treatments-xml/data/6D/99/E5/6D99E53EA8305EC8AF22BF81F61B4241.xml
2024-06-21 12:39:39 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1033.60686" ID-GBIF-Dataset="920fdd6f-b0e8-4059-8ff8-dd401a03625f" ID-PMC="PMC8084852" ID-Pensoft-Pub="1313-2970-1033-81" ID-Pensoft-UUID="2E6467D3D0025F778A77798A4072BCB5" ID-PubMed="33958921" ID-ZooBank="BEFB015F4A354987AD10DFEB7D4F91DD" ModsDocID="1313-2970-1033-81" checkinTime="1619185013811" checkinUser="pensoft" docAuthor="Pinho, Luiz Carlos &amp; Andersen, Trond" docDate="2021" docId="6D99E53EA8305EC8AF22BF81F61B4241" docLanguage="en" docName="ZooKeys 1033: 81-125" docOrigin="ZooKeys 1033" docPubDate="2021-04-22" docSource="http://dx.doi.org/10.3897/zookeys.1033.60686" docTitle="Nilothauma leccii Pinho &amp; Andersen 2021, sp. nov." docType="treatment" docUuid="CBFC7D88-17AB-4BF8-9CBA-2995FDAB798B" docUuidSource="ZooBank" docVersion="4" id="2E6467D3D0025F778A77798A4072BCB5" lastPageNumber="81" masterDocId="2E6467D3D0025F778A77798A4072BCB5" masterDocTitle="Neotropical Nilothauma Kieffer, 1921 (Diptera, Chironomidae): Key, eleven new species, re-descriptions, new combination and new records" masterLastPageNumber="125" masterPageNumber="81" pageNumber="81" updateTime="1668152459540" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Neotropical Nilothauma Kieffer, 1921 (Diptera, Chironomidae): Key, eleven new species, re-descriptions, new combination and new records</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pinho, Luiz Carlos</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9153-9997</mods:nameIdentifier>
<mods:affiliation>Laboratory of Systematics of Diptera, Department of Ecology and Zoology, Federal University of Santa Catarina, Campus Trindade, CEP 88040 - 900, Florianopolis, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">luizcarlospinho@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Andersen, Trond</mods:namePart>
<mods:affiliation>Department of Natural History, University Museum of Bergen, University of Bergen, P. O. Box 7800, NO- 5020, Bergen, Norway</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-22</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1033</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>81</mods:start>
<mods:end>125</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1033.60686</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1033.60686</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1033-81</mods:identifier>
<mods:identifier type="ZooBank">BEFB015F4A354987AD10DFEB7D4F91DD</mods:identifier>
<mods:identifier type="Pensoft-UUID">2E6467D3D0025F778A77798A4072BCB5</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181534243" LSID="urn:lsid:zoobank.org:act:CBFC7D88-17AB-4BF8-9CBA-2995FDAB798B" httpUri="http://treatment.plazi.org/id/6D99E53EA8305EC8AF22BF81F61B4241" lastPageNumber="81" pageId="0" pageNumber="81">
<subSubSection pageId="0" pageNumber="81" type="nomenclature">
<paragraph pageId="0" pageNumber="81">
<taxonomicName LSID="http://zoobank.org/CBFC7D88-17AB-4BF8-9CBA-2995FDAB798B" authority="Pinho &amp; Andersen, 2021" authorityName="Pinho &amp; Andersen" authorityYear="2021" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nilothauma leccii" order="Diptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="leccii" status="sp. nov.">Nilothauma leccii</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="81">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Nilothauma leccii sp. nov. adult male A hypopygium, dorsal view B hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="10.3897/zookeys.1033.60686.figure7" httpUri="https://binary.pensoft.net/fig/534700" pageId="0" pageNumber="81">Figures 7A, B</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution maps of Neotropical Nilothauma species A N. hamadae sp. nov., N. jupau sp. nov., N. terena sp. nov., N. yekwana sp. nov., N. complicatum Mendes &amp; Andersen, 2009 B N. leccii sp. nov., N. marianoi sp. nov., N. mateusi sp. nov., N. txukuyana sp. nov., N. maya sp. nov. C N. karitiana sp. nov., N. duena Roback, 1960, N. calori Mendes &amp; Andersen, 2009 D N. werekena sp. nov., N. zitoi Mendes &amp; Andersen, 2009, N. strebulosum (Adam &amp; Saether, 2000)." figureDoi="10.3897/zookeys.1033.60686.figure16" httpUri="https://binary.pensoft.net/fig/534709" pageId="0" pageNumber="81">, 16B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="type material">
<paragraph pageId="0" pageNumber="81">Type material.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Holotype</emphasis>
male, slide-mounted: Brazil,
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo,
<normalizedToken originalValue="São">Sao</normalizedToken>
<normalizedToken originalValue="Sebastião">Sebastiao</normalizedToken>
, Rio das Pedras,
<geoCoordinate degrees="23" direction="south" minutes="44" orientation="latitude" precision="15" seconds="27" value="-23.740833">23°44'27&quot;S</geoCoordinate>
,
<geoCoordinate degrees="45" direction="west" minutes="37" orientation="longitude" precision="15" seconds="12" value="-45.62">45°37'12&quot;W</geoCoordinate>
, 28.x.2005, light trap, A.R. Calor et al. leg. (UFSC).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="etymology">
<paragraph pageId="0" pageNumber="81">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="81">The specific epithet is a noun in the genitive case honouring Lucas Silveira Lecci, for his friendship and prolific fieldwork.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Diagnostic characters.</paragraph>
