="93">T. sipaliwini</emphasis>
</taxonomicName>
Short &amp; Kadosoe, 2011, 
<taxonomicName lsidName="T. striatus" pageId="0" pageNumber="93" rank="species" species="striatus">
<emphasis italics="true" pageId="0" pageNumber="93">T. striatus</emphasis>
</taxonomicName>
Short, 2013, and 
<taxonomicName lsidName="T. sulcatus" pageId="0" pageNumber="93" rank="species" species="sulcatus">
<emphasis italics="true" pageId="0" pageNumber="93">T. sulcatus</emphasis>
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Short &amp; 
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, 2007; see Table 
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Checklist of Tobochares species, their assigned species group, and known distribution. Asterisks (*) denote new country records for previously described species." httpUri="http://table.plazi.org/id/84B03C95B2FBCE7F53225ACCB19930A4" pageId="0" pageNumber="93" tableUuid="84B03C95B2FBCE7F53225ACCB19930A4">2</tableCitation>
.
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