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<document id="7A7001B8320EC16BB52E1687CD1B1467" ID-DOI="10.3897/phytokeys.246.125734" ID-publisher-id="125734" URI-arpha="6960A909-87C0-550E-A5A6-74B9B7185F58" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1726497190775" checkinUser="pensoft" docAuthor="Figueira, Maurício, Schindler, Bianca, Onofre de Araujo, Andréa, Chautems, Alain, Perret, Mathieu, da Silva, Nílber Gonçalves, Alves, Ruy José Válka &amp; Simon, Marcelo Fragomeni" docDate="2024" docId="B47757F37AAA5BB4B5C0AFA18DC9FF20" docLanguage="en" docName="PhytoKeys 246: 265-276" docOrigin="PhytoKeys 246" docSource="https://phytokeys.pensoft.net/article/125734/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.phytokeys" docStyleName="PensoftTaxPub.0000.journal_article.phytokeys" docTitle="Goyazia villosa R. A. Howard" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="276" masterDocId="6960A90987C0550EA5A674B9B7185F58" masterDocTitle="George Gardner’s enigmatic species Goyazia villosa (Gesneriaceae) finally rediscovered" masterLastPageNumber="276" masterPageNumber="265" pageNumber="265" updateTime="1726498837381" updateUser="admin">
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<mods:title id="FDBB3FBCB61E6DD5B2C61B72EA0ED41D">George Gardner’s enigmatic species Goyazia villosa (Gesneriaceae) finally rediscovered</mods:title>
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<mods:namePart id="4A483B6C733363F5A7634A03EFB220CB">Figueira, Maurício</mods:namePart>
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<mods:affiliation id="51006F1EBC78C80E2C7B884D57A7EB36">Programa de Pós-graduação em Botânica, Departamento de Botânica, Universidade de Brasília, 70919 - 970, Brasília, DF, Brazil</mods:affiliation>
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<mods:namePart id="50F70085FF724CFFDEA112AABBCEEF96">Schindler, Bianca</mods:namePart>
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<mods:affiliation id="C5D3ECA65801C932FEEC1B29AA83FBED">Programa de Pós-graduação em Botânica, Departamento de Botânica, Universidade de Brasília, 70919 - 970, Brasília, DF, Brazil</mods:affiliation>
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<mods:namePart id="D3B84FAC6FA3DEDD8242AC111B9C84BF">Onofre de Araujo, Andréa</mods:namePart>
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<mods:affiliation id="BFC42A2D36C73C69223AA46D0AC0D263">Universidade Federal de São Carlos, Rodovia João Leme dos Santos (SP- 264), km 110, Itinga, 18052 - 780, Sorocaba SP, Brazil</mods:affiliation>
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<mods:affiliation id="037D0ECDE46955D8EE0B50CBBCCFD92A">Conservatoire &amp; Jardin botaniques de Genève, 1292 Chambésy, Switzerland &amp; Department of Plant Sciences, University of Geneva, 1292 Chambésy, Switzerland</mods:affiliation>
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<mods:namePart id="36AE747E31451D898B613F8031766022">da Silva, Nílber Gonçalves</mods:namePart>
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<mods:affiliation id="80E610FFCB07E379C73C1671D0AC7BFC">Universidade Federal do Rio de Janeiro, Museu Nacional, Departamento de Botânica. Quinta da Boa Vista s / n, 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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<treatmentCitation id="71CB084502E63FACA8F47A7D4BB42A78" author="R. A. Howard" issue="3" journal="J. Arnold Arbor." page="367" score="9.270020995939838" volume="56" year="1975">
<taxonomicName id="5D37E983E067CB40D32F222C61DE8505" ID-CoL="3H4Z5" authority="(Gardner) R. A. Howard" authorityName="R. A. Howard" baseAuthorityName="Gardner" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="A3F7AB3BF736E3FE6537D02BEABCA7CB" italics="true">Goyazia villosa</emphasis>
(Gardner) R. A. Howard
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, J. Arnold Arbor. 56 (3): 367. 1975
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.
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≡ 
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<emphasis id="C69CB01043ADA5B6B4AE0FFD033137A3" italics="true">Tapina villosa</emphasis>
Gardner.
