treatments-xml/data/A7/E8/B1/A7E8B110FB875A6CBAAC65042B2B7CE5.xml
2024-06-21 12:46:51 +02:00

264 lines
27 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="A7E8B110FB875A6CBAAC65042B2B7CE5" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho titschacki Roewer 1949" docType="treatment" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>80</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>309</mods:start>
<mods:end>388</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/asp.80.e73829</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.80.e73829</mods:identifier>
<mods:identifier type="Pensoft-Pub">1864-8312-80-309</mods:identifier>
<mods:identifier type="ZooBank">D5C0468B99A14EF39237D9BC51A8BDA3</mods:identifier>
<mods:identifier type="Pensoft-UUID">11907D1B6F665D96B5FD2F9751431366</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:A7E8B110FB875A6CBAAC65042B2B7CE5" httpUri="http://treatment.plazi.org/id/A7E8B110FB875A6CBAAC65042B2B7CE5" lastPageNumber="309" pageId="0" pageNumber="309">
<subSubSection pageId="0" pageNumber="309" type="nomenclature">
<paragraph pageId="0" pageNumber="309">
3.23.
<taxonomicName LSID="A7E8B110-FB87-5A6C-BAAC-65042B2B7CE5" authority="Roewer, 1949" authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho titschacki" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="titschacki">Ayacucho titschacki Roewer, 1949</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">Figs 3D</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">, 7U, V</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">, 12I, J</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">, 23E</figureCitation>
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="reference_group">
<paragraph pageId="0" pageNumber="309">
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho titschacki" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="titschacki">Ayacucho titschacki</taxonomicName>
Roewer, 1949: 57 (desc.), figs. 114a (dorsal habitus), 114b (ventral habitus), 114c (chelicera, pedipalpus and carapace in lateral view); 114d (detail of coxa IV), 114e (sternum), 114f (chelicera in lateral view);
<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
: 144 (cat.).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="redescription">
<paragraph pageId="0" pageNumber="309">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
MALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=2) DSW: 3.0-4.1; DSL: 4.5-5.4; CL: 1.5-2.2. FIVL: 4.8-5.4. ChL: 2.5-4.0.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
: (Fig.
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">23E</figureCitation>
) Predominantly yellowish, with two longitudinal black stripes in the lateral region of dorsal scutum (carapace and lateral edges of the areas I-IV). Lateral margins of DS yellowish, with small black spots. Posterior margin of the dorsal scutum, free tergites and legs predominantly black. Chelicerae and pedipalpus predominantly dark yellow.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
) Alpha-type DSS. Anterior margin of carapace with median elevation totally covered with granules. Ocularium with two small acuminate tubercles, absent in some individuals. Areas I-IV with two pairs of small median tubercles; the lateral pair can be greatly reduced in some individuals. Posterior margin of dorsal scutum with a transverse row of 4-5 acuminate tubercles (larger than tubercles of areas I-IV). Free tergites I-III with a row of tubercles similar to the posterior margin of dorsal scutum, increasing the number of free tergite I to III.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
) Swollen or not in males. Segment I densely covered with granules throughout its length. Segment II predominantly smooth, with some hairs on the front face; finger with three teeth. Segment III with two teeth.
<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: With very small granules sparsely distributed on the dorsal surface of the femur and patella. Trochanter with a distal ventral setiferous tubercle. Femur with a ventrobasal setiferous tubercle and a row of six small ventromedian setiferous tubercles. Tibia: prolateral IIII, retrolateral IIII. Tarsus: prolateral IIi, retrolateral Iii.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxa I with dispersed small tubercles. Coxae II-III, predominantly smooth, with sparse granules. Coxa IV with granulation denser than coxae III-IV. Genital area smooth. Free sternites I-IV and anal operculum with small granules sparsely distributed.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
,
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7U, V</figureCitation>
) Coxa I with a prolateral apophysis; coxa II with a retrolateral and a prolateral apophysis. Coxa IV densely granulate, with larger tubercles distributed more densely in the prolateral portion. Trochanters I-IV with few granules, unarmed. Femora I-III, with granules distributed throughout its length. Femur IV with granules densely distributed in the dorsal and lateral, less heavily on the ventral surface. Tarsal formula: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=2) 7, 12, 7, 8.
