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<document id="9C8AA5D837E3BC95D7A0E0D41031E0F8" ID-CLB-Dataset="21907" ID-DOI="10.5852/ejt.2017.390" ID-GBIF-Dataset="54438cfa-5f3a-4ee3-85c8-00e453a6d641" ID-ISSN="2118-9773" ID-Zenodo-Dep="3839858" ID-ZooBank="4D1D1CA3-8345-4BA3-9C7C-5EBDD40752CE" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1590155668479" checkinUser="carolina" docAuthor="Wisshak, Max" docDate="2017" docId="8878B758BA369F694E3E23EEFCB9FD05" docLanguage="en" docName="ejt-2017-390.pdf.imf" docOrigin="European Journal of Taxonomy 390" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Pyritonema belua Wisshak 2017, isp. nov." docType="treatment" docVersion="13" lastPageNumber="66" masterDocId="7441CF20BA769F284C53200DFFF9FFC3" masterDocTitle="Taming an ichnotaxonomical Pandoras box: revision of dendritic and rosetted microborings (ichnofamily: Dendrinidae)" masterLastPageNumber="99" masterPageNumber="1" pageNumber="65" updateTime="1698840949990" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CC29C86457FBFD8DA685461CC4D03CC6">Taming an ichnotaxonomical Pandoras box: revision of dendritic and rosetted microborings (ichnofamily: Dendrinidae)</mods:title>
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<mods:title id="313EE4F7928DDF3CA1C803EEFF1DD745">European Journal of Taxonomy</mods:title>
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<mods:date id="1EAA6D9B6228521651AFC2217F52CFCE">2017</mods:date>
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<treatment id="8878B758BA369F694E3E23EEFCB9FD05" ID-DOI="http://doi.org/10.5281/zenodo.3853699" ID-GBIF-Taxon="164249293" ID-Zenodo-Dep="3853699" LSID="urn:lsid:plazi:treatment:8878B758BA369F694E3E23EEFCB9FD05" httpUri="http://treatment.plazi.org/id/8878B758BA369F694E3E23EEFCB9FD05" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
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<paragraph id="006E064EBA369F684E3E23EEFC3EFC3D" blockId="64.[361,1226,995,1093]" box="[621,967,995,1022]" pageId="64" pageNumber="65">
<heading id="5B26B122BA369F684E3E23EEFC3EFC3D" box="[621,967,995,1022]" centered="true" fontSize="11" level="2" pageId="64" pageNumber="65" reason="2">
<taxonomicName id="C7D17DCDBA369F684E3E23EEFCA4FC3D" authority="Wisshak, 2017" authorityName="Wisshak" authorityYear="2017" box="[621,861,995,1022]" class="Hexactinellida" genus="Pyritonema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lyssacinosida" pageId="64" pageNumber="65" phylum="Porifera" rank="species" species="belua" status="sp. nov.">
<emphasis id="32A5DA5CBA369F684E3E23EEFCA4FC3D" bold="true" box="[621,861,995,1022]" italics="true" pageId="64" pageNumber="65">Pyrodendrina belua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29966727BA369F684F3723E9FC3EFC3D" box="[868,967,996,1022]" pageId="64" pageNumber="65">isp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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</paragraph>
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<subSubSection id="48CB55C5BA369F684EA32427FC71FB4E" pageId="64" pageNumber="65" type="description">
<paragraph id="006E064EBA369F684EA32427FCBAFB86" blockId="64.[361,1226,995,1093]" box="[752,835,1066,1093]" pageId="64" pageNumber="65">
<figureCitation id="98EA1ACBBA369F684EA32427FCBAFB86" box="[752,835,1066,1093]" captionStart="Fig" captionStartId="65.[190,233,1825,1851]" captionTargetBox="[189,1401,999,1792]" captionTargetId="figure@65.[189,1401,999,1792]" captionTargetPageId="65" captionText="Fig. 25. Pyrodendrina belua isp. nov. A. SEM planar view of holotype, surrounded by a carpet of other microborings in an epoxy cast prepared from a bivalve shell sampled from the Middle Pennsylvanian Buckhorn Asphalt Lagerstätte in Oklahoma, USA. BC. Lateral views of holotype illustrating vertically oriented galleries, parts of which bearing perpendicular side braches. D. Detail of terminal widening and spiny ornamentation. E. Detail of slender prostrate galleries with terminal swellings and typical branching pattern; an anastomosis is developed at the lower right." figureDoi="http://doi.org/10.5281/zenodo.3839909" httpUri="https://zenodo.org/record/3839909/files/figure.png" pageId="64" pageNumber="65">Fig. 25</figureCitation>
</paragraph>
<paragraph id="006E064EBA369F684CEE247EFC71FB4E" blockId="64.[189,904,1139,1166]" box="[189,904,1139,1166]" pageId="64" pageNumber="65">
<treatmentCitationGroup id="20C12160BA369F684CEE247EFC71FB4E" box="[189,904,1139,1166]" pageId="64" pageNumber="65">
?
