409 lines
42 KiB
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409 lines
42 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/jhr.90.80150" ID-GBIF-Dataset="e637a41b-b775-4a5e-ab68-a0ff6c3111db" ID-Pensoft-Pub="1314-2607-90-101" ID-Pensoft-UUID="CB21DA90B0FB5683B712B7717D201AD6" ID-Zenodo-Dep="6508930" ID-ZooBank="13EC4891F1244E77BC22F6978445DAE7" ModsDocID="1314-2607-90-101" checkinTime="1651307098940" checkinUser="pensoft" docAuthor="Zhang, Bing-Lan, Yan, Cheng-Jin, van Achterberg, Cornelis, Peng, Yan-Qiong & Chen, Hua-Yan" docDate="2022" docId="47B53C532E3C5D28932C016A34882F81" docLanguage="en" docName="JourHymenoptRes 90: 101-128" docOrigin="Journal of Hymenoptera Research 90" docPubDate="2022-04-29" docSource="http://dx.doi.org/10.3897/jhr.90.80150" docTitle="Lycogaster umbonata Chen & van Achterberg 2022, sp. nov." docType="treatment" docUuid="1CA730CA-7C2D-4713-B62D-64BEF9397958" docUuidSource="ZooBank" docVersion="4" id="CB21DA90B0FB5683B712B7717D201AD6" lastPageNumber="101" masterDocId="CB21DA90B0FB5683B712B7717D201AD6" masterDocTitle="Integrated taxonomy unveils new species of Trigonalyidae (Insecta, Hymenoptera) from Yunnan, China" masterLastPageNumber="128" masterPageNumber="101" pageNumber="101" updateTime="1678757742368" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Integrated taxonomy unveils new species of Trigonalyidae (Insecta, Hymenoptera) from Yunnan, China</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zhang, Bing-Lan</mods:namePart>
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<mods:affiliation>School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yan, Cheng-Jin</mods:namePart>
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<mods:affiliation>Wenzhou Vocational School of Science and Technology, Wenzhou 325300, China & Wencheng Modern Agriculture and Health Industry Research Institute, Wenzhou 325300, China</mods:affiliation>
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<mods:nameIdentifier type="email">ycj0685@163.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>van Achterberg, Cornelis</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6495-4853</mods:nameIdentifier>
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<mods:affiliation>State Key Laboratory of Rice Biology and Ministry of Agriculture, Hangzhou 310058, China & Key Lab of Agricultural Entomology, Institute of Insect Sciences, Zhejiang University, Hangzhou 310058, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Peng, Yan-Qiong</mods:namePart>
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<mods:affiliation>CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Chen, Hua-Yan</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-0382-1635</mods:nameIdentifier>
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<mods:affiliation>Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China</mods:affiliation>
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<mods:nameIdentifier type="email">huayanc@scbg.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Journal of Hymenoptera Research</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-04-29</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>90</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>101</mods:start>
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<mods:end>128</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/jhr.90.80150</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.90.80150</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2607-90-101</mods:identifier>
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<treatment ID-GBIF-Taxon="194907407" LSID="urn:lsid:zoobank.org:act:1CA730CA-7C2D-4713-B62D-64BEF9397958" httpUri="http://treatment.plazi.org/id/47B53C532E3C5D28932C016A34882F81" lastPageNumber="101" pageId="0" pageNumber="101">
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<subSubSection pageId="0" pageNumber="101" type="nomenclature">
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<paragraph pageId="0" pageNumber="101">
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<taxonomicName LSID="http://zoobank.org/1CA730CA-7C2D-4713-B62D-64BEF9397958" authority="Chen & van Achterberg" authorityName="Chen & van Achterberg" authorityYear="2022" class="Insecta" family="Trigonalyidae" genus="Lycogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycogaster umbonata" order="Hymenoptera" pageId="0" pageNumber="101" phylum="Arthropoda" rank="species" species="umbonata" status="sp. nov.">Lycogaster umbonata Chen & van Achterberg</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="101">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="description">
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<paragraph pageId="0" pageNumber="101">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">Figs 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="diagnosis">
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<paragraph pageId="0" pageNumber="101">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="101">
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Antenna elongate and hardly widened medially (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4A</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6A</figureCitation>
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); second sternite of female with a strong lobe-shaped protuberance medio-apically (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5E</figureCitation>
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); mandibles with ivory patches (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4B</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6B</figureCitation>