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Type: 
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. Goyaz [
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]: in dry clefts of rocks near the summit of the Serra de Natividade, 
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, 
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. Gardner 3875
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000509791 image!; 
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<heading id="9FF286BD779AC983D8967DBA1CD68B08" reason="title">Description.</heading>
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Herb, rupicolous, with a perennial scaly rhizome at the base of the roots, stem 
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long, erect, unbranched, green 
<emphasis id="5495963C107F52C9A1EB6ECC17ABDB33" italics="true">in vivo</emphasis>
, villous. Leaves in basal pseudo-rosette or opposite and arranged in 2–5 pairs along the stem, anisophyllous, internodes 
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long, petiole 
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long, terete, green, villous, blade 1–3.5 × 
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, ovate-elliptic or elliptic, symmetrical, membranaceous, pubescent on both faces, tector trichomes unicellular or multicellular uniseriate, base obtuse or cuneate, apex acute or acuminate, margin serrate with 5–14 teeth, lateral veins 4–5 per side. Inflorescence composed of a single flower on leaf axil, pedicel 
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long, erect, villous; sepals subequal, 3–4 × 
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, linear-lanceolate, subulate at the apex, villous; corolla tubular-infundibuliform, pseudo-actinomorphic, 
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long, white at base, tube dark purple, lobes 5, 
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long, subequal, marked with 3 lines of purple and faint dots, with white towards the apex, margin subdentate, throat with wine red dots; stamens 4, filaments glabrous, anthers coherent, nectary dehiscent, annular; ovary superior, ovoid, villous, style 
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long, villous at base, glabrous at the apex, stigma stomatomorph. Fruit a dry loculicidal capsule, dehiscent at the apex, seeds black.
</paragraph>
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<paragraph id="523E4A3533BCA92E359B698E9850035E">
<heading id="F5D4E18B62A073346E208EAAE647106A" reason="title">Additional specimens examined.</heading>
</paragraph>
<paragraph id="859A5712D78BC4DFF6A7CDFAF61849B7">
<materialsCitation id="34FEA8D4FFED3530280BDE81D6C53EBF" collectingDate="2017-04-05" collectionCode="CEN, R" collectorName="R. J. V. Alves &amp; N. G. Silva" country="Brazil" elevation="295" latitude="-9.981944" location="Serra do Lajeado" longLatPrecision="21" longitude="-48.316666" municipality="Palmas" specimenCount="1" stateProvince="Tocantins">
<collectingCountry id="5494ED3D4297527C1254B18438851F1A" name="Brazil">Brazil</collectingCountry>
• 
<collectingRegion id="C1019FE7E83A660BDD6369E348975FB7" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
, 
<collectingMunicipality id="4F4AF02763CFAF44FC32BE44A3A50815">Palmas</collectingMunicipality>
, 
<location id="6BB706A934345E92E395FBED901C8E73" LSID="urn:lsid:plazi:treatment:B47757F37AAA5BB4B5C0AFA18DC9FF20:6BB706A934345E92E395FBED901C8E73" country="Brazil" latitude="-9.981944" longLatPrecision="21" longitude="-48.316666" municipality="Palmas" name="Serra do Lajeado" stateProvince="Tocantins">Serra do Lajeado</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EDGAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-48.316667,-9.981944]}">
<geoCoordinate id="8BFD7B0F3FBE4DC3D3F91DD07030B161" degrees="09" direction="south" minutes="58" orientation="latitude" precision="15" seconds="55" value="-9.981944">09 ° 58 ' 55 &quot; S</geoCoordinate>
, 
<geoCoordinate id="5D3AD4AB678AB9951AABB6CAAD32BDA0" degrees="48" direction="west" minutes="19" orientation="longitude" precision="15" seconds="00" value="-48.316666">48 ° 19 ' 00 &quot; W</geoCoordinate>
</named-content>
; 