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<emphasis id="32A5DA5CBA369F684C99247EFE36FB4E" box="[202,463,1139,1165]" italics="true" pageId="64" pageNumber="65">Semidendrina pulchra</emphasis>
</taxonomicName>
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<bibRefCitation id="64407BBFBA369F684DBF247EFCFDFB4E" author="Wisshak" box="[492,772,1139,1165]" etAl="et al." firstAuthor="Wisshak" pageId="64" pageNumber="65" pagination="21 - 54" refId="ref55181" refString="Wisshak M., Seuss B. &amp; Nutzel A. 2008. Evolutionary implications of an exceptionally preserved Carboniferous microboring assemblage in the Buckhorn Asphalt lagerstatte (Oklahoma, USA). In: Wisshak M. &amp; Tapanila L. (eds) Current Developments in Bioerosion: 21 - 54. Springer, Heidelberg. https: // doi. org / 10.1007 / 978 - 3 - 540 - 77598 - 0 _ 2" type="book chapter" year="2008">
Wisshak
<emphasis id="32A5DA5CBA369F684E0B2479FD6BFB4E" box="[600,658,1139,1165]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
2008: 43
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, fig. 9CE.
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</paragraph>
</subSubSection>
<subSubSection id="48CB55C5BA369F684CEE24B1FDF4FA84" pageId="64" pageNumber="65" type="diagnosis">
<paragraph id="006E064EBA369F684CEE24B1FECFFB15" blockId="64.[189,1398,1212,1351]" box="[189,310,1212,1238]" pageId="64" pageNumber="65">
<heading id="5B26B122BA369F684CEE24B1FECFFB15" bold="true" box="[189,310,1212,1238]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="32A5DA5CBA369F684CEE24B1FECFFB15" bold="true" box="[189,310,1212,1238]" pageId="64" pageNumber="65">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="006E064EBA369F684CEE24EAFDF4FA84" blockId="64.[189,1398,1212,1351]" pageId="64" pageNumber="65">Prostrate galleries slender, with bi- or trifurcations and rare anastomosis. Additionally, sub-vertical tunnels reach straight into the substrate. All galleries end in widened terminations and are ornamented with short spiny protrusions.</paragraph>
</subSubSection>
<subSubSection id="48CB55C5BA369F684CEE257BFC24FA1D" pageId="64" pageNumber="65" type="etymology">
<paragraph id="006E064EBA369F684CEE257BFEBDFA53" blockId="64.[189,1398,1398,1503]" box="[189,324,1398,1424]" pageId="64" pageNumber="65">
<heading id="5B26B122BA369F684CEE257BFEBDFA53" bold="true" box="[189,324,1398,1424]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="32A5DA5CBA369F684CEE257BFEBDFA53" bold="true" box="[189,324,1398,1424]" pageId="64" pageNumber="65">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="006E064EBA369F684CEE25ACFC24FA1D" blockId="64.[189,1398,1398,1503]" pageId="64" pageNumber="65">From the Latin belua, monster (for the most part in the sea), referring to the former working name “the beast” this trace had earned because of its “fur” and “vertical teeth”.</paragraph>
</subSubSection>
<subSubSection id="48CB55C5BA369F684CEE2600FCF8F920" pageId="64" pageNumber="65" type="materials_examined">
<paragraph id="006E064EBA369F684CEE2600FD8AF9E4" blockId="64.[189,627,1549,1575]" box="[189,627,1549,1575]" pageId="64" pageNumber="65">
<heading id="5B26B122BA369F684CEE2600FD8AF9E4" bold="true" box="[189,627,1549,1575]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="32A5DA5CBA369F684CEE2600FD8AF9E4" bold="true" box="[189,627,1549,1575]" pageId="64" pageNumber="65">Type material, locality and horizon</emphasis>
</heading>
</paragraph>
<paragraph id="006E064EBA369F684CEE2631FCF8F920" blockId="64.[189,1398,1595,1763]" pageId="64" pageNumber="65">
<materialsCitation id="B0B90C13BA369F684CEE2631FD07F920" ID-GBIF-Occurrence="2625219311" collectionCode="BSPG" country="United States of America" location="Bayerische Staatssammlung fur Palaontologie und Geologie Munchen" municipality="The" pageId="64" pageNumber="65" specimenCode="BSPG 2007 XII 12" specimenCount="2" stateProvince="Oklahoma" typeStatus="holotype">
The
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(
<figureCitation id="98EA1ACBBA369F684D212631FE30F995" box="[370,457,1595,1622]" captionStart="Fig" captionStartId="65.[190,233,1825,1851]" captionTargetBox="[189,1401,999,1792]" captionTargetId="figure@65.[189,1401,999,1792]" captionTargetPageId="65" captionText="Fig. 25. Pyrodendrina belua isp. nov. A. SEM planar view of holotype, surrounded by a carpet of other microborings in an epoxy cast prepared from a bivalve shell sampled from the Middle Pennsylvanian Buckhorn Asphalt Lagerstätte in Oklahoma, USA. BC. Lateral views of holotype illustrating vertically oriented galleries, parts of which bearing perpendicular side braches. D. Detail of terminal widening and spiny ornamentation. E. Detail of slender prostrate galleries with terminal swellings and typical branching pattern; an anastomosis is developed at the lower right." figureDoi="http://doi.org/10.5281/zenodo.3839909" httpUri="https://zenodo.org/record/3839909/files/figure.png" pageId="64" pageNumber="65">Fig. 25</figureCitation>
AE) is cast in epoxy of a bivalve shell from the Buckhorn Asphalt, Boggy Formation, Deese group, Desmoinesian, middle Pennsylvanian, sampled at the Buckhorn Asphalt Quarry,
<collectingRegion id="C215C8ACBA369F684D4E268FFE63F95F" box="[285,410,1666,1692]" country="United States of America" name="Oklahoma" pageId="64" pageNumber="65">Oklahoma</collectingRegion>
,
<collectingCountry id="78C646DEBA369F684DF5268FFE1DF95F" box="[422,484,1666,1692]" name="United States of America" pageId="64" pageNumber="65">USA</collectingCountry>
.
<collectingMunicipality id="E00A9C34BA369F684DBD268FFDE5F95F" box="[494,540,1666,1692]" pageId="64" pageNumber="65">The</collectingMunicipality>
epoxy cast is stored in the
<location id="050E5095BA369F684F09268FFE2CF97C" LSID="urn:lsid:plazi:treatment:8878B758BA369F694E3E23EEFCB9FD05:050E5095BA369F684F09268FFE2CF97C" country="United States of America" municipality="The" name="Bayerische Staatssammlung fur Palaontologie und Geologie Munchen" pageId="64" pageNumber="65" stateProvince="Oklahoma">Bayerische Staatssammlung für Paläontologie und Geologie München</location>
,
<collectingCountry id="78C646DEBA369F684DB326A8FDB4F97C" box="[480,589,1701,1727]" name="Germany" pageId="64" pageNumber="65">Germany</collectingCountry>
(
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d).
<location id="050E5095BA369F684F3626ABFC77F903" LSID="urn:lsid:plazi:treatment:8878B758BA369F694E3E23EEFCB9FD05:050E5095BA369F684F3626ABFC77F903" box="[869,910,1702,1728]" country="United States of America" municipality="The" name="For" pageId="64" pageNumber="65" stateProvince="Oklahoma">For</location>
further details about the
<typeStatus id="DF6AB8ECBA369F6848E026ABFB1FF903" box="[1203,1254,1702,1728]" pageId="64" pageNumber="65">type</typeStatus>
locality and its ichnological record, see
<bibRefCitation id="64407BBFBA369F684DAD26C5FD07F920" author="Wisshak" box="[510,766,1736,1763]" etAl="et al." firstAuthor="Wisshak" pageId="64" pageNumber="65" pagination="21 - 54" refId="ref55181" refString="Wisshak M., Seuss B. &amp; Nutzel A. 2008. Evolutionary implications of an exceptionally preserved Carboniferous microboring assemblage in the Buckhorn Asphalt lagerstatte (Oklahoma, USA). In: Wisshak M. &amp; Tapanila L. (eds) Current Developments in Bioerosion: 21 - 54. Springer, Heidelberg. https: // doi. org / 10.1007 / 978 - 3 - 540 - 77598 - 0 _ 2" type="book chapter" year="2008">
<location id="050E5095BA369F684DAD26C5FD9BF921" LSID="urn:lsid:plazi:treatment:8878B758BA369F694E3E23EEFCB9FD05:050E5095BA369F684DAD26C5FD9BF921" box="[510,610,1736,1762]" country="United States of America" municipality="The" name="Wisshak" pageId="64" pageNumber="65" stateProvince="Oklahoma">Wisshak</location>
<emphasis id="32A5DA5CBA369F684E3A26C4FD5DF921" box="[617,676,1736,1762]" italics="true" pageId="64" pageNumber="65">et al.</emphasis>
(2008)
</bibRefCitation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="48CB55C5BA369F694CEE271CFACEFEAF" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65" type="description">
<paragraph id="006E064EBA369F684CEE271CFEB7F8E8" blockId="64.[189,334,1809,1835]" box="[189,334,1809,1835]" pageId="64" pageNumber="65">
<heading id="5B26B122BA369F684CEE271CFEB7F8E8" bold="true" box="[189,334,1809,1835]" fontSize="11" level="3" pageId="64" pageNumber="65" reason="3">
<emphasis id="32A5DA5CBA369F684CEE271CFEB7F8E8" bold="true" box="[189,334,1809,1835]" pageId="64" pageNumber="65">Description</emphasis>
</heading>
</paragraph>
<paragraph id="006E064EBA369F694CEE274DFACEFEAF" blockId="64.[189,1399,1856,2024]" lastBlockId="65.[189,1399,267,365]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
The basal dendritic network of straight galleries displays uni- or bilateral ramification at 45° to 55° and rare anastomoses (
<figureCitation id="98EA1ACBBA369F684DCB276EFDDEF8BD" box="[408,551,1891,1918]" captionStart="Fig" captionStartId="65.[190,233,1825,1851]" captionTargetBox="[189,1401,999,1792]" captionTargetId="figure@65.[189,1401,999,1792]" captionTargetPageId="65" captionText="Fig. 25. Pyrodendrina belua isp. nov. A. SEM planar view of holotype, surrounded by a carpet of other microborings in an epoxy cast prepared from a bivalve shell sampled from the Middle Pennsylvanian Buckhorn Asphalt Lagerstätte in Oklahoma, USA. BC. Lateral views of holotype illustrating vertically oriented galleries, parts of which bearing perpendicular side braches. D. Detail of terminal widening and spiny ornamentation. E. Detail of slender prostrate galleries with terminal swellings and typical branching pattern; an anastomosis is developed at the lower right." figureDoi="http://doi.org/10.5281/zenodo.3839909" httpUri="https://zenodo.org/record/3839909/files/figure.png" pageId="64" pageNumber="65">Fig. 25A, E</figureCitation>
). All galleries end in slightly widened terminations (
<figureCitation id="98EA1ACBBA369F6848C0276EFB10F8BD" box="[1171,1257,1891,1918]" captionStart="Fig" captionStartId="65.[190,233,1825,1851]" captionTargetBox="[189,1401,999,1792]" captionTargetId="figure@65.[189,1401,999,1792]" captionTargetPageId="65" captionText="Fig. 25. Pyrodendrina belua isp. nov. A. SEM planar view of holotype, surrounded by a carpet of other microborings in an epoxy cast prepared from a bivalve shell sampled from the Middle Pennsylvanian Buckhorn Asphalt Lagerstätte in Oklahoma, USA. BC. Lateral views of holotype illustrating vertically oriented galleries, parts of which bearing perpendicular side braches. D. Detail of terminal widening and spiny ornamentation. E. Detail of slender prostrate galleries with terminal swellings and typical branching pattern; an anastomosis is developed at the lower right." figureDoi="http://doi.org/10.5281/zenodo.3839909" httpUri="https://zenodo.org/record/3839909/files/figure.png" pageId="64" pageNumber="65">Fig. 25</figureCitation>
AE). From these basal galleries, additional tunnels of slightly larger diameter, with or without perpendicular side branches, penetrate the substrate sub-vertically and likewise end in widened terminations (
<figureCitation id="98EA1ACBBA369F68488927A7FAD6F807" box="[1242,1327,1961,1988]" captionStart="Fig" captionStartId="65.[190,233,1825,1851]" captionTargetBox="[189,1401,999,1792]" captionTargetId="figure@65.[189,1401,999,1792]" captionTargetPageId="65" captionText="Fig. 25. Pyrodendrina belua isp. nov. A. SEM planar view of holotype, surrounded by a carpet of other microborings in an epoxy cast prepared from a bivalve shell sampled from the Middle Pennsylvanian Buckhorn Asphalt Lagerstätte in Oklahoma, USA. BC. Lateral views of holotype illustrating vertically oriented galleries, parts of which bearing perpendicular side braches. D. Detail of terminal widening and spiny ornamentation. E. Detail of slender prostrate galleries with terminal swellings and typical branching pattern; an anastomosis is developed at the lower right." figureDoi="http://doi.org/10.5281/zenodo.3839909" httpUri="https://zenodo.org/record/3839909/files/figure.png" pageId="64" pageNumber="65">Fig. 25</figureCitation>
BD). The entire trace has a bulged surface texture and is ornamented with short spiny protrusions (
<figureCitation id="98EA1ACBBA369F6848AF27C0FA90F82B" box="[1276,1385,1997,2024]" captionStart="Fig" captionStartId="65.[190,233,1825,1851]" captionTargetBox="[189,1401,999,1792]" captionTargetId="figure@65.[189,1401,999,1792]" captionTargetPageId="65" captionText="Fig. 25. Pyrodendrina belua isp. nov. A. SEM planar view of holotype, surrounded by a carpet of other microborings in an epoxy cast prepared from a bivalve shell sampled from the Middle Pennsylvanian Buckhorn Asphalt Lagerstätte in Oklahoma, USA. BC. Lateral views of holotype illustrating vertically oriented galleries, parts of which bearing perpendicular side braches. D. Detail of terminal widening and spiny ornamentation. E. Detail of slender prostrate galleries with terminal swellings and typical branching pattern; an anastomosis is developed at the lower right." figureDoi="http://doi.org/10.5281/zenodo.3839909" httpUri="https://zenodo.org/record/3839909/files/figure.png" pageId="64" pageNumber="65">Fig. 25D</figureCitation>
). The
<typeStatus id="DF6AB8ECBA379F694CA02106FEA3FEE6" box="[243,346,267,293]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
measures
<quantity id="C729ABABBA379F694D842106FDCAFEE5" box="[471,563,267,294]" metricMagnitude="-1" metricUnit="m" metricValue="5.9399999999999995" pageId="65" pageNumber="66" unit="mm" value="594.0">594 µm</quantity>
in length and
<quantity id="C729ABABBA379F694E8D2106FCC3FEE5" box="[734,826,267,294]" metricMagnitude="-1" metricUnit="m" metricValue="2.96" pageId="65" pageNumber="66" unit="mm" value="296.0">296 µm</quantity>
in width, and has a maximum penetration depth of
<quantity id="C729ABABBA379F694C8F2123FEC3FE8A" box="[220,314,302,329]" metricMagnitude="-1" metricUnit="m" metricValue="2.35" pageId="65" pageNumber="66" unit="mm" value="235.0">235 µm</quantity>
. The horizontal galleries measure 20.9 ±
<quantity id="C729ABABBA379F694F4A2123FC95FE8A" box="[793,876,302,329]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="65" pageNumber="66" unit="mm" value="6.5">6.5 µm</quantity>
in thickness (n = 10) and the vertical tunnels are slightly thicker, 24.3 ±
<quantity id="C729ABABBA379F694DAA215FFDB4FEAF" box="[505,589,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="65" pageNumber="66" unit="mm" value="5.8">5.8 µm</quantity>
(n = 10). The spiny protrusions measure up to
<quantity id="C729ABABBA379F694827215FFB39FEAF" box="[1140,1216,338,365]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="65" pageNumber="66" unit="mm" value="15.0">15 µm</quantity>
in length.
</paragraph>
</subSubSection>
<subSubSection id="48CB55C5BA379F694CEE2181FCB9FD05" pageId="65" pageNumber="66" type="discussion">
<paragraph id="006E064EBA379F694CEE2181FED6FE65" blockId="65.[189,303,396,422]" box="[189,303,396,422]" pageId="65" pageNumber="66">
<heading id="5B26B122BA379F694CEE2181FED6FE65" bold="true" box="[189,303,396,422]" fontSize="11" level="3" pageId="65" pageNumber="66" reason="3">
<emphasis id="32A5DA5CBA379F694CEE2181FED6FE65" bold="true" box="[189,303,396,422]" pageId="65" pageNumber="66">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="006E064EBA379F694CEE21B7FDFFFDFD" blockId="65.[189,1399,442,574]" pageId="65" pageNumber="66">
Originally tentatively assigned to
<taxonomicName id="C7D17DCDBA379F694E1921B7FC90FE17" authorityName="Wisshak" authorityYear="2017" baseAuthorityName="Fischer" baseAuthorityYear="1875" box="[586,873,442,468]" class="Monothalamea" family="Astrorhizidae" genus="Nododendrina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Astrorhizida" pageId="65" pageNumber="66" phylum="Foraminifera" rank="species" species="europaea">
<emphasis id="32A5DA5CBA379F694E1921B7FC90FE17" box="[586,873,442,468]" italics="true" pageId="65" pageNumber="66">Nododendrina europaea</emphasis>
</taxonomicName>
(as
<emphasis id="32A5DA5CBA379F694FCA21B7FB66FE17" box="[921,1183,442,468]" italics="true" pageId="65" pageNumber="66">
Semidendrina
<taxonomicName id="C7D17DCDBA379F69481721B7FB66FE17" box="[1092,1183,442,468]" pageId="65" pageNumber="66" rank="species" species="pulchra">pulchra</taxonomicName>
</emphasis>
), this trace is now recognised as a different ichnospecies. It is clearly distinguished from
<taxonomicName id="C7D17DCDBA379F69484421D3FB52FE34" authorityName="Wisshak" authorityYear="2017" baseAuthorityName="Fischer" baseAuthorityYear="1875" box="[1047,1195,478,503]" class="Monothalamea" family="Astrorhizidae" genus="Nododendrina" higherTaxonomySource="GBIF" kingdom="Chromista" order="Astrorhizida" pageId="65" pageNumber="66" phylum="Foraminifera" rank="species" species="europaea">
<emphasis id="32A5DA5CBA379F69484421D3FB52FE34" box="[1047,1195,478,503]" italics="true" pageId="65" pageNumber="66">N. europaea</emphasis>
</taxonomicName>
by the lack of a vertically oriented main node, by the almost complete lack of anastomosis, and by the presence of many vertically oriented galleries.
</paragraph>
<paragraph id="006E064EBA379F694CEE2268FCB9FD05" blockId="65.[189,1399,613,710]" pageId="65" pageNumber="66">
Distinguished from other ichnospecies of
<emphasis id="32A5DA5CBA379F694EEC2268FC98FDBC" box="[703,865,613,639]" italics="true" pageId="65" pageNumber="66">Pyrodendrina</emphasis>
by its straight and slender galleries, which always widen near their termination. The central node is least distinct in this species, and the hairy ornament is less pronounced than in
<taxonomicName id="C7D17DCDBA379F694E3822A1FD2FFD05" box="[619,726,684,710]" class="Hexactinellida" genus="Pyritonema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lyssacinosida" pageId="65" pageNumber="66" phylum="Porifera" rank="species" species="villosa" status="sp. nov.">
<emphasis id="32A5DA5CBA379F694E3822A1FD2FFD05" box="[619,726,684,710]" italics="true" pageId="65" pageNumber="66">P. villosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="29966727BA379F694E8E22A6FCB9FD05" box="[733,832,683,710]" pageId="65" pageNumber="66">isp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>