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); second tergite of female strongly shiny and with elliptical depressions antero-laterally (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5D</figureCitation>
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); wing membrane subhyaline and without darkened (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4A</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5A</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6A</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="101">
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<emphasis bold="true" pageId="0" pageNumber="101">Figure 6.</emphasis>
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<taxonomicName authorityName="Chen & van Achterberg" authorityYear="2022" class="Insecta" family="Trigonalyidae" genus="Lycogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycogaster umbonata" order="Hymenoptera" pageId="0" pageNumber="101" phylum="Arthropoda" rank="species" species="umbonata">
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<emphasis italics="true" pageId="0" pageNumber="101">Lycogaster umbonata</emphasis>
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</taxonomicName>
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Chen & van Achterberg, sp. nov., paratype, male (En-419022)
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<emphasis bold="true" pageId="0" pageNumber="101">A</emphasis>
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habitus, lateral aspect
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<emphasis bold="true" pageId="0" pageNumber="101">B</emphasis>
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head, anterior aspect
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<emphasis bold="true" pageId="0" pageNumber="101">C</emphasis>
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metasoma, dorsal aspect
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<emphasis bold="true" pageId="0" pageNumber="101">D</emphasis>
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metasoma, lateral aspect
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<emphasis bold="true" pageId="0" pageNumber="101">E</emphasis>
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metasoma, ventral aspect.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="101" type="description">
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<paragraph pageId="0" pageNumber="101">Description.</paragraph>
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<paragraph pageId="0" pageNumber="101">
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<emphasis bold="true" pageId="0" pageNumber="101">Female.</emphasis>
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Holotype, length of body 5.1 mm (of fore wing 4.1 mm).
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</paragraph>
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<paragraph pageId="0" pageNumber="101">
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<emphasis bold="true" pageId="0" pageNumber="101">
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<emphasis italics="true" pageId="0" pageNumber="101">Head</emphasis>
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.
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</emphasis>
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Antenna with 21 segments, elongate and hardly widened medially (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4A</figureCitation>
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); frons spaced punctate and shiny, with distinct median depression (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4B</figureCitation>
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); vertex and temple largely smooth and shiny with sparse and fine punctures (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4B, C</figureCitation>
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); head subparallel-sided behind eyes, distinctly narrowed posteriorly and 1.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as mesoscutum (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4C</figureCitation>
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); dorsal length of eye approximately as long as temple (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4C</figureCitation>
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); occipital carina narrow lamelliform and with weak crenulae medio-dorsally (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4C</figureCitation>
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); supra-antennal elevations hardly developed as a thin rim and smooth, 0.3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as scapus (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4C</figureCitation>
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); clypeus emarginate and comparatively thin medio-ventrally (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4B</figureCitation>
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).
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</paragraph>
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<paragraph pageId="0" pageNumber="101">
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<emphasis bold="true" pageId="0" pageNumber="101">
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<emphasis italics="true" pageId="0" pageNumber="101">Mesosoma</emphasis>
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.