<quantity id="983CDE08362FAA7740E9CE47AAEA3C26" metricMagnitude="2" metricUnit="m" metricValue="2.95" unit="m" value="295.0">
<elevation id="4DC51B44CDC64838A204D1C67E115F35" metricMagnitude="2" metricUnit="m" metricValue="2.95" unit="m" value="295.0">295 m</elevation>
</quantity>
, fl.; 
<date id="DF264DCDB119A53F45FDE9EEAFFA8282" value="2017-04-05">
<collectingDate id="82B66B5A0E9FB715DB3C1C109D6B21E9" value="2017-04-05">5 Apr 2017</collectingDate>
</date>
; 
<emphasis id="036070B25526A0913CBF259D2DA34967" italics="true">
<collectorName id="823976BCC39FFCBE5BDC8D44A0B95D28">R. J. V. Alves</collectorName>
&amp; 
<collectorName id="57EBB77DDE57A1CCE97D7A3809940916">N. G. Silva</collectorName>
12586
</emphasis>
(
<collectionCode id="922045FC424EF8A2AEC59BF9D9BDC089" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" type="Herbarium">CEN</collectionCode>
, 
<collectionCode id="6E659449C1B63EFF6D2285798810AC94" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
)
</materialsCitation>
; • 
<materialsCitation id="1D73E486D707F99B417EFC85AC103183" collectingDate="2021-03-02" collectionCode="CEN, RB" collectorName="B. Schindler &amp; M. Figueira &amp; M. F. Simon &amp; V. F. Gomes &amp; W. B. Silva" country="Brazil" county="Ponte Alta do Tocantins" elevation="467" latitude="-11.156111" location="Cachoeira da Fumaca" longLatPrecision="21" longitude="-47.011944" municipality="ESEC Serra Geral do Tocantins" specimenCount="1" stateProvince="Tocantins">
<collectingRegion id="1C7CAB79BECA5B3F916DF945120B0656" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
, 
<collectingCounty id="47F82AEA157FD14111E3FEFFCD9081D4">Ponte Alta do Tocantins</collectingCounty>
, 
<collectingMunicipality id="4304927AFB4233BFB58BBFEE3A4ED15D">ESEC Serra Geral do Tocantins</collectingMunicipality>
, 
<collectingMunicipality id="0F981FF2F866552371E7E4B66AD09E32">rio das Balsas</collectingMunicipality>
, 
<location id="AA3496D0A952093958FD4553F98E6075" LSID="urn:lsid:plazi:treatment:B47757F37AAA5BB4B5C0AFA18DC9FF20:AA3496D0A952093958FD4553F98E6075" country="Brazil" county="Ponte Alta do Tocantins" latitude="-11.156111" longLatPrecision="21" longitude="-47.011944" municipality="ESEC Serra Geral do Tocantins" name="Cachoeira da Fumaca" stateProvince="Tocantins">Cachoeira da Fumaça</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ENGAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.011944,-11.156111]}">
<geoCoordinate id="6D973947EA990AAF17BC843BC8048B1C" degrees="11" direction="south" minutes="09" orientation="latitude" precision="15" seconds="22" value="-11.156111">11 ° 09 ' 22 &quot; S</geoCoordinate>
, 
<geoCoordinate id="42BB053522978F2ED1E1964A8D63C6FA" degrees="47" direction="west" minutes="00" orientation="longitude" precision="15" seconds="43" value="-47.011944">47 ° 00 ' 43 &quot; W</geoCoordinate>
</named-content>
; 
<quantity id="DF3D3854CF9C5C3C013F43AE7570905C" metricMagnitude="2" metricUnit="m" metricValue="4.67" unit="m" value="467.0">
<elevation id="995E0CB6084725842AF9BAA694E09E82" metricMagnitude="2" metricUnit="m" metricValue="4.67" unit="m" value="467.0">467 m</elevation>
</quantity>
, fl. and fr.; 
<date id="F11B8E6D393FAD21BC735D0AD4D53916" value="2021-03-02">
<collectingDate id="9FD0454B3481B9C3C3C0F04DE634A538" value="2021-03-02">2 Mar 2021</collectingDate>
</date>
; 
<emphasis id="AEBC3B520E499AF2C990205418B07E5F" italics="true">
<collectorName id="3A1183C7588C4595B0957ED9276BEB8A">B. Schindler</collectorName>
</emphasis>
, 
<emphasis id="C7253B08FD51565757CF29DB0BEBA7E8" italics="true">
<collectorName id="CB048F6DBEC00D70F73DE0B8DD741D55">M. Figueira</collectorName>
</emphasis>
, 
<emphasis id="00D5DF68BF516C5FB8B7965E36414384" italics="true">
<collectorName id="AE121422655180F92F24B5488A530DD4">M. F. Simon</collectorName>
</emphasis>
, 
<emphasis id="E199D593C09497A78C9F0C87F1B212E1" italics="true">
<collectorName id="05C896942150A987E47EE9E8D0C9040B">V. F. Gomes</collectorName>
</emphasis>
, 
<emphasis id="B9433BFA6A23C4A1461039CBA689307E" italics="true">
<collectorName id="50731C034D53A9AF572B6037B020D7E9">W. B. Silva</collectorName>
41
</emphasis>
(
<collectionCode id="3A74C7C9052033A59C3EE0BF3D20CB2E" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" type="Herbarium">CEN</collectionCode>