<paragraph pageId="0" pageNumber="81">The male can be separated from its congeners by its large size combined with unmarked wing; spatulate anal point; superior volsella leaf-shapped; inferior volsella with strong, split setae and digitiform and strongly setose gonostylus.</paragraph>
<caption doi="10.3897/zookeys.1033.60686.figure7" httpUri="https://binary.pensoft.net/fig/534700" pageId="0" pageNumber="81" start="Figure 7" startId="F7">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Figure 7.</emphasis>
<taxonomicName authorityName="Pinho &amp; Andersen" authorityYear="2021" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nilothauma leccii" order="Diptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="leccii">
<emphasis italics="true" pageId="0" pageNumber="81">Nilothauma leccii</emphasis>
</taxonomicName>
sp. nov. adult male
<emphasis bold="true" pageId="0" pageNumber="81">A</emphasis>
hypopygium, dorsal view
<emphasis bold="true" pageId="0" pageNumber="81">B</emphasis>
hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="description">
<paragraph pageId="0" pageNumber="81">Description.</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Male imago (n = 1).</emphasis>
Total length 4.35 mm. Wing length 1.96 mm. Total length/wing length 2.22. Wing length/length of profemur 2.45.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Colouration</emphasis>
.
</emphasis>
Thorax and legs brown, abdomen light brown. Wing membrane without dark markings.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Antenna</emphasis>
.
</emphasis>
AR = 0.16. Thirteenth flagellomere 124
<normalizedToken originalValue="µm">µm</normalizedToken>
long.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Head</emphasis>
.
</emphasis>
Temporal setae 6 in single row. Clypeus with 15 setae. Tentorium 113
<normalizedToken originalValue="µm">µm</normalizedToken>
long, maximum width 25
<normalizedToken originalValue="µm">µm</normalizedToken>
. Stipes not measurable. Palp segment lengths (in
<normalizedToken originalValue="µm">µm</normalizedToken>
): 37, 33, 74, 107, 138. Third palpomere with 2 sensilla clavata subapically, longest about 25
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Fifth palpomere/third palpomere 1.86.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Thorax</emphasis>
.
</emphasis>
Antepronotum with 4 setae. Dorsocentrals 17 partly biserial posterior, acrostichals 14, prealars 6. Scutellum with 13 setae.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Wing</emphasis>
.
</emphasis>
VR = 1.53. Brachiolum with 2 setae, R with 15 setae, R1 with 24 setae, R4+5 with 31 setae, remaining veins bare.
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
<emphasis italics="true" pageId="0" pageNumber="81">Legs</emphasis>
.
</emphasis>
Spur of fore tibia 65
<normalizedToken originalValue="µm">µm</normalizedToken>
long including 41
<normalizedToken originalValue="µm">µm</normalizedToken>
long scale. Mid-tibia with 1 spur, 47
<normalizedToken originalValue="µm">µm</normalizedToken>
long; hind tibia with 2 spurs, 43 and 65
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Combs of mid-tibia 29
<normalizedToken originalValue="µm">µm</normalizedToken>
long, of hind tibia 47
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Width at apex of fore tibia 69
<normalizedToken originalValue="µm">µm</normalizedToken>
, of mid-tibia 73
<normalizedToken originalValue="µm">µm</normalizedToken>
, of hind tibia 89
<normalizedToken originalValue="µm">µm</normalizedToken>
. Lengths and proportions of legs as in Table
<tableCitation captionStart="Table 6" captionStartId="T6" captionText="Table 6. Lengths (in μm) and proportions of leg segments in Nilothauma leccii sp. nov., adult male (n = 1)." httpUri="http://table.plazi.org/id/B11614E273B37F373B2C74854E601A12" pageId="0" pageNumber="81" tableUuid="B11614E273B37F373B2C74854E601A12">6</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="B11614E273B37F373B2C74854E601A12" httpUri="http://table.plazi.org/id/B11614E273B37F373B2C74854E601A12" pageId="0" pageNumber="81" start="Table 6" startId="T6">
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">Table 6.</emphasis>
Lengths (in
<normalizedToken originalValue="μm">μm</normalizedToken>
) and proportions of leg segments in
<taxonomicName authorityName="Pinho &amp; Andersen" authorityYear="2021" class="Insecta" family="Chironomidae" genus="Nilothauma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nilothauma leccii" order="Diptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="leccii">
<emphasis italics="true" pageId="0" pageNumber="81">Nilothauma leccii</emphasis>
</taxonomicName>
sp. nov., adult male (n = 1).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="81">
<table pageId="0" pageNumber="81">
<tr pageId="0" pageNumber="81">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">Fe</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">Ti</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">ta1</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">ta2</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">ta3</th>
<th colspan="1" pageId="0" pageNumber="81" rowspan="1">ta4</th>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">801</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">670</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">972</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">490</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">384</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">286</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">874</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">605</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">400</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">180</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">139</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">90</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">964</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">989</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">596</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">302</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">261</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">155</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">ta5</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">LR</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">BV</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">SV</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="81">BR</emphasis>
</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p1</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">139</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">1.45</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">1.88</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">1.51</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">1.7</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p2</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">65</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0.66</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.97</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.69</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4.3</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
<tr pageId="0" pageNumber="81">
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">p3</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">98</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">0.60</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.09</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">3.27</td>
<td colspan="1" pageId="0" pageNumber="81" rowspan="1">4.8</td>
<td colspan="1" isEmpty="true" pageId="0" pageNumber="81" rowspan="1">-</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" italics="true" pageId="0" pageNumber="81">Hypopygium</emphasis>
(Fig.
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Nilothauma leccii sp. nov. adult male A hypopygium, dorsal view B hypopygium with anal point and tergite IX removed, dorsal aspect to the left and ventral aspect to the right." figureDoi="10.3897/zookeys.1033.60686.figure7" httpUri="https://binary.pensoft.net/fig/534700" pageId="0" pageNumber="81">7A, B</figureCitation>
). Tergite IX without dorsal lobes, posterior margin subtriangular with 7 setae to each side of the anal point. Anal point spatulate, 59
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 10
<normalizedToken originalValue="µm">µm</normalizedToken>
wide basally, 17
<normalizedToken originalValue="µm">µm</normalizedToken>
wide medially. Tergite bands not continuous. Laterosternite IX with 4 setae. Phallapodeme 61
<normalizedToken originalValue="µm">µm</normalizedToken>
long; transverse sternapodeme 35
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Gonocoxite 171
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Inferior volsella weakly curved, 113
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 15
<normalizedToken originalValue="µm">µm</normalizedToken>
wide subapically, with microtrichia and 7 strong, apically split setae. Superior volsella leaf-shaped, 69
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 10
<normalizedToken originalValue="µm">µm</normalizedToken>
wide at base, 23
<normalizedToken originalValue="µm">µm</normalizedToken>
wide medially, covered with microtrichia and with few weak setae ventrally and along inner margin. Median volsella narrow, 48
<normalizedToken originalValue="µm">µm</normalizedToken>
long, covered with microtrichia and with 4 setae on tubercles at apex, setae about 23
<normalizedToken originalValue="µm">µm</normalizedToken>
long. Gonostylus digitiform, strongly setose, 163
<normalizedToken originalValue="µm">µm</normalizedToken>
long, 37
<normalizedToken originalValue="µm">µm</normalizedToken>
wide medially. HR = 1.05. HV = 2.67.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="female adult and immatures">
<paragraph pageId="0" pageNumber="81">Female adult and immatures.</paragraph>
<paragraph pageId="0" pageNumber="81">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="81" type="distribution">
<paragraph pageId="0" pageNumber="81">Distribution</paragraph>
<paragraph pageId="0" pageNumber="81">
<emphasis bold="true" pageId="0" pageNumber="81">
(Fig.
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Distribution maps of Neotropical Nilothauma species A N. hamadae sp. nov., N. jupau sp. nov., N. terena sp. nov., N. yekwana sp. nov., N. complicatum Mendes &amp; Andersen, 2009 B N. leccii sp. nov., N. marianoi sp. nov., N. mateusi sp. nov., N. txukuyana sp. nov., N. maya sp. nov. C N. karitiana sp. nov., N. duena Roback, 1960, N. calori Mendes &amp; Andersen, 2009 D N. werekena sp. nov., N. zitoi Mendes &amp; Andersen, 2009, N. strebulosum (Adam &amp; Saether, 2000)." figureDoi="10.3897/zookeys.1033.60686.figure16" httpUri="https://binary.pensoft.net/fig/534709" pageId="0" pageNumber="81">16B</figureCitation>
).
</emphasis>
Only known from
<normalizedToken originalValue="São">Sao</normalizedToken>
Paulo State in Brazil.
</paragraph>
</subSubSection>
</treatment>
</document>