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</emphasis>
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Length of mesosoma 1.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as its height (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5C</figureCitation>
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); mesopleuron below transverse mesopleural groove rugose to punctate anteriorly, largely smooth and shiny posteriorly, above groove similar but rugose anteriorly (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5C</figureCitation>
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); notauli narrow anteriorly, widened posteriorly and coarsely crenulate; middle lobe of mesoscutum sparsely and rather finely punctate anteriorly, coarsely punctate at posterior end, lateral lobes sparsely and rather finely punctate (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5B</figureCitation>
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); scutellar sulcus wide, both medially and laterally and coarsely crenulate (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5B</figureCitation>
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); scutellum largely smooth with sparsely and finely punctate, distinctly shiny, rather flat and anteriorly near level of mesoscutum; metascutellum shiny and smooth medio-posteriorly (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5B</figureCitation>
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); propodeum largely obliquely rugulose to rugose with smooth interspaces anterio-laterally (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5B</figureCitation>
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); posterior propodeal carina thick lamelliform and distinctly arched, foramen approximately 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
|
||
as wide as high medially (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">
|
||
<emphasis bold="true" pageId="0" pageNumber="101">
|
||
<emphasis italics="true" pageId="0" pageNumber="101">Wings</emphasis>
|
||
.
|
||
</emphasis>
|
||
Fore wing: length of vein 1-M 1.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as vein 1-SR; second submarginal cell 1.4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as third submarginal cell (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">
|
||
<emphasis bold="true" pageId="0" pageNumber="101">
|
||
<emphasis italics="true" pageId="0" pageNumber="101">Metasoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
First tergite 0.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as its apical width, smooth, with rather distinct elliptical depression medially and basal half with several distinct coarse transverse rugae medially (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5D</figureCitation>
|
||
); second tergite with elliptical depressions antero-laterally; second-fifth tergites largely smooth with sparse fine punctures (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5D</figureCitation>
|
||
); sternites sparsely and finely punctate (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5F</figureCitation>
|
||
); second sternite strongly convex in lateral view, with a strong lobe-shaped protuberance medio-apically (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5E</figureCitation>
|
||
); third sternite about 0.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
as long as second sternite, with distinct ledge (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5F</figureCitation>
|
||
); hypopygium truncate apically (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">
|
||
<emphasis bold="true" pageId="0" pageNumber="101">
|
||
<emphasis italics="true" pageId="0" pageNumber="101">Colour</emphasis>
|
||
.
|
||
</emphasis>
|
||
Black; mandibular teeth and palpi dark brown; anterior-basal part of mandible ivory (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4B</figureCitation>
|
||
); distal of fumer, tibia, tarsus and claw yellow to brownish (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4A</figureCitation>
|
||
); pterostigma dark brown; wing membrane subhyaline and without darkened (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">
|
||
<emphasis bold="true" pageId="0" pageNumber="101">Male.</emphasis>
|
||
Paratype, length of body 5.9 mm, of fore wing 4.5 mm; antenna with 21 segments and without tyloids (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6A</figureCitation>
|
||
); vertex densely and coarsely punctate; sculpture of mesosoma coarser than that in female (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6B</figureCitation>
|
||
); second tergite with a round depression antero-medially; second-fifth tergites and sternites densely and rather fine punctate, except sparsely and coarsely punctate in big round area of third tergite medially (Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6C-E</figureCitation>
|
||
); genitalia extruded (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6D</figureCitation>
|
||
); ivory part of mandible larger than that of female; clypeus with ivory patches (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6B</figureCitation>
|
||
); large part of hind tibia, hind tarsus and hind claw brown (Fig.
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6A</figureCitation>
|
||
), not like that in female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101" type="etymology">
|
||
<paragraph pageId="0" pageNumber="101">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">
|
||
Named after the strong lobe-shaped protuberance of female second sternite: from "
|
||
<emphasis italics="true" pageId="0" pageNumber="101">umbo</emphasis>
|
||
" (Latin for "rounded protuberance").