, 
<collectionCode id="93578F4EE3B8B854DD551C389F66A8BB" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" type="Herbarium">RB</collectionCode>
)
</materialsCitation>
; • 
<materialsCitation id="4DC26C9FF13A236758C3FE3659DD2382" collectingDate="2022-03-15" collectionCode="CEN, G, R, RB, UB" collectorName="M. Figueira &amp; B. Schindler &amp; M. F. Simon &amp; R. R. Souza &amp; V. F. Gomes &amp; W. B. Silva" country="Brazil" county="Ponte Alta do Tocantins" elevation="465" latitude="-11.156111" location="Cachoeira da Fumaca" longLatPrecision="21" longitude="-47.0125" municipality="Ponte Alta" specimenCode="G 00447873" specimenCount="1" stateProvince="Tocantins">
idem; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6GAE" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-47.012500,-11.156111]}">
<geoCoordinate id="765142BF189C3BE8FA8AE4FDC7258DEA" degrees="11" direction="south" minutes="09" orientation="latitude" precision="15" seconds="22" value="-11.156111">11 ° 09 ' 22 &quot; S</geoCoordinate>
, 
<geoCoordinate id="39687D794648F6D98DDEDDCC12C5B434" degrees="47" direction="west" minutes="00" orientation="longitude" precision="15" seconds="45" value="-47.0125">47 ° 00 ' 45 &quot; W</geoCoordinate>
</named-content>
; 
<quantity id="E91DB2FAC5A3DAD35389BCDBB5232169" metricMagnitude="2" metricUnit="m" metricValue="4.65" unit="m" value="465.0">
<elevation id="8C92A66A1BD323DCEFA8CA502CFCF85D" metricMagnitude="2" metricUnit="m" metricValue="4.65" unit="m" value="465.0">465 m</elevation>
</quantity>
, fl.; 
<date id="D9BA54168C89E2450B2BA55AB11BAB75" value="2022-03-15">
<collectingDate id="2CC109C601C6C36483E25FF6345EF941" value="2022-03-15">15 Mar 2022</collectingDate>
</date>
; 
<emphasis id="FCBC0933E573DEBD61149268A4BF72F4" italics="true">
<collectorName id="AEF8889A732FB34EAEBFC2B5111BD7B4">M. Figueira</collectorName>
</emphasis>
, 
<emphasis id="B5D1A3DED5928D6C334E0B469420DFC7" italics="true">
<collectorName id="26F5839AE7373402D9D461AAEAD72F37">B. Schindler</collectorName>
</emphasis>
, 
<emphasis id="CA6EC3E0169D90C00F4F3D4D5C0DC480" italics="true">
<collectorName id="44FFEB7426BFB7888288AF10B331BEA3">M. F. Simon</collectorName>
</emphasis>
, 
<emphasis id="95FB9557D2402B8938C96B692F084C38" italics="true">
<collectorName id="6DF9A4FFD188227C67610BEF79CB130F">R. R. Souza</collectorName>
</emphasis>
, 
<emphasis id="8E762D01825F8D10A0820F8B843638BD" italics="true">
<collectorName id="D79ECD0D58F98D57BC170665CBB3C689">V. F. Gomes</collectorName>
</emphasis>
, 
<emphasis id="A014D125E8F51D2BDEB6D6DB4B52974F" italics="true">
<collectorName id="5BD3E35E3C337B24515AFD257587EA58">W. B. Silva</collectorName>
1808
</emphasis>
(
<collectionCode id="7D3DCE63811FD9B74633E8DE9F7D9646" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" type="Herbarium">CEN</collectionCode>
, 
<collectionCode id="EA947737C317F0C763C801D2DF5A336A" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G</collectionCode>
[barcode] 
<specimenCode id="410BE6E0CCB292BDFE237F731D2BB3ED" collectionCode="G" country="Switzerland" lsid="urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Genève" type="Herbarium">G 00447873</specimenCode>
, 
<collectionCode id="2E722860EABDD309B17DAD45939A884E" country="Chile" name="Departamento de Geologia, Universidad de Chile">R</collectionCode>
, 
<collectionCode id="CAA68F7787A73133FAF6F65A3F565FF6" country="Brazil" lsid="urn:lsid:biocol.org:col:15528" name="Jardim Botânico do Rio de Janeiro" type="Herbarium">RB</collectionCode>
, 
<collectionCode id="CBC08B435ECBDC31AB8EA8A29302FBCF" country="France" name="Laboratoire de Biostratigraphie">UB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EQHAE" type="distribution">
<paragraph id="353743AAD73406928445A2F26AA21FCB">
<heading id="99ECCA38B8BD945F9AE0CEB8D25503D4" reason="title">Distribution and habitat.</heading>
</paragraph>
<paragraph id="C62AD0CE983FC2D2354969898F2AF7E0">
The species is endemic to the state of 
<collectingRegion id="6E268D85919CA4907C9F42E054045C23" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
, 
<collectingCountry id="69EFE0F6889BFEBD186613B73EDA21B8" name="Brazil">Brazil</collectingCountry>
(Fig. 