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">12I, J</figureCitation>
) VP rectangular; distal margin slightly concave, with short lateral-apical projections; slightly concave in lateral view. MS C1-C5 (or C6) subapical long and straight; MS A1 median short and straight; MS B1 sub basal long and straight (longer than MS A and shorter than MS C). Lateral sacs short and apically acuminated, with short T3-like microsetae. Stylus slightly thick, apically inflated, dorsally inclined, with apical tiny projections. Dorsal process absent. Promontory convex. -
<emphasis bold="true" pageId="0" pageNumber="309">
FEMALE:
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=4) DSW: 3.0-4.4; DSL: 4.5-5.4; CL: 1.9-4.7. FIVL: 3.7-4.5. ChL: 1.7-4.0. Swollen chelicerae in some specimens, like males. Tubercles of femur of pedipalpus slightly smaller. Ocularium tubercles smaller than in male. Granules on femur IV thinner when compared with those of male. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=4) 6-7, 10-11, 7, 7-8.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="diagnosis">
<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="309">
Similar to
<taxonomicName authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho glauberrochai" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="glauberrochai">
<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho glauberrochai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
,
<taxonomicName genus="A." lsidName="A. pomacocha" pageId="0" pageNumber="309" rank="species" species="pomacocha">
<emphasis italics="true" pageId="0" pageNumber="309">A. pomacocha</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
,
<taxonomicName genus="A." lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
and
<taxonomicName genus="A." lsidName="A. vargasllosai" pageId="0" pageNumber="309" rank="species" species="vargasllosai">
<emphasis italics="true" pageId="0" pageNumber="309">A. vargasllosai</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
in the following combination of characteristics: dorsal scutum densely granulate; ocularium and areas I-IV of DS unarmed or armed with tiny tubercles, slightly greater than granules; posterior margin of DS and free tergites I-III with median rows of acuminate tubercles (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
). It differs from the previously mentioned species in the following combination of characteristics: ocularium with two small acuminate tubercles and densely granulate (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3D</figureCitation>
); male femur IV unarmed (Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7U, V</figureCitation>
; unlike
<taxonomicName genus="A." lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
); penis VP rectangular; five-six pairs of subapical MS C; one pair of MS A and B (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Penis of Metasarcidae, dorsal and lateral views, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D Ayacucho inermis (Roewer, 1957) comb. nov.; E - F Ayacucho pasolinii sp. nov.; G - H Ayacucho spielbergi sp. nov.; I - J Ayacucho titschacki Roewer, 1949; K - L Ayacucho triarmatus nom. nov.; Legend bars = 0.1 mm." figureDoi="10.3897/asp.80.e73829.figure12" httpUri="https://binary.pensoft.net/fig/729886" pageId="0" pageNumber="309">12I, J</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="remarks">
<paragraph pageId="0" pageNumber="309">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="309">The male holotype and paratypes examined by Roewer (SMF RII 8589/20) are actually females.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="distribution">
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="309">
(Fig.
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
) PERU.
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
.
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
, Ocollo and Virgem de Cacharras de Cocha.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
<emphasis italics="true" pageId="0" pageNumber="309">Type material</emphasis>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="PERU" location="PERU" specimenCount="♀" typeStatus="Holotype">
<typeStatus>Holotype</typeStatus>
<specimenCount></specimenCount>
, '
<collectingCountry name="Peru">PERU</collectingCountry>
,
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
,
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
, without date and leg. (SMF RII 8589/20)
</materialsCitation>
</emphasis>
-
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount></specimenCount>
,
<normalizedToken originalValue="ditto">'ditto'</normalizedToken>
(SMF RII 8589/20).
<emphasis bold="true" pageId="0" pageNumber="309">
<emphasis italics="true" pageId="0" pageNumber="309">Additional material</emphasis>
.
</emphasis>
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="PERU" latitude="-13.029722" location="Virgem de Cacharras de Cocha" longLatPrecision="21" longitude="-73.87056" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<specimenCount type="male">1 ♂</specimenCount>
,
<specimenCount type="female">2 ♀</specimenCount>
'
<collectingCountry name="Peru">PERU</collectingCountry>
,
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
,
<location LSID="urn:lsid:plazi:treatment:A7E8B110FB875A6CBAAC65042B2B7CE5:2E96CA51C1AE6E9164E346767C20A6A2" country="PERU" latitude="-13.029722" longLatPrecision="21" longitude="-73.87056" name="Virgem de Cacharras de Cocha">Virgem de Cacharras de Cocha</location>
,
<geoCoordinate degrees="13" direction="south" minutes="01" orientation="latitude" precision="15" seconds="47" value="-13.029722">13°0147″S</geoCoordinate>
<geoCoordinate degrees="73" direction="west" minutes="52" orientation="longitude" precision="15" seconds="14" value="-73.87056">73°5214″W</geoCoordinate>
, without date and leg. (MZSP 36972)
</materialsCitation>
.
<materialsCitation collectingDate="2011-04-29" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, D. Silva" country="PERU" latitude="-13.331388" location="Near Ocollo" longLatPrecision="1" longitude="-74.50489" specimenCount="2" specimenCount-female="2">
<specimenCount type="female">2 ♀</specimenCount>
'
<collectingCountry name="Peru">PERU</collectingCountry>
,
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">Ayacucho</taxonomicName>
,
<location LSID="urn:lsid:plazi:treatment:A7E8B110FB875A6CBAAC65042B2B7CE5:C2719E395FA3B21594C16ABEB1F7B3C0" country="PERU" latitude="-13.331388" longLatPrecision="1" longitude="-74.50489" name="Near Ocollo">Near Ocollo</location>
,
<geoCoordinate degrees="13" direction="south" minutes="19" orientation="latitude" precision="1" seconds="53.0" value="-13.331388">13°1953.0″S</geoCoordinate>
<geoCoordinate degrees="74" direction="west" minutes="30" orientation="longitude" precision="1" seconds="17,6" value="-74.50489">74°3017,6″ W</geoCoordinate>
,
<collectingDate value="2011-04-29">29/IV/2011</collectingDate>
,
<collectorName>R. Pinto-da-Rocha</collectorName>
,
<collectorName>A. Benedetti</collectorName>
,
<collectorName>J. Ochoa</collectorName>
&amp;
<collectorName>D. Silva</collectorName>
leg. (MUSM)
</materialsCitation>
;
<specimenCount type="male">1 ♂</specimenCount>
,
<specimenCount type="female">2 ♀</specimenCount>
,
<normalizedToken originalValue="ditto">'ditto'</normalizedToken>
(MUBI);
<specimenCount type="male">1 ♂</specimenCount>
,
<specimenCount type="female">5 ♀</specimenCount>
<normalizedToken originalValue="ditto">'ditto'</normalizedToken>
(MZSP 36973).
</paragraph>
</subSubSection>
</treatment>
</document>