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="101">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">
|
||
<materialsCitation accessionNumber="OM057968" collectingDate="2020-09-15" collectingDateMax="2020-09-30" collectingDateMin="2020-09-15" collectorName="Lang Yi, GenBank" country="China" elevation="2390" latitude="25.299238" location="Longyang Dist." longLatPrecision="1" longitude="98.76982" municipality="Baoshan City" specimenCount="1" stateProvince="Yunnan" typeStatus="Holotype">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">
|
||
<typeStatus>Holotype</typeStatus>
|
||
</emphasis>
|
||
, female,
|
||
<collectingCountry name="China">China</collectingCountry>
|
||
:
|
||
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
|
||
,
|
||
<collectingMunicipality>Baoshan City</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:47B53C532E3C5D28932C016A34882F81:7576F16052638F4637C2DCE9950C3CF5" country="China" latitude="25.299238" longLatPrecision="1" longitude="98.76982" municipality="Baoshan City" name="Longyang Dist." stateProvince="Yunnan">Longyang Dist.</location>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.3899999999999997" unit="m" value="2390.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.3899999999999997" unit="m" value="2390.0">2390 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate degrees="25" direction="north" minutes="17" orientation="latitude" precision="1" seconds="57.26" value="25.299238">25°17'57.26"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="98" direction="east" minutes="46" orientation="longitude" precision="1" seconds="11.35" value="98.76982">98°46'11.35"E</geoCoordinate>
|
||
,
|
||
<collectingDate value="2020-09-15" valueMax="2020-09-30" valueMin="2020-09-15">15-30.IX.2020</collectingDate>
|
||
,
|
||
<collectorName>Lang Yi</collectorName>
|
||
leg., MT, SYSBM En-419021 (deposited in SYSBM);
|
||
<collectorName>GenBank</collectorName>
|
||
:
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OM057968">OM057968</accessionNumber>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation accessionNumber="OM057967" collectingDate="2020-07-30" collectingDateMax="2020-08-15" collectingDateMin="2020-07-30" collectorName="Lang Yi, GenBank" country="China" elevation="2390" latitude="25.299238" location="Longyang Dist." longLatPrecision="1" longitude="98.76982" municipality="Baoshan City" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="Paratype">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="101">
|
||
<typeStatus>Paratype</typeStatus>
|
||
</emphasis>
|
||
,
|
||
<specimenCount type="male">1 male</specimenCount>
|
||
,
|
||
<collectingCountry name="China">China</collectingCountry>
|
||
:
|
||
<collectingRegion country="China" name="Yunnan">Yunnan</collectingRegion>
|
||
,
|
||
<collectingMunicipality>Baoshan City</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:47B53C532E3C5D28932C016A34882F81:F69FEF649C05CF5ED9E2A5099A33C869" country="China" latitude="25.299238" longLatPrecision="1" longitude="98.76982" municipality="Baoshan City" name="Longyang Dist." stateProvince="Yunnan">Longyang Dist.</location>
|
||
,
|
||
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.3899999999999997" unit="m" value="2390.0">
|
||
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.3899999999999997" unit="m" value="2390.0">2390 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate degrees="25" direction="north" minutes="17" orientation="latitude" precision="1" seconds="57.26" value="25.299238">25°17'57.26"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="98" direction="east" minutes="46" orientation="longitude" precision="1" seconds="11.35" value="98.76982">98°46'11.35"E</geoCoordinate>
|
||
,
|
||
<collectingDate value="2020-07-30" valueMax="2020-08-15" valueMin="2020-07-30">30.VII-15.VIII.2020</collectingDate>
|
||
,
|
||
<collectorName>Lang Yi</collectorName>
|
||
leg.,MT, SYSBM En-419022 (SYSBM);
|
||
<collectorName>GenBank</collectorName>
|
||
:
|
||
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/OM057967">OM057967</accessionNumber>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="101" type="distribution">
|
||
<paragraph pageId="0" pageNumber="101">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">China (Yunnan). Collected at 2390 m.</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">
|
||
The key to Chinese species of
|
||
<taxonomicName authorityName="Shuckard" authorityYear="1841" class="Insecta" family="Trigonalyidae" genus="Lycogaster" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Lycogaster" order="Hymenoptera" pageId="0" pageNumber="101" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="101">Lycogaster</emphasis>