<figureCitation id="694900524F11BAB86A700BE56861AEE7" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Location of the type collection of Goyazia villosa made in 1840 (circle) and new records made in the municipality of Palmas in 2017 (diamond) and Estação Ecológica Serra Geral do Tocantins (EESGT) in 2021 and 2022 (triangle). Brazilian states: BA (Bahia), MA (Maranhão), PI (Piauí), TO (Tocantins). Track of G. Gardner based on the Atlas dos Viajantes no Brasil (https://viajantes.bbm.usp.br/). Map produced with QGIS (QGIS Development Team 2024)." figureDoi="10.3897/phytokeys.246.125734.figure1" httpUri="https://binary.pensoft.net/fig/1134771">1</figureCitation>
). It has been recorded in the municipalities of Natividade, Palmas, and Ponte Alta do 
<collectingRegion id="971C86C72ECDB8F1A7E7CFE3FC8C40C4" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
. Recent records were made in a humid and shaded hillside, in the Serra do Lajeado, at around 
<quantity id="BC73985FB76F408F7F49475B964FC06C" metricMagnitude="2" metricUnit="m" metricValue="2.95" unit="m" value="295.0">295 m</quantity>
of altitude (
<emphasis id="C680ABFE3AAECF0820DEAAB1D4FDCC21" italics="true">R. J. V. Alves &amp; N. G. Silva 12586</emphasis>
), and also in a riparian forest on shaded humid sandstone walls in the locality known as Cachoeira da Fumaça, in the southwest corner of the 
<abbrev id="ABBRID0E3HAE" xlink_title="Estação Ecológica Serra Geral do Tocantins">
<collectionCode id="CEC9E3AD942A411C81C048E622BAFF43">EESGT</collectionCode>
</abbrev>
, at around 
<quantity id="8EB79205ED8BE8D8D273E5D8E0807F22" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</quantity>
of altitude (
<emphasis id="2554B733305B17276FDFED993533914F" italics="true">B. Schindler et al. 41 and M. Figueira et al. 1808</emphasis>
) (Fig. 
<figureCitation id="D16FF15FB210D6485F4CCE786810FA50" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitat and habit of Goyazia A Cachoeira da Fumaça at the Estação Ecológica Serra Geral do Tocantins B habitat of G. villosa on rocky outcrops C frontal view of G. villosa attached to sandstone rocks D lateral view of individuals of G. petraea E, F habit of G. rupicola. Photographs: A – C, E M. Figueira and B. Schindler D and F A. O. Araujo; photos from collections: B, C M. Figueira et al. 1808 D A. O. Araujo et al. 1065-24 E M. Figueira et al. 1897 F A. O. Araujo et al. 1082." figureDoi="10.3897/phytokeys.246.125734.figure3" httpUri="https://binary.pensoft.net/fig/1134773">3 A – C</figureCitation>
). Description of the habitat by Gardner on the specimen label indicates that the plant was collected “ in dry clefts of rocks near the summit of the Serra de Natividade ”. In contrast, our field observations suggest that 
<taxonomicName id="9D5A50F4C20D43877A0D1875166FC8CA" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="1ADAED31EF23642B7F1599AE8A637A6A" italics="true">Goyazia villosa</emphasis>
</taxonomicName>
grows in humid and shaded sites. Preference of 
<taxonomicName id="D077BE15671B61DB0F018E059084F650" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="BA210AB20E113602653411C2BBB17A4A" italics="true">G. villosa</emphasis>
</taxonomicName>
for drier sites is however not unlikely because of its rhizomes with fleshy scales (Fig. 
<figureCitation id="4CC0246C898AE3410C17EE2981B941BA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Goyazia villosa (Gardner) R. A. Howard A habit B detail of rhizome with fleshy scales C detail of leaf trichomes on the adaxial surface D front view of corolla E lateral view of corolla F detail of sepals, ovary, and ring nectary (corolla removed) G stamen H immature fruit I open capsule. Photographs by M. Figueira and B. Schindler from M. Figueira et al. 1808." figureDoi="10.3897/phytokeys.246.125734.figure2" httpUri="https://binary.pensoft.net/fig/1134772">2 B</figureCitation>
) that allow the plant to become dormant during the dry season, and grow and reproduce during the rainy season. This feature has already been reported for other genera of 
<taxonomicName id="947AB7F6A00EFD27EC65D996182D26AF" rank="subTribe" subTribe="Gloxiniinae">Gloxiniinae</taxonomicName>
(
<bibRefCitation id="637179D3D0E2C636F62E9D89CC2E3910" DOI="10.1002/tax.591019" author="Araujo" etAl="et al." firstAuthor="Araujo" issue="1" journalOrPublisher="Taxon" pagination="203-208" refId="B7" refString="Araújo AO, Souza VC, Perret M (2010) Chautemsia calcicola: A new genus and species of Gloxinieae (Gesneriaceae) from Minas Gerais, Brazil. Taxon 59 (1): 203–208. https://doi.org/10.1002/tax.591019" title="Chautemsia calcicola: A new genus and species of Gloxinieae (Gesneriaceae) from Minas Gerais, Brazil." volume="59" year="2010">Araújo et al. 2010</bibRefCitation>
).
</paragraph>
<caption id="C2D214BFBB9C334609259F786D2F46EB" ID-DOI="10.3897/phytokeys.246.125734.figure2" ID-arpha="103EB178-907A-599A-A428-E2DB4C620D1C" httpUri="https://binary.pensoft.net/fig/1134772" startId="F2">
<paragraph id="84AEF8B5259FCCB397AD7AAD578FE6FE">
<label id="1F9FB0DDCDF2631A3F4E1C96DF3E93B1">Figure 2.</label>
</paragraph>
<paragraph id="1759701AA78306B6D801193DADB3F86E">
<taxonomicName id="E4059236A010AC169B121E0BE8CD1980" authority="(Gardner) R. A. Howard A" authorityName="R. A. Howard A" baseAuthorityName="Gardner" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="DA478FB905BA0EB01239D4A823AA16CD" italics="true">Goyazia villosa</emphasis>
(Gardner) R. A. Howard 
<emphasis id="4A1F8B4C977A1B7323B55C79D6AA5CD6" bold="true">A</emphasis>
</taxonomicName>
habit 
<emphasis id="315B3DFB1729432489937A2DEFF9CF35" bold="true">B</emphasis>
detail of rhizome with fleshy scales 
<emphasis id="5B912F4AA1198FCA1891E441B44650BD" bold="true">C</emphasis>
detail of leaf trichomes on the adaxial surface 
<emphasis id="DA185AFE674C74C8559CB7CF242BCE7E" bold="true">D</emphasis>
front view of corolla 
<emphasis id="67BF3AA36D92065B92AAAC5FB2C8B966" bold="true">E</emphasis>
lateral view of corolla 
<emphasis id="5F96E9AAD53D2936721C74757D5019B1" bold="true">F</emphasis>
detail of sepals, ovary, and ring nectary (corolla removed) 
<emphasis id="D85CDC38B997C733451D7BA6A1130CAB" bold="true">G</emphasis>
stamen 
<emphasis id="B5F5E699E4962C73F1094C68401DB771" bold="true">H</emphasis>
immature fruit 
<emphasis id="E4F8536A90F11A4EED54398FECE374F5" bold="true">I</emphasis>
open capsule. Photographs by M. Figueira and B. Schindler from M. Figueira et al. 1808.
</paragraph>
</caption>
<caption id="3F8D1AB68FB8A3842E8C2FFA9E5E875F" ID-DOI="10.3897/phytokeys.246.125734.figure3" ID-arpha="D2714A06-A991-5988-878E-4606A1B1561C" httpUri="https://binary.pensoft.net/fig/1134773" startId="F3">
<paragraph id="5C8B9D9C59E51B79C6B7AE5003EB1EDB">
<label id="12093231298CD520923ADF8769B9BA08">Figure 3.</label>
</paragraph>
<paragraph id="E591CA6BB86C764AEC54CA811D27068B">
Habitat and habit of 
<taxonomicName id="56D5C357A4C1C226E73DA2ABD06D05A1" authority="A Cachoeira da Fumaca" authorityName="A Cachoeira da Fumaca" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="genus">
<emphasis id="99A15DD9960360FA9DC8F948BFBCB82F" italics="true">Goyazia</emphasis>
<emphasis id="656FE43118E596B6080D57113ADF03B7" bold="true">A</emphasis>
Cachoeira da Fumaça
</taxonomicName>
at the Estação Ecológica Serra Geral do Tocantins 
<emphasis id="2693BD974892E0E9B00C164D1D0C4130" bold="true">B</emphasis>
habitat of 
<taxonomicName id="A74CCDC056E4893E3EAB4FCA3F3E984D" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="E589149CFF47D001973FE5479E2E6848" italics="true">G. villosa</emphasis>
</taxonomicName>
on rocky outcrops 
<emphasis id="6FD0648D2670304302F32769AF3856CD" bold="true">C</emphasis>
frontal view of 
<taxonomicName id="D8250B0E2176A1C9BD95FFF3C576297B" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="1FFADAFD2F7467390E897E5DDC872C28" italics="true">G. villosa</emphasis>
</taxonomicName>
attached to sandstone rocks 
<emphasis id="ACF38A8B76A3B629EF7FE6EB867355FB" bold="true">D</emphasis>
lateral view of individuals of 
<taxonomicName id="F45929830AE8C4294EF7C1C03D4F8449" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="petraea">
<emphasis id="E9EFC7537F27A99B51A910569757E3E8" italics="true">G. petraea</emphasis>
</taxonomicName>
<emphasis id="AC71F977F11BE38B2C6E0E725BCA1837" bold="true">E, F</emphasis>
habit of 
<taxonomicName id="185DB4921296CE9BB8469F42644C039F" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="rupicola">
<emphasis id="A298F6E57B81906B7AD2773D3398FEAF" italics="true">G. rupicola</emphasis>
</taxonomicName>
. Photographs: 
<emphasis id="4255D73E7C92DF8619D9CF69CE21A700" bold="true">A – C</emphasis>
, 
<emphasis id="A56624FA28AF8F96A02DED9FA8CDB194" bold="true">E</emphasis>
M. Figueira and B. Schindler 
<emphasis id="6A85FC6597A0B4AFF0A544BDFACD53DC" bold="true">D</emphasis>
and 
<emphasis id="009CBCB30199135D2A8FD082C42DA5C6" bold="true">F</emphasis>
A. O. Araujo; photos from collections: 
<emphasis id="185F7AD505B7FA9DBADC0FDF416D2C36" bold="true">B, C</emphasis>
M. Figueira et al. 1808 
<emphasis id="3CDC9638F0CDABB095DB8CCC528ABF02" bold="true">D</emphasis>
A. O. Araujo et al. 1065-24 
<emphasis id="548FE407D1CC8BDDD3CF480ECB52A01C" bold="true">E</emphasis>
M. Figueira et al. 1897 
<emphasis id="1BE936707A74CD655DAF8062BB9EEFB1" bold="true">F</emphasis>
A. O. Araujo et al. 1082.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0ENNAE" type="phenology">
<paragraph id="E9E8CE58E143148CC2C587267B896787">
<heading id="35420830DC463FB3EAC69D31528582B5" reason="title">Phenology.</heading>
</paragraph>
<paragraph id="55A898941AC3A7B34739B0CB0FBCC8EE">Flowers and fruits between December and April during the rainy season.</paragraph>
</subSubSection>
<subSubSection id="SECID0ESNAE" type="etymology">
<paragraph id="2BFE7BCD59D8598212D3000B5C23EE1F">
<heading id="C47D66A540DAD335727A66BDD3477CF0" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="CFBA421897A8DE34FF8A42F20018B6B9">
The genus name is a reference to the state of 
<collectingRegion id="D25C7C33237B8FAAA8CA0496A7427E42" country="Brazil" name="Goias">Goiás</collectingRegion>
(formerly spelled as Goyaz in old Portuguese orthography). The plant was collected in an area that now belongs to the state of 
<collectingRegion id="20BD8B42372C680621AA8A0E01E1DCCE" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
. The specific epithet refers to the trichomes found throughout the plant.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EXNAE" type="conservation status">
<paragraph id="4E850ECDE3CCC5B4ABFD195745D0D18B">
<heading id="ABDF1AA0C56967FE117EBE230A308C67" reason="title">Conservation status assessment.</heading>
</paragraph>
<paragraph id="52B00445B2FDAC01A863FC7B537F8444">
<taxonomicName id="C6E5B039234A82FA73BBE36C6786285B" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="E7EB50807639F6927E6241B7D60EB6BF" italics="true">Goyazia villosa</emphasis>
</taxonomicName>
is currently known from only three localities, where it inhabits a very specific habitat. It has an 
<abbrev id="ABBRID0EIOAE" xlink_title="Area of Occupancy">AOO</abbrev>
of 
<quantity id="F8D343847FBFFD248B69792AC67EB3EB" metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
<superScript id="6E4ED978CA647260022909F2C5CE323C">2</superScript>
and 
<abbrev id="ABBRID0EOOAE" xlink_title="Extent of Occurrence">EOO</abbrev>
<quantity id="F794CF6E950CD21D78EFB31EF41288C9" metricMagnitude="6" metricUnit="m" metricValue="9.10344" unit="km" value="9103.44">9103.44 km</quantity>
<superScript id="70686E46B602DCA6DDD399390F13C9DA">2</superScript>
. Analysis of land use within 
<taxonomicName id="7FAABE4A0ACED26A4104ED5F1BD06851" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="EE11E97527F602368B364607903C5436" italics="true">G. villosa</emphasis>
</taxonomicName>
range (
<abbrev id="ABBRID0E6OAE" xlink_title="Extent of Occurrence">
<collectionCode id="06C2C84B9C509ABE3E9AAAA614F0D015">EOO</collectionCode>
</abbrev>
) revealed a reduction of 
<quantity id="FE6DE2A55842DBF24D4D29328A796C6F" metricMagnitude="6" metricUnit="m" metricValue="1.616" unit="km" value="1616.0">1616 km</quantity>
<superScript id="11A6BA6FBC34C6777D69368CFDE93440">2</superScript>
over 35 years, which represents an increment of anthropic areas from 9.0 % in 1986 to 17.7 % in 2021. Considering its 
<abbrev id="ABBRID0EFPAE" xlink_title="Area of Occupancy">AOO</abbrev>
&lt;
<quantity id="753987907182013A9EEA40BA1E4088AF" metricMagnitude="5" metricUnit="m" metricValue="5.0" unit="km" value="500.0">500 km</quantity>
<superScript id="43F92C4F66F7A66DD8F0310A4666D339">2</superScript>
, number of locations ≤ 5, and continuing decline in extent of occurrence, we preliminary assign a category of endangered (EN) for 
<taxonomicName id="A5DF19AD32C638A4E6F82F5BE18923C2" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="08B7193820EE7E721AC800074402B3D2" italics="true">G. villosa</emphasis>
</taxonomicName>
under IUCN criterion B 2 ab (i).
</paragraph>
<paragraph id="FC9BF1B3763C8885CCDBB545A40A33E1">
Although known from only three localities and inhabiting a very specific habitat, 
<taxonomicName id="F30A50B73485A26A794A751FFF6D1BD7" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="B8156C0D8E8A78ECAE1B6A434305B657" italics="true">G. villosa</emphasis>
</taxonomicName>
occurs in a region that still harbors much of its native vegetation, despite the observed vegetation loss over its range. The two newly reported populations are between 100 and 
<quantity id="D4137B85DC19E2FCFCEEEC6ADE7DA115" metricMagnitude="5" metricUnit="m" metricValue="2.0" unit="km" value="200.0">200 km</quantity>
from the type collection. The area covered within its current range harbors a number of unexplored sites of potential occurrence of 
<taxonomicName id="A0530D3980E47510439BEFAE2168708C" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="44D4BB8FFE614812D5487941364FC474" italics="true">G. villosa</emphasis>
</taxonomicName>
, including several mountain ranges, which could increase its 
<abbrev id="ABBRID0EOQAE" xlink_title="Area of Occupancy">AOO</abbrev>
. Recently documented habitats of 
<taxonomicName id="B5834B1C9FA84964EE4E6DD3CF14C170" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="0A951953409628DE5CA0CDA7BE37DE92" italics="true">G. villosa</emphasis>
</taxonomicName>
are either unsuitable for agriculture (rocky outcrops) or protected by environmental regulations (riparian forests), which reduces the chances of loss of this specific habitat. The overall extent of occurrence of 
<taxonomicName id="54B61089C726D166A874D96E7167EFEE" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="59BC2371C9B59DC6CB5D84DED3027759" italics="true">G. villosa</emphasis>
</taxonomicName>
in eastern 
<collectingRegion id="A31C5F72BFD3F5406E0CDC5A3912F789" country="Brazil" name="Tocantins">Tocantins</collectingRegion>
includes a large protected area (
<abbrev id="ABBRID0EIRAE" xlink_title="Estação Ecológica Serra Geral do Tocantins">
<collectionCode id="8AF5C23100AC545AF2654C421AD39EE6">EESGT</collectionCode>
</abbrev>
). One negative aspect in the conservation of this species is that the population found in Cachoeira da Fumaça within the 
<abbrev id="ABBRID0EMRAE" xlink_title="Estação Ecológica Serra Geral do Tocantins">
<collectionCode id="F84309EBF4343A816A3A891DFD571C69">EESGT</collectionCode>
</abbrev>
may suffer from trampling since a patch of individuals of 
<taxonomicName id="96660CE259296CDEFB890259906D888D" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="9864A328D96635ED20F065B7D219D721" italics="true">G. villosa</emphasis>
</taxonomicName>
is located along a trail used by the numerous visitors to the waterfall. The rarity of 
<taxonomicName id="76CD2A0643B614A5B3C239BF99AC278F" class="Magnoliopsida" family="Gesneriaceae" genus="Goyazia" kingdom="Plantae" order="Lamiales" phylum="Tracheophyta" rank="species" species="villosa">
<emphasis id="75AFE4D088B6E850D8F2A05857602E1D" italics="true">G. villosa</emphasis>
</taxonomicName>
is probably a result of strong habitat specificity and insufficient collecting effort across its geographic range, which could explain the paucity of records. Finally, more sampling is needed to understand its entire geographical range.
</paragraph>
</subSubSection>
</treatment>
</document>