|
||
</taxonomicName>
|
||
published by
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.385.6560" author="Chen, H-Y" journalOrPublisher="ZooKeys" pageId="0" pageNumber="101" pagination="1 - 207" refId="B6" refString="Chen, H-Y, van Achterberg, C, He, J-H, Xu, Z-F, 2014. A revision of the Chinese Trigonalyidae (Hymenoptera, Trigonalyoidea). ZooKeys 385: 1 - 207, DOI: https://doi.org/10.3897/zookeys.385.6560" title="A revision of the Chinese Trigonalyidae (Hymenoptera, Trigonalyoidea)." url="https://doi.org/10.3897/zookeys.385.6560" volume="385" year="2014">Chen et al. (2014)</bibRefCitation>
|
||
could be updated to accommodate
|
||
<taxonomicName lsidName="L. umbonata" pageId="0" pageNumber="101" rank="species" species="umbonata">
|
||
<emphasis italics="true" pageId="0" pageNumber="101">L. umbonata</emphasis>
|
||
</taxonomicName>
|
||
by replacing couplet 2 as follows:
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="101">
|
||
<table inLine="true" pageId="0" pageNumber="101">
|
||
<tr pageId="0" pageNumber="101">
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">2</td>
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
|
||
Female antenna elongate and hardly widened medially (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4A</figureCitation>
|
||
); second tergite smooth and strongly shiny (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5D</figureCitation>
|
||
); medial third of fore wing subhyaline (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A habitus, lateral aspect B head, anterior aspect C head, dorsal aspect." figureDoi="10.3897/jhr.90.80150.figure4" httpUri="https://binary.pensoft.net/fig/677741" pageId="0" pageNumber="101">4A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5A</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Lycogaster umbonata Chen & van Achterberg, sp. nov., paratype, male (En- 419022) A habitus, lateral aspect B head, anterior aspect C metasoma, dorsal aspect D metasoma, lateral aspect E metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure6" httpUri="https://binary.pensoft.net/fig/677743" pageId="0" pageNumber="101">6A</figureCitation>
|
||
) or slightly darkened
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="101">2</emphasis>
|
||
'
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="101">
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">Female antenna less elongate and distinctly widened medially; second tergite punctate and moderately shiny; medial third of fore wing largely dark brown</td>
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="101">3</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="101">
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">2'</td>
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">Medio-apical protuberance of second sternite with pair of small submedial acute teeth close to each other; posterior end of middle lobe of mesoscutum rugose</td>
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="101">
|
||
<taxonomicName lsidName="L. angustula" pageId="0" pageNumber="101" rank="species" species="angustula">
|
||
<emphasis italics="true" pageId="0" pageNumber="101">L. angustula</emphasis>
|
||
</taxonomicName>
|
||
Chen, van Achterberg, He & Xu
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="101">
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">-</td>
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
|
||
Medio-apical protuberance of second sternite obtuse, lamelliform and convex medio-apically, without small submedial acute teeth (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5E</figureCitation>
|
||
); posterior end of middle lobe of mesoscutum largely smooth with sparse fine punctures (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Lycogaster umbonata Chen & van Achterberg, sp. nov., holotype, female (En- 419021) A wings B mesosoma, dorsal aspect C mesosoma, lateral aspect D metasoma, dorsal aspect E metasoma, lateral aspect F metasoma, ventral aspect." figureDoi="10.3897/jhr.90.80150.figure5" httpUri="https://binary.pensoft.net/fig/677742" pageId="0" pageNumber="101">5B</figureCitation>
|
||
)
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="101" rowspan="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="101">
|
||
<taxonomicName lsidName="L. umbonata" pageId="0" pageNumber="101" rank="species" species="umbonata">
|
||
<emphasis italics="true" pageId="0" pageNumber="101">L. umbonata</emphasis>
|
||
</taxonomicName>
|
||
Chen & van Achterberg, sp. nov.
|
||
</emphasis>
|
||
</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |