treatments-xml/data/47/66/B6/4766B618E87CFFC375C5FBDFF79EFCB7.xml

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<mods:title id="CF0F56278F287E31395258E4C96DAF8B">A new species of Monstrilla (Crustacea: Copepoda: Monstrilloida: Monstrillidae) collected by the R / V Albatross (1908) from the Sulu Sea, Philippines</mods:title>
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<taxonomicName id="08CF7C8DE87CFFCB75C5FBDFF4EDFBDC" authority="Suárez-Morales, 2022" authorityName="Suárez-Morales" authorityYear="2022" box="[488,724,1052,1078]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="1" pageNumber="3042" phylum="Arthropoda" rank="species" species="albatrossi" status="sp. nov.">Monstrilla albatrossi</taxonomicName>
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<taxonomicNameLabel id="E6886667E87CFFCB76F1FBDFF505FBDC" box="[732,828,1052,1078]" pageId="1" pageNumber="3042" rank="species">sp. nov.</taxonomicNameLabel>
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http:// 
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urn:lsid:zoobank.org:pub: 
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<emphasis id="FDBBDB1CE87CFFCB766FFB72F4D8FB21" bold="true" box="[578,737,1201,1227]" italics="true" pageId="1" pageNumber="3042">
(
<figureCitation id="57F41B8BE87CFFCB7666FB72F4E1FB21" box="[587,728,1201,1227]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="3.[160,225,1482,1505]" captionStartId-1="4.[159,224,1339,1362]" captionStartId-2="5.[160,225,1153,1176]" captionStartId-3="6.[160,225,1638,1661]" captionTargetBox-0="[189,1151,146,1437]" captionTargetBox-1="[165,1117,215,1304]" captionTargetBox-2="[164,1151,143,1026]" captionTargetBox-3="[165,1061,144,1603]" captionTargetId-0="figure-45@3.[163,1153,142,1450]" captionTargetId-1="figure-85@4.[163,1153,142,1307]" captionTargetId-2="figure-117@5.[163,1153,142,1121]" captionTargetId-3="figure-1@6.[163,1153,142,1606]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Figure 1. Monstrilla albatrossi sp. nov., holotype adult male from Mindanao, Philippines. (a) Habitus, dorsal view; (b) same, lateral view; (c) same, ventral view. Scale bars: 100 μm." captionText-1="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" captionText-2="Figure 3. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Right antennule, ventral view; (b) left antennule, dorsal view. Setal elements on segments 1–4 labelled following nomenclature by Grygier and Ohtsuka (1995); armature of fifth segment labelled following Huys et al. (2007); ae2 on distal segment follows Huys et al.’s (2007) sequence notation of antennulary aesthetascs. Arrows indicate proximal rounded protuberance on fifth antennulary segment. Scale bars: 75 μm." captionText-3="Figure 4. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Leg 1, anterior view; (b) leg 2, anterior view; (c) leg 3, anterior view; (d) leg 4 anterior view. Scale bars: 75 μm." figureDoi-0="http://doi.org/10.5281/zenodo.6456864" figureDoi-1="http://doi.org/10.5281/zenodo.6456866" figureDoi-2="http://doi.org/10.5281/zenodo.6456868" figureDoi-3="http://doi.org/10.5281/zenodo.6456870" httpUri-0="https://zenodo.org/record/6456864/files/figure.png" httpUri-1="https://zenodo.org/record/6456866/files/figure.png" httpUri-2="https://zenodo.org/record/6456868/files/figure.png" httpUri-3="https://zenodo.org/record/6456870/files/figure.png" pageId="1" pageNumber="3042">Figures 1–4</figureCitation>
)
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<emphasis id="FDBBDB1CE87CFFCB748DFAD0F746FAC7" bold="true" box="[160,383,1299,1325]" italics="true" pageId="1" pageNumber="3042">Material examined</emphasis>
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<materialsCitation id="7FA70D53E87CFFCB748DFAF5F521FA98" ID-GBIF-Occurrence="3751237301" collectionCode="USNM" pageId="1" pageNumber="3042" specimenCode="USNM-74005" specimenCount="1" typeStatus="holotype">
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<typeStatus id="1074B9ACE87CFFCB748DFAF5F734FABA" box="[160,269,1334,1360]" pageId="1" pageNumber="3042" type="holotype">Holotype</typeStatus>
.
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<specimenCount id="D9C9CC87E87CFFCB750CFAF4F7EEFABA" box="[289,471,1335,1360]" pageId="1" pageNumber="3042">One adult male</specimenCount>
, vial deposited at the National Museum of Natural History, Smithsonian Institution, Washington, DC (
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)
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.
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<emphasis id="FDBBDB1CE87CFFCB748DFA6AF70FFA29" bold="true" box="[160,310,1449,1475]" italics="true" pageId="1" pageNumber="3042">
<typeStatus id="1074B9ACE87CFFCB748DFA6AF6E1FA29" box="[160,216,1449,1475]" pageId="1" pageNumber="3042">Type</typeStatus>
locality
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Panabutan Pt., west coast of Mindanao, 
<collectingCountry id="B7D8479EE87CFFCB7643FA0FF4D5FA0F" box="[622,748,1484,1509]" name="Philippines" pageId="1" pageNumber="3042">Philippines</collectingCountry>
, plankton trawl, surface waters. R/V 
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cruise sta. 5133 (~ 
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7° 35 
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06.39 
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N
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, 
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122° 08 
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05.31 
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E
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; coll. 
<date id="BB7121CEE87CFFCB7741FA2CF21DF9E2" box="[876,1060,1519,1544]" pageId="1" pageNumber="3042" value="1908-02-06">6 February 1908</date>
)).
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<subSubSection id="87D55485E87CFFCB748DF9FDF57CF92A" pageId="1" pageNumber="3042" type="etymology">
<paragraph id="CF70070EE87CFFCB748DF9FDF724F9B2" blockId="1.[160,1156,1598,1728]" box="[160,285,1598,1624]" pageId="1" pageNumber="3042">
<emphasis id="FDBBDB1CE87CFFCB748DF9FDF724F9B2" bold="true" box="[160,285,1598,1624]" italics="true" pageId="1" pageNumber="3042">Etymology</emphasis>
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The species is named in reference to the illustrious steamer R/V 
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of the 
<collectingCountry id="B7D8479EE87CFFCB7015F9A1F6DDF974" name="United States of America" pageId="1" pageNumber="3042">United States</collectingCountry>
Commission of Fish and Fisheries, operating in the western Pacific in 1887–1909 and sampling the rich pelagic copepod fauna of this region.
</paragraph>
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<emphasis id="FDBBDB1CE87FFFC8748DFF4DF72DFF42" bold="true" box="[160,276,142,168]" italics="true" pageId="2" pageNumber="3043">Diagnosis</emphasis>
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Male 
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<emphasis id="FDBBDB1CE87FFFC874CDFF71F775FF21" box="[224,332,178,203]" italics="true" pageId="2" pageNumber="3043">Monstrilla</emphasis>
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with distinct cuticular ornamentation of cephalothorax represented by wide, complete belt-like striation pattern covering 1/3 of post-oral cephalothoracic surface plus intercalate fields of small papillae on dorsal, ventral, and lateral surfaces, fifth antennulary segment with rounded proximal protuberance. Genital complex 
<typeStatus id="1074B9ACE87FFFC87029FED9F20FFED9" box="[1028,1078,282,307]" pageId="2" pageNumber="3043">type</typeStatus>
II with corrugate margin connecting both lappets and acute apical opercular processes. Fifth legs represented by pair of rounded processes on ventral surface of fifth pedigerous somite.
</paragraph>
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<paragraph id="CF70070EE87FFFC8748DFE77F7F4FE24" blockId="2.[160,1156,436,1709]" box="[160,461,436,462]" pageId="2" pageNumber="3043">
<emphasis id="FDBBDB1CE87FFFC8748DFE77F7F4FE24" bold="true" box="[160,461,436,462]" italics="true" pageId="2" pageNumber="3043">Description of adult male</emphasis>
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Body shape and tagmosis as usual in male 
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<emphasis id="FDBBDB1CE87FFFC8765BFE14F4D9FE1A" box="[630,736,471,496]" italics="true" pageId="2" pageNumber="3043">Monstrilla</emphasis>
</taxonomicName>
(Suárez-Morales 2000; 
<bibRefCitation id="AB5E7AFFE87FFFC877F1FE14F7EFFDF9" author="Suarez-Morales E &amp; Castellanos-Osorio I" pageId="2" pageNumber="3043" pagination="111 - 123" refId="ref5016" refString="Suarez-Morales E, Castellanos-Osorio I. 2019. A new species of Monstrilla (Copepoda, Monstrilloida, Monstrillidae) from the plankton of a bay system of the northwestern Caribbean, with a key to species. ZooKeys. 876: 111 - 123. doi: 10.3897 / zookeys. 876.38400" type="journal article" year="2019">Suárez-Morales and Castellanos-Osorio 2019</bibRefCitation>
). Total body length (measured in dorsal view from anterior end of cephalothorax to posterior margin of anal somite): 
<quantity id="0837AAEBE87FFFC876D3FDDFF55BFDDF" box="[766,866,540,565]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="3043" unit="mm" value="1.25">1.25 mm</quantity>
. First pedigerous thoracic somite incorporated into cephalothorax. Cephalothorax short, relatively robust, representing slightly more than 47% of total body length (
<figureCitation id="57F41B8BE87FFFC87681FDA1F539FD91" box="[684,768,610,635]" captionStart="Figure 1" captionStartId="3.[160,225,1482,1505]" captionTargetBox="[189,1151,146,1437]" captionTargetId="figure-45@3.[163,1153,142,1450]" captionTargetPageId="3" captionText="Figure 1. Monstrilla albatrossi sp. nov., holotype adult male from Mindanao, Philippines. (a) Habitus, dorsal view; (b) same, lateral view; (c) same, ventral view. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6456864" httpUri="https://zenodo.org/record/6456864/files/figure.png" pageId="2" pageNumber="3043">Figure 1</figureCitation>
(a–c)). Oral cone located 45% of the way back along ventral surface of cephalothorax (oc, 
<figureCitation id="57F41B8BE87FFFC876D4FD47F562FD77" box="[761,859,644,669]" captionStart="Figure 1" captionStartId="3.[160,225,1482,1505]" captionTargetBox="[189,1151,146,1437]" captionTargetId="figure-45@3.[163,1153,142,1450]" captionTargetPageId="3" captionText="Figure 1. Monstrilla albatrossi sp. nov., holotype adult male from Mindanao, Philippines. (a) Habitus, dorsal view; (b) same, lateral view; (c) same, ventral view. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6456864" httpUri="https://zenodo.org/record/6456864/files/figure.png" pageId="2" pageNumber="3043">Figures 1</figureCitation>
(a, b) and 2(b,c)). Cephalic region anteriorly wide, forehead flat. Ocelli present, pigment cups small, round in dorsal view, separated by one eye diameter, poorly developed, almost unpigmented. Small oval hyaline bodies (hb in 
<figureCitation id="57F41B8BE87FFFC875B7FD2FF7C1FCEF" box="[410,504,748,773]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="2" pageNumber="3043">Figure 2</figureCitation>
(d)) (see 
<bibRefCitation id="AB5E7AFFE87FFFC87676FD2FF57BFCEF" author="Suarez-Morales E" box="[603,834,748,773]" pageId="2" pageNumber="3043" pagination="92 - 103" refId="ref4924" refString="Suarez-Morales E. 2018. Monstrilloid copepods: the best of three worlds. Bull South Calif Acad Sci. 107 (2): 92 - 103." type="journal article" year="2018">Suárez-Morales 2018</bibRefCitation>
) adjacent to rounded ocelli. Cuticular ornamentation of cephalothorax including field of transverse papillae on dorsal forehead surface (
<figureCitation id="57F41B8BE87FFFC87541FCF1F7F3FCA1" box="[364,458,818,843]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="2" pageNumber="3043">Figure 2</figureCitation>
(d)). In addition, usual pair of nipple-like processes on ventral surface between oral cone and antennule bases; processes small, with adjacent field of faint cuticular striae (
<figureCitation id="57F41B8BE87FFFC8757EFCB4F796FC7A" box="[339,431,887,912]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="2" pageNumber="3043">Figure 2</figureCitation>
(a–c)). Cephalothorax with continuous belt of shallow transverse cuticular striae covering nearly 1/3 of its post-oral half (sb in 
<figureCitation id="57F41B8BE87FFFC8771EFC59F5B5FC59" box="[819,908,922,947]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="2" pageNumber="3043">Figure 2</figureCitation>
(a–c)); cephalothoracic striated belt with intercalate fields of small scattered papillae on ventral, dorsal and lateral surfaces (if in 
<figureCitation id="57F41B8BE87FFFC87565FC1CF790FC12" box="[328,425,991,1016]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="2" pageNumber="3043">Figure 2</figureCitation>
(a–c)) Oral cone weakly produced, with adjacent field of faint, transverse striae (oc in 
<figureCitation id="57F41B8BE87FFFC8758AFBC1F43FFBF1" box="[423,518,1026,1051]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="2" pageNumber="3043">Figure 2</figureCitation>
(b, c)). Rounded medial protuberance present on ventral surface between nipple-like processes and antennule bases; protuberance moderately produced, with notch, visible in lateral view (mp in 
<figureCitation id="57F41B8BE87FFFC876FBFB84F508FB8A" box="[726,817,1095,1120]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="2" pageNumber="3043">Figure 2</figureCitation>
(b, c)).
</paragraph>
<paragraph id="CF70070EE87FFFC874EDFBA9F5C5FB22" blockId="2.[160,1156,436,1709]" pageId="2" pageNumber="3043">
Urosome consisting of five somites: fifth pedigerous somite (largest of urosome), genital somite (with genital complex), two free somites and short anal somite. Ratio of lengths of urosomites (from proximal to distal): 26.8:23.3:25:17.7:7.2 = 100 (
<figureCitation id="57F41B8BE87FFFC87748FB6CF586FB22" box="[869,959,1199,1224]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="2" pageNumber="3043">Figure 2</figureCitation>
(e, f)).
</paragraph>
<paragraph id="CF70070EE87FFFC974EDFB11F26BF977" blockId="2.[160,1156,436,1709]" lastBlockId="3.[160,1155,1599,1693]" lastPageId="3" lastPageNumber="3044" pageId="2" pageNumber="3043">
Antennules representing about 35% of total body length, and almost 72% of cephalothorax length. As usual in male 
<taxonomicName id="08CF7C8DE87FFFC8761EFB37F4A6FAE7" authorityName="Dana" authorityYear="1849" box="[563,671,1268,1293]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="2" pageNumber="3043" phylum="Arthropoda" rank="genus">
<emphasis id="FDBBDB1CE87FFFC8761EFB37F4A6FAE7" box="[563,671,1268,1293]" italics="true" pageId="2" pageNumber="3043">Monstrilla</emphasis>
</taxonomicName>
, antennules five-segmented, geniculation between segments 4 and 5 (
<figureCitation id="57F41B8BE87FFFC875C6FAD4F470FADA" box="[491,585,1303,1328]" captionStart="Figure 3" captionStartId="5.[160,225,1153,1176]" captionTargetBox="[164,1151,143,1026]" captionTargetId="figure-117@5.[163,1153,142,1121]" captionTargetPageId="5" captionText="Figure 3. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Right antennule, ventral view; (b) left antennule, dorsal view. Setal elements on segments 1–4 labelled following nomenclature by Grygier and Ohtsuka (1995); armature of fifth segment labelled following Huys et al. (2007); ae2 on distal segment follows Huys et al.’s (2007) sequence notation of antennulary aesthetascs. Arrows indicate proximal rounded protuberance on fifth antennulary segment. Scale bars: 75 μm." figureDoi="http://doi.org/10.5281/zenodo.6456868" httpUri="https://zenodo.org/record/6456868/files/figure.png" pageId="2" pageNumber="3043">Figure 3</figureCitation>
(a, b)). Following setal nomenclature proposed by 
<bibRefCitation id="AB5E7AFFE87FFFC8748DFAF9F7E0FAB9" author="Grygier MJ &amp; Ohtsuka S" box="[160,473,1338,1363]" pageId="2" pageNumber="3043" pagination="703 - 719" refId="ref4307" refString="Grygier MJ, Ohtsuka S. 1995. SEM observation of the nauplius of Monstrilla hamatapex, new species, from Japan and an example of upgraded descriptive standards for monstrilloid copepods. J. Crustacean Biol. 15: 703 - 719. https: // doi. org / 10.1163 / 193724095 X 00118" type="journal article" year="1995">Grygier and Ohtsuka (1995)</bibRefCitation>
for female monstrilloid copepod antennules, seta 1 moderately long, almost reaching half length of succeeding second segment. Elements 2v1–3 and 2d1,2 setiform, long, element IId long, thick, with protuberant distal socket. Third segment short, subquadrate, with setiform, flexible medial element 3 reaching about proximal 1/3 of fourth segment. Setae IIId, IIIv present, long. Fourth segment armed with nine elements: 4d1,2, 4aes, 4v1–3, IVd, Vv, Vm. Elements 4v1,2 twice as long as element 4v3. Distal segment lacking modified setae, with conspicuous rounded proximal protuberance (arrows in 
<figureCitation id="57F41B8BE87FFFC87533F9EFF747F9AF" box="[286,382,1580,1605]" captionStart="Figure 3" captionStartId="5.[160,225,1153,1176]" captionTargetBox="[164,1151,143,1026]" captionTargetId="figure-117@5.[163,1153,142,1121]" captionTargetPageId="5" captionText="Figure 3. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Right antennule, ventral view; (b) left antennule, dorsal view. Setal elements on segments 1–4 labelled following nomenclature by Grygier and Ohtsuka (1995); armature of fifth segment labelled following Huys et al. (2007); ae2 on distal segment follows Huys et al.’s (2007) sequence notation of antennulary aesthetascs. Arrows indicate proximal rounded protuberance on fifth antennulary segment. Scale bars: 75 μm." figureDoi="http://doi.org/10.5281/zenodo.6456868" httpUri="https://zenodo.org/record/6456868/files/figure.png" pageId="2" pageNumber="3043">Figure 3</figureCitation>
(a, b)) and armed with 11 setal elements (sensu 
<bibRefCitation id="AB5E7AFFE87FFFC87790F9EFF245F9AF" author="Huys R &amp; Llewellyn-Hughes J &amp; Conroy-Dalton S &amp; Olson PD &amp; Spinks JN &amp; Johnston DA" box="[957,1148,1580,1605]" pageId="2" pageNumber="3043" pagination="368 - 378" refId="ref4384" refString="Huys R, Llewellyn-Hughes J, Conroy-Dalton S, Olson PD, Spinks JN, Johnston DA. 2007. Extraordinary host switching in siphonostomatoid copepods and the demise of the Monstrilloida: integrating molecular data, ontogeny and antennulary morphology. Mol Phylogenet Evol. 43: 368 - 378. doi: 10.1016 / j. ympev. 2007.02.004" type="journal article" year="2007">Huys et al. 2007</bibRefCitation>
) including setae 1–6 and unmodified elements A–C plus short, slender aesthetasc 6aes (sensu 
<bibRefCitation id="AB5E7AFFE87FFFC874DEF9B1F41DF961" author="Grygier MJ &amp; Ohtsuka S" box="[243,548,1650,1675]" pageId="2" pageNumber="3043" pagination="703 - 719" refId="ref4307" refString="Grygier MJ, Ohtsuka S. 1995. SEM observation of the nauplius of Monstrilla hamatapex, new species, from Japan and an example of upgraded descriptive standards for monstrilloid copepods. J. Crustacean Biol. 15: 703 - 719. https: // doi. org / 10.1163 / 193724095 X 00118" type="journal article" year="1995">Grygier and Ohtsuka 1995</bibRefCitation>
), or ae2 (sensu 
<bibRefCitation id="AB5E7AFFE87FFFC876C8F9B1F59AF961" author="Huys R &amp; Llewellyn-Hughes J &amp; Conroy-Dalton S &amp; Olson PD &amp; Spinks JN &amp; Johnston DA" box="[741,931,1650,1675]" pageId="2" pageNumber="3043" pagination="368 - 378" refId="ref4384" refString="Huys R, Llewellyn-Hughes J, Conroy-Dalton S, Olson PD, Spinks JN, Johnston DA. 2007. Extraordinary host switching in siphonostomatoid copepods and the demise of the Monstrilloida: integrating molecular data, ontogeny and antennulary morphology. Mol Phylogenet Evol. 43: 368 - 378. doi: 10.1016 / j. ympev. 2007.02.004" type="journal article" year="2007">Huys et al. 2007</bibRefCitation>
) in apical position. Sockets on middle and distal surfaces of fourth and fifth antennulary segments (indicated by asterisks in 
<figureCitation id="57F41B8BE87EFFC9756FF9FCF7A6F9B2" box="[322,415,1599,1624]" captionStart="Figure 3" captionStartId="5.[160,225,1153,1176]" captionTargetBox="[164,1151,143,1026]" captionTargetId="figure-117@5.[163,1153,142,1121]" captionTargetPageId="5" captionText="Figure 3. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Right antennule, ventral view; (b) left antennule, dorsal view. Setal elements on segments 1–4 labelled following nomenclature by Grygier and Ohtsuka (1995); armature of fifth segment labelled following Huys et al. (2007); ae2 on distal segment follows Huys et al.’s (2007) sequence notation of antennulary aesthetascs. Arrows indicate proximal rounded protuberance on fifth antennulary segment. Scale bars: 75 μm." figureDoi="http://doi.org/10.5281/zenodo.6456868" httpUri="https://zenodo.org/record/6456868/files/figure.png" pageId="3" pageNumber="3044">Figure 3</figureCitation>
(b)) suggest loss of some (likely four) setal elements, particularly of the ‘b’ group (sensu 
<bibRefCitation id="AB5E7AFFE87EFFC975A9F9A1F49EF991" author="Grygier MJ &amp; Ohtsuka S" box="[388,679,1634,1659]" pageId="3" pageNumber="3044" pagination="703 - 719" refId="ref4307" refString="Grygier MJ, Ohtsuka S. 1995. SEM observation of the nauplius of Monstrilla hamatapex, new species, from Japan and an example of upgraded descriptive standards for monstrilloid copepods. J. Crustacean Biol. 15: 703 - 719. https: // doi. org / 10.1163 / 193724095 X 00118" type="journal article" year="1995">Grygier and Ohtsuka 1995</bibRefCitation>
) in the 
<typeStatus id="1074B9ACE87EFFC976D6F9A1F566F991" box="[763,863,1634,1659]" pageId="3" pageNumber="3044" type="holotype">holotype</typeStatus>
specimen. Length ratio of antennular segments (proximal to distal): 11.4:19.8:11.4:27.8:29.6 = 100 (
<figureCitation id="57F41B8BE87EFFC977E1F947F210F977" box="[972,1065,1668,1693]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="3" pageNumber="3044">Figure 2</figureCitation>
(a)).
</paragraph>
<caption id="9BB05786E87EFFC9748DFA09F59BFA14" ID-DOI="http://doi.org/10.5281/zenodo.6456864" ID-Zenodo-Dep="6456864" httpUri="https://zenodo.org/record/6456864/files/figure.png" pageId="3" pageNumber="3044" startId="3.[160,225,1482,1505]" targetBox="[189,1151,146,1437]" targetPageId="3">
<paragraph id="CF70070EE87EFFC9748DFA09F59BFA14" blockId="3.[160,1157,1482,1534]" pageId="3" pageNumber="3044">
<emphasis id="FDBBDB1CE87EFFC9748DFA09F6C2FA0B" bold="true" box="[160,251,1482,1505]" pageId="3" pageNumber="3044">Figure 1.</emphasis>
<taxonomicName id="08CF7C8DE87EFFC9752FFA09F7FCFA0B" authority="Suárez-Morales, 2022" authorityName="Suárez-Morales" authorityYear="2022" box="[258,453,1482,1505]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="3" pageNumber="3044" phylum="Arthropoda" rank="species" species="albatrossi" status="sp. nov.">
<emphasis id="FDBBDB1CE87EFFC9752FFA09F7FCFA0B" box="[258,453,1482,1505]" italics="true" pageId="3" pageNumber="3044">Monstrilla albatrossi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6886667E87EFFC975E1FA09F425FA0B" box="[460,540,1482,1505]" pageId="3" pageNumber="3044" rank="species">sp. nov.</taxonomicNameLabel>
, holotype adult male from Mindanao, Philippines. (a) Habitus, dorsal view; (b) same, lateral view; (c) same, ventral view. Scale bars: 100 μm.
</paragraph>
</caption>
<caption id="9BB05786E879FFCE74B2FAF8F5F4FA40" ID-DOI="http://doi.org/10.5281/zenodo.6456866" ID-Zenodo-Dep="6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="4" pageNumber="3045" startId="4.[159,224,1339,1362]" targetBox="[165,1117,215,1304]" targetPageId="4">
<paragraph id="CF70070EE879FFCE74B2FAF8F5F4FA40" blockId="4.[159,1156,1339,1450]" pageId="4" pageNumber="3045">
<emphasis id="FDBBDB1CE879FFCE74B2FAF8F6C5FAB8" bold="true" box="[159,252,1339,1362]" pageId="4" pageNumber="3045">Figure 2.</emphasis>
<taxonomicName id="08CF7C8DE879FFCE7529FAF8F7FEFAB8" authority="Suárez-Morales, 2022" authorityName="Suárez-Morales" authorityYear="2022" box="[260,455,1339,1362]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="4" pageNumber="3045" phylum="Arthropoda" rank="species" species="albatrossi" status="sp. nov.">
<emphasis id="FDBBDB1CE879FFCE7529FAF8F7FEFAB8" box="[260,455,1339,1362]" italics="true" pageId="4" pageNumber="3045">Monstrilla albatrossi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6886667E879FFCE75FDFAF8F418FAB8" box="[464,545,1339,1362]" pageId="4" pageNumber="3045" rank="species">sp. nov.</taxonomicNameLabel>
, holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu 
<bibRefCitation id="AB5E7AFFE879FFCE76E4FAB5F5A1FA67" author="Suarez-Morales E" box="[713,920,1398,1421]" pageId="4" pageNumber="3045" pagination="92 - 103" refId="ref4924" refString="Suarez-Morales E. 2018. Monstrilloid copepods: the best of three worlds. Bull South Calif Acad Sci. 107 (2): 92 - 103." type="journal article" year="2018">Suárez-Morales 2018</bibRefCitation>
): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm
</paragraph>
</caption>
<paragraph id="CF70070EE879FFCE74EDFA39F2BDF92A" blockId="4.[159,1156,1530,1728]" pageId="4" pageNumber="3045">
First incorporated pedigerous thoracic somite and succeeding three pedigers each bearing well-developed biramous swimming legs (
<figureCitation id="57F41B8BE879FFCE764EF9DEF48CF9DC" box="[611,693,1565,1590]" captionStart="Figure 1" captionStartId="3.[160,225,1482,1505]" captionTargetBox="[189,1151,146,1437]" captionTargetId="figure-45@3.[163,1153,142,1450]" captionTargetPageId="3" captionText="Figure 1. Monstrilla albatrossi sp. nov., holotype adult male from Mindanao, Philippines. (a) Habitus, dorsal view; (b) same, lateral view; (c) same, ventral view. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6456864" httpUri="https://zenodo.org/record/6456864/files/figure.png" pageId="4" pageNumber="3045">Figure 1</figureCitation>
(b, c)). Swimming legs 1–4 with triarticulate endopodites and exopodites and same armament pattern, except for leg 1 exopodite, with one seta fewer on distal segment than in other legs (
<figureCitation id="57F41B8BE879FFCE76DAF9A1F568F991" box="[759,849,1634,1659]" captionStart="Figure 4" captionStartId="6.[160,225,1638,1661]" captionTargetBox="[165,1061,144,1603]" captionTargetId="figure-1@6.[163,1153,142,1606]" captionTargetPageId="6" captionText="Figure 4. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Leg 1, anterior view; (b) leg 2, anterior view; (c) leg 3, anterior view; (d) leg 4 anterior view. Scale bars: 75 μm." figureDoi="http://doi.org/10.5281/zenodo.6456870" httpUri="https://zenodo.org/record/6456870/files/figure.png" pageId="4" pageNumber="3045">Figure 4</figureCitation>
(a–d)). Exopods longer than endopods. Distal spiniform seta of third exopod with denticles along outer margin, inner margin weakly setulose in all legs. Coxae joined by smooth rectangular coupler (
<figureCitation id="57F41B8BE879FFCE7028F964F261F92A" box="[1029,1112,1703,1728]" captionStart="Figure 1" captionStartId="3.[160,225,1482,1505]" captionTargetBox="[189,1151,146,1437]" captionTargetId="figure-45@3.[163,1153,142,1450]" captionTargetPageId="3" captionText="Figure 1. Monstrilla albatrossi sp. nov., holotype adult male from Mindanao, Philippines. (a) Habitus, dorsal view; (b) same, lateral view; (c) same, ventral view. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6456864" httpUri="https://zenodo.org/record/6456864/files/figure.png" pageId="4" pageNumber="3045">Figure 1</figureCitation>
(c)).
</paragraph>
<caption id="9BB05786E878FFCF748DFB42F45AFAC0" ID-DOI="http://doi.org/10.5281/zenodo.6456868" ID-Zenodo-Dep="6456868" httpUri="https://zenodo.org/record/6456868/files/figure.png" pageId="5" pageNumber="3046" startId="5.[160,225,1153,1176]" targetBox="[164,1151,143,1026]" targetPageId="5">
<paragraph id="CF70070EE878FFCF748DFB42F45AFAC0" blockId="5.[160,1156,1153,1322]" pageId="5" pageNumber="3046">
<emphasis id="FDBBDB1CE878FFCF748DFB42F6C5FB72" bold="true" box="[160,252,1153,1176]" pageId="5" pageNumber="3046">Figure 3.</emphasis>
<taxonomicName id="08CF7C8DE878FFCF7529FB42F7FEFB72" authority="Suárez-Morales, 2022" authorityName="Suárez-Morales" authorityYear="2022" box="[260,455,1153,1176]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="5" pageNumber="3046" phylum="Arthropoda" rank="species" species="albatrossi" status="sp. nov.">
<emphasis id="FDBBDB1CE878FFCF7529FB42F7FEFB72" box="[260,455,1153,1176]" italics="true" pageId="5" pageNumber="3046">Monstrilla albatrossi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6886667E878FFCF75FDFB42F418FB72" box="[464,545,1153,1176]" pageId="5" pageNumber="3046" rank="species">sp. nov.</taxonomicNameLabel>
, holotype adult male from off Mindanao, Philippines, female holotype. (a) Right antennule, ventral view; (b) left antennule, dorsal view. Setal elements on segments 1–4 labelled following nomenclature by 
<bibRefCitation id="AB5E7AFFE878FFCF7683FB78F5F3FB38" author="Grygier MJ &amp; Ohtsuka S" box="[686,970,1211,1234]" pageId="5" pageNumber="3046" pagination="703 - 719" refId="ref4307" refString="Grygier MJ, Ohtsuka S. 1995. SEM observation of the nauplius of Monstrilla hamatapex, new species, from Japan and an example of upgraded descriptive standards for monstrilloid copepods. J. Crustacean Biol. 15: 703 - 719. https: // doi. org / 10.1163 / 193724095 X 00118" type="journal article" year="1995">Grygier and Ohtsuka (1995)</bibRefCitation>
; armature of fifth segment labelled following 
<bibRefCitation id="AB5E7AFFE878FFCF7590FB1AF44AFB1A" author="Huys R &amp; Llewellyn-Hughes J &amp; Conroy-Dalton S &amp; Olson PD &amp; Spinks JN &amp; Johnston DA" box="[445,627,1241,1264]" pageId="5" pageNumber="3046" pagination="368 - 378" refId="ref4384" refString="Huys R, Llewellyn-Hughes J, Conroy-Dalton S, Olson PD, Spinks JN, Johnston DA. 2007. Extraordinary host switching in siphonostomatoid copepods and the demise of the Monstrilloida: integrating molecular data, ontogeny and antennulary morphology. Mol Phylogenet Evol. 43: 368 - 378. doi: 10.1016 / j. ympev. 2007.02.004" type="journal article" year="2007">Huys et al. (2007)</bibRefCitation>
; ae2 on distal segment follows Huys et al.’s (2007) sequence notation of antennulary aesthetascs. Arrows indicate proximal rounded protuberance on fifth antennulary segment. Scale bars: 75 μm.
</paragraph>
</caption>
<paragraph id="CF70070EE878FFCF748DFAA1F575FA2A" blockId="5.[159,1155,1378,1595]" pageId="5" pageNumber="3046">
Basis separated from coxa by diagonal articulation. Basis of swimming legs 1, 2 and 4 with short basipodal seta; seta on leg 3 being thicker and 2.5–3 times as long as in other legs (
<figureCitation id="57F41B8BE878FFCF748AFA64F6C6FA2A" box="[167,255,1447,1472]" captionStart="Figure 4" captionStartId="6.[160,225,1638,1661]" captionTargetBox="[165,1061,144,1603]" captionTargetId="figure-1@6.[163,1153,142,1606]" captionTargetPageId="6" captionText="Figure 4. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Leg 1, anterior view; (b) leg 2, anterior view; (c) leg 3, anterior view; (d) leg 4 anterior view. Scale bars: 75 μm." figureDoi="http://doi.org/10.5281/zenodo.6456870" httpUri="https://zenodo.org/record/6456870/files/figure.png" pageId="5" pageNumber="3046">Figure 4</figureCitation>
(a–d)). All natatory setae lightly and biserially plumose.
</paragraph>
<paragraph id="CF70070EE878FFCF74EDFA09F572F9D1" blockId="5.[159,1155,1378,1595]" pageId="5" pageNumber="3046">Armament formula of legs 1–4: basis exopod endopod Leg 1 0-1 I-1; 0-1; I, 2, 2 0-1; 0-1; 1, 2, 2 Legs 2–4 0-1 I-1; 0-1; I, 2, 3 0-1; 0-1;1,2,2</paragraph>
<paragraph id="CF70070EE878FFCD74EDF9A1F20FFF07" blockId="5.[160,1155,1634,1728]" lastBlockId="7.[160,1156,142,375]" lastPageId="7" lastPageNumber="3048" pageId="5" pageNumber="3046">
Fifth legs absent, but represented by pair of low rounded protuberances on ventral surface of fifth pedigerous somite (arrows in 
<figureCitation id="57F41B8BE878FFCF76BDF947F4D2F977" box="[656,747,1668,1693]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="5" pageNumber="3046">Figure 2</figureCitation>
(e, f)). Genital somite carrying genital complex consisting of short, thick medially curved shaft bearing paired distal genital lappets reaching distal margin of succeeding urosomite (
<figureCitation id="57F41B8BE87AFFCD773AFF4DF548FF4D" box="[791,881,142,167]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="7" pageNumber="3048">Figure 2</figureCitation>
(e, f)). Lappets divergent, digitiform, tapering distally into apical acute opercular processes (op in 
<figureCitation id="57F41B8BE87AFFCD77DAFF72F260FF20" box="[1015,1113,177,202]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="7" pageNumber="3048">Figure 2</figureCitation>
(g)); margin connecting both lappets curved, coarsely and evenly rugose (
<figureCitation id="57F41B8BE87AFFCD7783FF17F235FF07" box="[942,1036,212,237]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="7" pageNumber="3048">Figure 2</figureCitation>
(g)).
</paragraph>
<caption id="9BB05786E87BFFCC748DF9A5F7B6F952" ID-DOI="http://doi.org/10.5281/zenodo.6456870" ID-Zenodo-Dep="6456870" httpUri="https://zenodo.org/record/6456870/files/figure.png" pageId="6" pageNumber="3047" startId="6.[160,225,1638,1661]" targetBox="[165,1061,144,1603]" targetPageId="6">
<paragraph id="CF70070EE87BFFCC748DF9A5F7B6F952" blockId="6.[160,1156,1638,1720]" pageId="6" pageNumber="3047">
<emphasis id="FDBBDB1CE87BFFCC748DF9A5F6C5F997" bold="true" box="[160,252,1638,1661]" pageId="6" pageNumber="3047">Figure 4.</emphasis>
<taxonomicName id="08CF7C8DE87BFFCC7529F9A5F7FEF997" authority="Suárez-Morales, 2022" authorityName="Suárez-Morales" authorityYear="2022" box="[260,455,1638,1661]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="6" pageNumber="3047" phylum="Arthropoda" rank="species" species="albatrossi" status="sp. nov.">
<emphasis id="FDBBDB1CE87BFFCC7529F9A5F7FEF997" box="[260,455,1638,1661]" italics="true" pageId="6" pageNumber="3047">Monstrilla albatrossi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6886667E87BFFCC75FDF9A5F418F997" box="[464,545,1638,1661]" pageId="6" pageNumber="3047" rank="species">sp. nov.</taxonomicNameLabel>
, holotype adult male from off Mindanao, Philippines, female holotype. (a) Leg 1, anterior view; (b) leg 2, anterior view; (c) leg 3, anterior view; (d) leg 4 anterior view. Scale bars: 75 μm.
</paragraph>
</caption>
<paragraph id="CF70070EE87AFFCD74EDFF35F496FE9D" blockId="7.[160,1156,142,375]" pageId="7" pageNumber="3048">
Caudal rami subrectangular, with distal margin slightly wider than proximal, ramus approximately 1.3 times as long as wide. Each ramus bearing six well-developed caudal setae (setae I–VI sensu 
<bibRefCitation id="AB5E7AFFE87AFFCD75B3FEFFF49CFEBF" author="Huys R &amp; Boxshall GA" box="[414,677,316,341]" pageId="7" pageNumber="3048" refId="ref4363" refString="Huys R, Boxshall GA. 1991. Copepod evolution. London: The Ray Society; p. 468." type="journal volume" year="1991">Huys and Boxshall 1991</bibRefCitation>
), seta VI being shortest. Caudal setae II and III about as long as medial seta IV (
<figureCitation id="57F41B8BE87AFFCD7604FE9DF4B9FE9D" box="[553,640,350,375]" captionStart="Figure 1" captionStartId="3.[160,225,1482,1505]" captionTargetBox="[189,1151,146,1437]" captionTargetId="figure-45@3.[163,1153,142,1450]" captionTargetPageId="3" captionText="Figure 1. Monstrilla albatrossi sp. nov., holotype adult male from Mindanao, Philippines. (a) Habitus, dorsal view; (b) same, lateral view; (c) same, ventral view. Scale bars: 100 μm." figureDoi="http://doi.org/10.5281/zenodo.6456864" httpUri="https://zenodo.org/record/6456864/files/figure.png" pageId="7" pageNumber="3048">Figure 1</figureCitation>
(a)).
</paragraph>
<paragraph id="CF70070EE87AFFCD748DFE60F6CFFE57" blockId="7.[160,275,419,480]" box="[160,246,419,445]" pageId="7" pageNumber="3048">
<emphasis id="FDBBDB1CE87AFFCD748DFE60F6CFFE57" bold="true" box="[160,246,419,445]" italics="true" pageId="7" pageNumber="3048">Female</emphasis>
</paragraph>
<paragraph id="CF70070EE87AFFCD748DFE04F72AFE0A" blockId="7.[160,275,419,480]" box="[160,275,455,480]" pageId="7" pageNumber="3048">Unknown.</paragraph>
</subSubSection>
<subSubSection id="87D55485E87AFFC3748DFDCFF79EFCB7" lastPageId="9" lastPageNumber="3050" pageId="7" pageNumber="3048" type="discussion">
<paragraph id="CF70070EE87AFFCD748DFDCFF73FFDCC" blockId="7.[160,1156,524,1728]" box="[160,262,524,550]" pageId="7" pageNumber="3048">
<emphasis id="FDBBDB1CE87AFFCD748DFDCFF73FFDCC" bold="true" box="[160,262,524,550]" italics="true" pageId="7" pageNumber="3048">Remarks</emphasis>
</paragraph>
<paragraph id="CF70070EE87AFFCD748DFDECF4E0FCD1" blockId="7.[160,1156,524,1728]" pageId="7" pageNumber="3048">
The 
<typeStatus id="1074B9ACE87AFFCD74E2FDECF70AFDA2" box="[207,307,559,584]" pageId="7" pageNumber="3048" type="holotype">holotype</typeStatus>
specimen is still in good condition for taxonomic study considering the long time elapsed (113 years) since it was collected. However, the cephalothorax appears damaged (depressed), probably by forceps during handling, and several antennular setal elements are missing, particularly on the last segment. This is why only the structures on antennule segments 1–4 were homologised with the pattern proposed by 
<bibRefCitation id="AB5E7AFFE87AFFCD702CFD79F774FD1F" author="Grygier MJ &amp; Ohtsuka S" pageId="7" pageNumber="3048" pagination="703 - 719" refId="ref4307" refString="Grygier MJ, Ohtsuka S. 1995. SEM observation of the nauplius of Monstrilla hamatapex, new species, from Japan and an example of upgraded descriptive standards for monstrilloid copepods. J. Crustacean Biol. 15: 703 - 719. https: // doi. org / 10.1163 / 193724095 X 00118" type="journal article" year="1995">Grygier and Ohtsuka (1995)</bibRefCitation>
for female monstrilloids. But mixing of the two patterns (i.e. using the female notation for male antennules) should be avoided. The specimen was not dissected during examination and returned whole to its vial.
</paragraph>
<paragraph id="CF70070EE87AFFCD74EDFC86F5FEFAA9" blockId="7.[160,1156,524,1728]" pageId="7" pageNumber="3048">
Prior to this record of 
<taxonomicName id="08CF7C8DE87AFFCD759CFC86F401FCB4" authority="Suárez-Morales, 2022" authorityName="Suárez-Morales" authorityYear="2022" box="[433,568,837,862]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="albatrossi" status="sp. nov.">
<emphasis id="FDBBDB1CE87AFFCD759CFC86F401FCB4" box="[433,568,837,862]" italics="true" pageId="7" pageNumber="3048">M. albatrossi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6886667E87AFFCD7612FC86F4ACFCB4" box="[575,661,837,862]" pageId="7" pageNumber="3048" rank="species">sp. nov.</taxonomicNameLabel>
only six other species of 
<taxonomicName id="08CF7C8DE87AFFCD7787FC86F22DFCB4" authorityName="Dana" authorityYear="1849" box="[938,1044,837,862]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="genus">
<emphasis id="FDBBDB1CE87AFFCD7787FC86F22DFCB4" box="[938,1044,837,862]" italics="true" pageId="7" pageNumber="3048">Monstrilla</emphasis>
</taxonomicName>
had been reported from the 
<collectingCountry id="B7D8479EE87AFFCD7552FCA4F7C0FC6A" box="[383,505,871,896]" name="Philippines" pageId="7" pageNumber="3048">Philippines</collectingCountry>
area (Suárez-Morales 2000): 
<taxonomicName id="08CF7C8DE87AFFCD7760FCA4F2B9FC6A" authority="A. Scott, 1909" authorityName="A. Scott" authorityYear="1909" box="[845,1152,871,896]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="cymbula">
<emphasis id="FDBBDB1CE87AFFCD7760FCA4F5F6FC6A" box="[845,975,871,896]" italics="true" pageId="7" pageNumber="3048">M. cymbula</emphasis>
A. Scott, 1909
</taxonomicName>
, 
<taxonomicName id="08CF7C8DE87AFFCD748DFC49F42BFC49" authority="Suarez-Morales, 1994" authorityName="Suarez-Morales" authorityYear="1994" box="[160,530,906,931]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="FDBBDB1CE87AFFCD748DFC49F718FC49" box="[160,289,906,931]" italics="true" pageId="7" pageNumber="3048">M. elongata</emphasis>
<bibRefCitation id="AB5E7AFFE87AFFCD7505FC49F42BFC49" author="Suarez-Morales E" box="[296,530,906,931]" pageId="7" pageNumber="3048" pagination="262 - 267" refId="ref4592" refString="Suarez-Morales E. 1994. Monstrilla elongata, a new monstrilloid copepod (Crustacea: Copepoda: Monstrilloida) from a reef lagoon of the Caribbean coast of Mexico. Proc Biol Soc Wash. 107 (2): 262 - 267." type="journal article" year="1994">Suárez-Morales, 1994</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="08CF7C8DE87AFFCD760DFC49F5BCFC49" authority="Suarez-Morales, 2000" authorityName="Suarez-Morales" authorityYear="2000" box="[544,901,906,931]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="grygieri">
<emphasis id="FDBBDB1CE87AFFCD760DFC49F4A8FC49" box="[544,657,906,931]" italics="true" pageId="7" pageNumber="3048">M. grygieri</emphasis>
Suárez-Morales, 2000
</taxonomicName>
, 
<taxonomicName id="08CF7C8DE87AFFCD77BDFC49F746FC2C" authority="(Claus, 1863)" baseAuthorityName="Claus" baseAuthorityYear="1863" class="Hexanauplia" family="Monstrillidae" genus="Caromiobenella" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="helgolandica">
<emphasis id="FDBBDB1CE87AFFCD77BDFC49F6D7FC2C" italics="true" pageId="7" pageNumber="3048">Caromiobenella helgolandica</emphasis>
(Claus, 1863)
</taxonomicName>
(as 
<taxonomicName id="08CF7C8DE87AFFCD7584FC6EF46BFC2C" baseAuthorityName="Claus" baseAuthorityYear="1863" box="[425,594,941,966]" class="Hexanauplia" family="Monstrillidae" genus="Caromiobenella" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="helgolandica">
<emphasis id="FDBBDB1CE87AFFCD7584FC6EF46BFC2C" box="[425,594,941,966]" italics="true" pageId="7" pageNumber="3048">M. helgolandica</emphasis>
</taxonomicName>
), 
<taxonomicName id="08CF7C8DE87AFFCD7648FC6EF550FC2C" authority="(Scott, 1909)" baseAuthorityName="Scott" baseAuthorityYear="1909" box="[613,873,941,966]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="leucopis">
<emphasis id="FDBBDB1CE87AFFCD7648FC6EF4E2FC2C" box="[613,731,941,966]" italics="true" pageId="7" pageNumber="3048">M. leucopis</emphasis>
(Scott, 1909)
</taxonomicName>
and 
<taxonomicName id="08CF7C8DE87AFFCD77B2FC6EF2B9FC2C" authority="Dana, 1849" authorityName="Dana" authorityYear="1849" box="[927,1152,941,966]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="viridis">
<emphasis id="FDBBDB1CE87AFFCD77B2FC6EF5C4FC2C" box="[927,1021,941,966]" italics="true" pageId="7" pageNumber="3048">M. viridis</emphasis>
Dana, 1849
</taxonomicName>
. 
<taxonomicName id="08CF7C8DE87AFFCD748DFC0CF755FC02" authorityName="A. Scott" authorityYear="1909" box="[160,364,975,1000]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="cymbula">
<emphasis id="FDBBDB1CE87AFFCD748DFC0CF755FC02" box="[160,364,975,1000]" italics="true" pageId="7" pageNumber="3048">Monstrilla cymbula</emphasis>
</taxonomicName>
was described from a female specimen collected from the 
<collectingRegion id="0D0BC9ECE87AFFCD702DFC0CF209FC02" box="[1024,1072,975,1000]" country="Philippines" name="Sulu" pageId="7" pageNumber="3048">Sulu</collectingRegion>
Islands (Scott, 1909). 
<taxonomicName id="08CF7C8DE87AFFCD751BFC31F7CFFBE1" authorityName="Suarez-Morales" authorityYear="2000" box="[310,502,1010,1035]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="grygieri">
<emphasis id="FDBBDB1CE87AFFCD751BFC31F7CFFBE1" box="[310,502,1010,1035]" italics="true" pageId="7" pageNumber="3048">Monstrilla grygieri</emphasis>
</taxonomicName>
was recorded from a female found in Panabutan, 
<collectingRegion id="0D0BC9ECE87AFFCD7008FC31F26CFBE1" box="[1061,1109,1010,1035]" country="Philippines" name="Sulu" pageId="7" pageNumber="3048">Sulu</collectingRegion>
Sea (see Suárez-Morales 2000). Far East records of 
<taxonomicName id="08CF7C8DE87AFFCD76B2FBD6F52EFBC4" baseAuthorityName="Scott" baseAuthorityYear="1909" box="[671,791,1045,1070]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="leucopis">
<emphasis id="FDBBDB1CE87AFFCD76B2FBD6F52EFBC4" box="[671,791,1045,1070]" italics="true" pageId="7" pageNumber="3048">M. leucopis</emphasis>
</taxonomicName>
(
<bibRefCitation id="AB5E7AFFE87AFFCD7709FBD6F58BFBC4" author="Wilson CB" box="[804,946,1045,1070]" pageId="7" pageNumber="3048" pagination="141 - 441" refId="ref5387" refString="Wilson CB. 1950. Contributions to the biology of the Philippine Archipelago and adjacent regions. Copepods gathered by the United States fisheries steamer &quot; Albatross &quot; from 1887 to 1909, chiefly in the Pacific Ocean. U S Nat Mus Bull. 100: 141 - 441." type="journal article" year="1950">Wilson, 1950</bibRefCitation>
; 
<bibRefCitation id="AB5E7AFFE87AFFCD7793FBD6F26BFBC4" author="McAlice BJ" box="[958,1106,1045,1070]" pageId="7" pageNumber="3048" pagination="627 - 634" refId="ref4499" refString="McAlice BJ. 1985. On the male of Monstrilla helgolandica Claus (Copepoda, Monstrilloida). J Crustacean Biol. 5: 627 - 634. doi: 10.2307 / 1548240" type="journal article" year="1985">McAlice 1985</bibRefCitation>
) are unlikely to represent this species because it appears to be restricted to 
<collectingCountry id="B7D8479EE87AFFCD77E0FBF4F21BFBBA" box="[973,1058,1079,1104]" name="Norway" pageId="7" pageNumber="3048">Norway</collectingCountry>
(SuárezMorales 2010). The record of the Caribbean 
<taxonomicName id="08CF7C8DE87AFFCD76B3FB99F51BFB99" authorityName="Suarez-Morales" authorityYear="1994" box="[670,802,1114,1139]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="FDBBDB1CE87AFFCD76B3FB99F51BFB99" box="[670,802,1114,1139]" italics="true" pageId="7" pageNumber="3048">M. elongata</emphasis>
</taxonomicName>
from the 
<collectingCountry id="B7D8479EE87AFFCD77B7FB99F22BFB99" box="[922,1042,1114,1139]" name="Philippines" pageId="7" pageNumber="3048">Philippines</collectingCountry>
area (see Suárez-Morales 2000) is highly dubious, and the 
<specimenCount id="D9C9CC87E87AFFCD76CFFBBEF56BFB7C" box="[738,850,1149,1174]" pageId="7" pageNumber="3048" type="adult">two adult</specimenCount>
females deposited at the National Museum of Natural History (USNM 73856) most probably represent an undescribed species. The 
<collectingCountry id="B7D8479EE87AFFCD75ADFB01F7C1FB31" box="[384,504,1218,1243]" name="Philippines" pageId="7" pageNumber="3048">Philippines</collectingCountry>
record of 
<taxonomicName id="08CF7C8DE87AFFCD7647FB01F598FB31" baseAuthorityName="Claus" baseAuthorityYear="1863" box="[618,929,1218,1243]" class="Hexanauplia" family="Monstrillidae" genus="Caromiobenella" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="helgolandica">
<emphasis id="FDBBDB1CE87AFFCD7647FB01F598FB31" box="[618,929,1218,1243]" italics="true" pageId="7" pageNumber="3048">Caromiobenella helgolandica</emphasis>
</taxonomicName>
(as 
<taxonomicName id="08CF7C8DE87AFFCD77E0FB01F24FFB31" baseAuthorityName="Claus" baseAuthorityYear="1863" box="[973,1142,1218,1243]" class="Hexanauplia" family="Monstrillidae" genus="Caromiobenella" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="helgolandica">
<emphasis id="FDBBDB1CE87AFFCD77E0FB01F24FFB31" box="[973,1142,1218,1243]" italics="true" pageId="7" pageNumber="3048">M. helgolandica</emphasis>
</taxonomicName>
), also from the 
<collectingRegion id="0D0BC9ECE87AFFCD7568FB26F74CFB14" box="[325,373,1253,1278]" country="Philippines" name="Sulu" pageId="7" pageNumber="3048">Sulu</collectingRegion>
Islands (Scott 1909), is unsurprising because of the presumed wide distribution of this nominal species, currently deemed to represent a species complex (
<bibRefCitation id="AB5E7AFFE87AFFCD7488FAE9F7F3FAA9" author="Grygier MJ &amp; Ohtsuka S" box="[165,458,1322,1347]" pageId="7" pageNumber="3048" pagination="703 - 719" refId="ref4307" refString="Grygier MJ, Ohtsuka S. 1995. SEM observation of the nauplius of Monstrilla hamatapex, new species, from Japan and an example of upgraded descriptive standards for monstrilloid copepods. J. Crustacean Biol. 15: 703 - 719. https: // doi. org / 10.1163 / 193724095 X 00118" type="journal article" year="1995">Grygier and Ohtsuka 1995</bibRefCitation>
; 
<bibRefCitation id="AB5E7AFFE87AFFCD75FAFAE9F482FAA9" author="Suarez-Morales E" box="[471,699,1322,1347]" pageId="7" pageNumber="3048" pagination="55 - 67" refId="ref4840" refString="Suarez-Morales E. 2010. On the taxonomic status of Monstrilla leucopis Sars (Crustacea: Copepoda: Monstrilloida) from Norway, with comments on the male of M. longiremis Giesbrecht. Zootaxa. 2510: 55 - 67. doi: 10.11646 / zootaxa. 2510.1.3" type="journal article" year="2010">Suárez-Morales 2010</bibRefCitation>
, 
<bibRefCitation id="AB5E7AFFE87AFFCD76E5FAE9F4C7FAA9" author="Suarez-Morales E" box="[712,766,1322,1347]" pageId="7" pageNumber="3048" pagination="22915" refId="ref4889" refString="Suarez-Morales E. 2011. Diversity of the Monstrilloida (Crustacea: Copepoda). PLoS One. 6 (8): e 22915. doi: 10.1371 / journal. pone. 0022915" type="journal article" year="2011">2011</bibRefCitation>
; 
<bibRefCitation id="AB5E7AFFE87AFFCD7726FAE9F583FAA9" author="Jeon D &amp; Lee W &amp; Soh HY" box="[779,954,1322,1347]" pageId="7" pageNumber="3048" pagination="45 - 65" refId="ref4445" refString="Jeon D, Lee W, Soh HY. 2018. A new genus and two new species of monstrilloid copepods (Copepoda: Monstrillidae): integrating morphological, molecular phylogenetic and ecological evidence. J Crustacean Biol. 38: 45 - 65. doi: 10.1093 / jcbiol / rux 095" type="journal article" year="2018">Jeon et al. 2018</bibRefCitation>
).
</paragraph>
<paragraph id="CF70070EE87AFFC274EDFA8EF72BFF07" blockId="7.[160,1156,524,1728]" lastBlockId="8.[160,1156,142,1658]" lastPageId="8" lastPageNumber="3049" pageId="7" pageNumber="3048">
<bibRefCitation id="AB5E7AFFE87AFFCD74EDFA8EF75FFA8C" author="McAlice BJ" box="[192,358,1357,1382]" pageId="7" pageNumber="3048" pagination="627 - 634" refId="ref4499" refString="McAlice BJ. 1985. On the male of Monstrilla helgolandica Claus (Copepoda, Monstrilloida). J Crustacean Biol. 5: 627 - 634. doi: 10.2307 / 1548240" type="journal article" year="1985">McAlice (1985)</bibRefCitation>
reidentified 
<specimenCount id="D9C9CC87E87AFFCD75D5FA8EF45CFA8C" box="[504,613,1357,1382]" pageId="7" pageNumber="3048" type="male">four male</specimenCount>
specimens of 
<taxonomicName id="08CF7C8DE87AFFCD7726FA8EF21DFA8C" authority="(Sars, 1921)" baseAuthorityName="Sars" baseAuthorityYear="1921" box="[779,1060,1357,1382]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="serricornis">
<emphasis id="FDBBDB1CE87AFFCD7726FA8EF5A2FA8C" box="[779,923,1357,1382]" italics="true" pageId="7" pageNumber="3048">M. serricornis</emphasis>
(Sars, 1921)
</taxonomicName>
(now in 
<taxonomicName id="08CF7C8DE87AFFCD748DFAACF445FA62" authority="Jeon, Soh and Lee, 2018" authorityName="Jeon, Soh and Lee" authorityYear="2018" box="[160,636,1391,1416]" class="Hexanauplia" family="Monstrillidae" genus="Caromiobenella" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="genus">
<emphasis id="FDBBDB1CE87AFFCD748DFAACF774FA62" box="[160,333,1391,1416]" italics="true" pageId="7" pageNumber="3048">Caromiobenella</emphasis>
<bibRefCitation id="AB5E7AFFE87AFFCD7574FAACF445FA62" author="Jeon D &amp; Lee W &amp; Soh HY" box="[345,636,1391,1416]" pageId="7" pageNumber="3048" pagination="45 - 65" refId="ref4445" refString="Jeon D, Lee W, Soh HY. 2018. A new genus and two new species of monstrilloid copepods (Copepoda: Monstrillidae): integrating morphological, molecular phylogenetic and ecological evidence. J Crustacean Biol. 38: 45 - 65. doi: 10.1093 / jcbiol / rux 095" type="journal article" year="2018">Jeon, Soh and Lee, 2018</bibRefCitation>
</taxonomicName>
, collected from the Buritari Lagoon, 
<collectingCountry id="B7D8479EE87AFFCD701AFAACF6D6FA41" name="Kiribati" pageId="7" pageNumber="3048">Gilbert Islands</collectingCountry>
, now 
<collectingCountry id="B7D8479EE87AFFCD751AFA51F7BEFA41" box="[311,391,1426,1451]" name="Kiribati" pageId="7" pageNumber="3048">Kiribati</collectingCountry>
) during the ‘Albatross’ cruise as 
<taxonomicName id="08CF7C8DE87AFFCD7729FA51F598FA41" authorityName="McMurrich" authorityYear="1917" box="[772,929,1426,1451]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="canadensis">
<emphasis id="FDBBDB1CE87AFFCD7729FA51F598FA41" box="[772,929,1426,1451]" italics="true" pageId="7" pageNumber="3048">M. canadensis</emphasis>
</taxonomicName>
(= 
<taxonomicName id="08CF7C8DE87AFFCD77E1FA51F24FFA41" baseAuthorityName="Claus" baseAuthorityYear="1863" box="[972,1142,1426,1451]" class="Hexanauplia" family="Monstrillidae" genus="Caromiobenella" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="helgolandica">
<emphasis id="FDBBDB1CE87AFFCD77E1FA51F24FFA41" box="[972,1142,1426,1451]" italics="true" pageId="7" pageNumber="3048">C. helgolandica</emphasis>
</taxonomicName>
). 
<bibRefCitation id="AB5E7AFFE87AFFCD748DFA76F75BFA24" author="Sekiguchi H" box="[160,354,1461,1486]" pageId="7" pageNumber="3048" pagination="24 - 34" refId="ref4532" refString="Sekiguchi H. 1982. Monstrilloid copepods from Ago bay, central Japan. Proc Japan Soc Syst Zool. 22: 24 - 34." type="journal article" year="1982">Sekiguchi (1982)</bibRefCitation>
recorded several males of 
<taxonomicName id="08CF7C8DE87AFFCD7682FA76F57CFA24" baseAuthorityName="Sars" baseAuthorityYear="1921" box="[687,837,1461,1486]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="serricornis">
<emphasis id="FDBBDB1CE87AFFCD7682FA76F57CFA24" box="[687,837,1461,1486]" italics="true" pageId="7" pageNumber="3048">M. serricornis</emphasis>
</taxonomicName>
in central 
<collectingCountry id="B7D8479EE87AFFCD77E3FA76F22DFA24" box="[974,1044,1461,1486]" name="Japan" pageId="7" pageNumber="3048">Japan</collectingCountry>
; this was probably the first confirmed record of the species in the area, since the male reported by 
<bibRefCitation id="AB5E7AFFE87AFFCD74E8FA39F766F9F9" author="Wilson CB" box="[197,351,1530,1555]" pageId="7" pageNumber="3048" pagination="141 - 441" refId="ref5387" refString="Wilson CB. 1950. Contributions to the biology of the Philippine Archipelago and adjacent regions. Copepods gathered by the United States fisheries steamer &quot; Albatross &quot; from 1887 to 1909, chiefly in the Pacific Ocean. U S Nat Mus Bull. 100: 141 - 441." type="journal article" year="1950">Wilson (1950)</bibRefCitation>
as 
<taxonomicName id="08CF7C8DE87AFFCD75A5FA39F421F9F9" baseAuthorityName="Sars" baseAuthorityYear="1921" box="[392,536,1530,1555]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="serricornis">
<emphasis id="FDBBDB1CE87AFFCD75A5FA39F421F9F9" box="[392,536,1530,1555]" italics="true" pageId="7" pageNumber="3048">M. serricornis</emphasis>
</taxonomicName>
in the 
<collectingCountry id="B7D8479EE87AFFCD7643FA39F4D0F9F9" box="[622,745,1530,1555]" name="Philippines" pageId="7" pageNumber="3048">Philippines</collectingCountry>
was probably a misidentification of either 
<taxonomicName id="08CF7C8DE87AFFCD74C3F9DEF7B7F9DC" authorityName="McMurrich" authorityYear="1917" box="[238,398,1565,1590]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="canadensis">
<emphasis id="FDBBDB1CE87AFFCD74C3F9DEF7B7F9DC" box="[238,398,1565,1590]" italics="true" pageId="7" pageNumber="3048">M. canadensis</emphasis>
</taxonomicName>
or the species described here (see Grygier 1995). Furthermore, 
<bibRefCitation id="AB5E7AFFE87AFFCD748DF9FCF77EF9B2" author="McAlice BJ" box="[160,327,1599,1624]" pageId="7" pageNumber="3048" pagination="627 - 634" refId="ref4499" refString="McAlice BJ. 1985. On the male of Monstrilla helgolandica Claus (Copepoda, Monstrilloida). J Crustacean Biol. 5: 627 - 634. doi: 10.2307 / 1548240" type="journal article" year="1985">McAlice (1985)</bibRefCitation>
stated that neither specimen from Caldera Bay, Mindanao, identified as 
<taxonomicName id="08CF7C8DE87AFFCD748DF9A1F700F991" baseAuthorityName="Sars" baseAuthorityYear="1921" box="[160,313,1634,1659]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="serricornis">
<emphasis id="FDBBDB1CE87AFFCD748DF9A1F700F991" box="[160,313,1634,1659]" italics="true" pageId="7" pageNumber="3048">M. serricornis</emphasis>
</taxonomicName>
by 
<bibRefCitation id="AB5E7AFFE87AFFCD7542F9A1F429F991" author="Wilson CB" box="[367,528,1634,1659]" pageId="7" pageNumber="3048" pagination="141 - 441" refId="ref5387" refString="Wilson CB. 1950. Contributions to the biology of the Philippine Archipelago and adjacent regions. Copepods gathered by the United States fisheries steamer &quot; Albatross &quot; from 1887 to 1909, chiefly in the Pacific Ocean. U S Nat Mus Bull. 100: 141 - 441." type="journal article" year="1950">Wilson (1950)</bibRefCitation>
bears the slightest resemblance to that species. In 
<taxonomicName id="08CF7C8DE87AFFCD748DF946F70BF974" baseAuthorityName="Sars" baseAuthorityYear="1921" box="[160,306,1669,1694]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="species" species="serricornis">
<emphasis id="FDBBDB1CE87AFFCD748DF946F70BF974" box="[160,306,1669,1694]" italics="true" pageId="7" pageNumber="3048">M. serricornis</emphasis>
</taxonomicName>
, as in all other species of 
<taxonomicName id="08CF7C8DE87AFFCD7641F946F52EF974" authorityName="Jeon, Soh and Lee" authorityYear="2018" box="[620,791,1669,1694]" class="Hexanauplia" family="Monstrillidae" genus="Caromiobenella" kingdom="Animalia" order="Monstrilloida" pageId="7" pageNumber="3048" phylum="Arthropoda" rank="genus">
<emphasis id="FDBBDB1CE87AFFCD7641F946F52EF974" box="[620,791,1669,1694]" italics="true" pageId="7" pageNumber="3048">Caromiobenella</emphasis>
</taxonomicName>
, the inner distal margin of the terminal antennular segment bears a set of distinctive, conspicuous serrate processes (
<bibRefCitation id="AB5E7AFFE875FFC2748BFF4DF763FF4D" author="Jeon D &amp; Lee W &amp; Soh HY" box="[166,346,142,167]" pageId="8" pageNumber="3049" pagination="45 - 65" refId="ref4445" refString="Jeon D, Lee W, Soh HY. 2018. A new genus and two new species of monstrilloid copepods (Copepoda: Monstrillidae): integrating morphological, molecular phylogenetic and ecological evidence. J Crustacean Biol. 38: 45 - 65. doi: 10.1093 / jcbiol / rux 095" type="journal article" year="2018">Jeon et al. 2018</bibRefCitation>
), absent in 
<taxonomicName id="08CF7C8DE875FFC275CDFF4DF455FF4D" authority="Suárez-Morales, 2022" authorityName="Suárez-Morales" authorityYear="2022" box="[480,620,142,167]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="albatrossi" status="sp. nov.">
<emphasis id="FDBBDB1CE875FFC275CDFF4DF455FF4D" box="[480,620,142,167]" italics="true" pageId="8" pageNumber="3049">M. albatrossi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E6886667E875FFC27659FF4DF4F7FF4D" box="[628,718,142,167]" pageId="8" pageNumber="3049" rank="species">sp. nov.</taxonomicNameLabel>
, but the general resemblance of these specimens in body shape and size may have led to Wilson’s misidentification of the specimen.
</paragraph>
<paragraph id="CF70070EE875FFC274EDFF35F55DFE35" blockId="8.[160,1156,142,1658]" pageId="8" pageNumber="3049">
The male specimen studied here can be included in the genus 
<taxonomicName id="08CF7C8DE875FFC277A0FF35F5C0FEE5" authorityName="Dana" authorityYear="1849" box="[909,1017,246,271]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="genus">
<emphasis id="FDBBDB1CE875FFC277A0FF35F5C0FEE5" box="[909,1017,246,271]" italics="true" pageId="8" pageNumber="3049">Monstrilla</emphasis>
</taxonomicName>
by virtue of the combined presence of two urosomites between the genital somite and the anal somite, the structure of its genital complex, the presence of well-developed or reduced fifth legs, geniculate antennules lacking discernible modifications, the possession of six caudal setae in both sexes, and five-branched antennulary setae sometimes present. This combination of characters does not occur in any other known genus of monstrilloids (
<bibRefCitation id="AB5E7AFFE875FFC2748BFE05F760FE35" author="Jeon D &amp; Lee W &amp; Soh HY" box="[166,345,454,479]" pageId="8" pageNumber="3049" pagination="45 - 65" refId="ref4445" refString="Jeon D, Lee W, Soh HY. 2018. A new genus and two new species of monstrilloid copepods (Copepoda: Monstrillidae): integrating morphological, molecular phylogenetic and ecological evidence. J Crustacean Biol. 38: 45 - 65. doi: 10.1093 / jcbiol / rux 095" type="journal article" year="2018">Jeon et al. 2018</bibRefCitation>
; 
<bibRefCitation id="AB5E7AFFE875FFC27548FE05F56EFE35" author="Suarez-Morales E &amp; Castellanos-Osorio I" box="[357,855,454,479]" pageId="8" pageNumber="3049" pagination="111 - 123" refId="ref5016" refString="Suarez-Morales E, Castellanos-Osorio I. 2019. A new species of Monstrilla (Copepoda, Monstrilloida, Monstrillidae) from the plankton of a bay system of the northwestern Caribbean, with a key to species. ZooKeys. 876: 111 - 123. doi: 10.3897 / zookeys. 876.38400" type="journal article" year="2019">Suárez-Morales and Castellanos-Osorio 2019</bibRefCitation>
).
</paragraph>
<paragraph id="CF70070EE875FFC274EDFE2AF56FFAA8" blockId="8.[160,1156,142,1658]" pageId="8" pageNumber="3049">
The new species can be easily recognised by its unique cephalothorax cuticular ornamentation. There are only a few known monstrillids with a complete or incomplete belt-like cephalothorax striation pattern; the clearest example is 
<taxonomicName id="08CF7C8DE875FFC277B8FDEDF2BAFDAD" authorityName="Isaac" authorityYear="1974" box="[917,1155,558,583]" class="Hexanauplia" family="Monstrillidae" genus="Cymbasoma" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="striatum">
<emphasis id="FDBBDB1CE875FFC277B8FDEDF2BAFDAD" box="[917,1155,558,583]" italics="true" pageId="8" pageNumber="3049">Cymbasoma striatum</emphasis>
</taxonomicName>
from south-eastern Britain, which shows a wide striation field covering half the length of its cephalothorax and running around the cephalothorax (see 
<bibRefCitation id="AB5E7AFFE875FFC277ADFDB7F247FD67" author="Suarez-Morales E" box="[896,1150,628,653]" pageId="8" pageNumber="3049" pagination="139 - 149" refId="ref4718" refString="Suarez-Morales E. 2000 b. Redescription of two species of Cymbasoma from Southwest Britain and from Indonesia (Copepoda: Monstrilloida) with notes on taxonomy. Beaufortia / Bull Zool Mus Amsterdam. 50: 139 - 149." type="journal article" year="2000">Suárez-Morales 2000b</bibRefCitation>
, figs 1, 6, 7). Also, the widespread nominal species 
<taxonomicName id="08CF7C8DE875FFC276D8FD55F6E2FD38" authority="Giesbrecht, 1893" authorityName="Giesbrecht" authorityYear="1893" class="Hexanauplia" family="Monstrillidae" genus="Cymbasoma" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="FDBBDB1CE875FFC276D8FD55F5C3FD45" box="[757,1018,662,687]" italics="true" pageId="8" pageNumber="3049">Cymbasoma reticulatus</emphasis>
Giesbrecht, 1893
</taxonomicName>
, the Caribbean 
<taxonomicName id="08CF7C8DE875FFC2759FFD7AF5D0FD38" authority="Suarez-Morales and Gasca, 1998" authorityName="Suarez-Morales and Gasca" authorityYear="1998" box="[434,1001,697,722]" class="Hexanauplia" family="Monstrillidae" genus="Cymbasoma" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="bowmani">
<emphasis id="FDBBDB1CE875FFC2759FFD7AF47AFD38" box="[434,579,697,722]" italics="true" pageId="8" pageNumber="3049">C. bowmani</emphasis>
<bibRefCitation id="AB5E7AFFE875FFC27678FD7AF5D0FD38" author="Suarez-Morales E &amp; Gasca R" box="[597,1001,697,722]" pageId="8" pageNumber="3049" pagination="433 - 439" refId="ref5110" refString="Suarez-Morales E, Gasca R. 1998. Cymbasoma bowmani n. sp., a new monstrilloid (Copepoda: Monstrilloida) from a Caribbean reef, with notes on species variation. J Mar Syst. 15 (1 - 4): 433 - 439. doi: 10.1016 / S 0924 - 7963 (97) 00092 - 4" type="journal article" year="1998">Suárez-Morales and Gasca, 1998</bibRefCitation>
</taxonomicName>
, the Pacific 
<taxonomicName id="08CF7C8DE875FFC2748DFD1FF573FD1F" authority="Suarez-Morales and Morales-Ramirez, 2003" authorityName="Suarez-Morales and Morales-Ramirez" authorityYear="2003" box="[160,842,732,757]" class="Hexanauplia" family="Monstrillidae" genus="Cymbasoma" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="concepcionae">
<emphasis id="FDBBDB1CE875FFC2748DFD1FF76CFD1F" box="[160,341,732,757]" italics="true" pageId="8" pageNumber="3049">C. concepcionae</emphasis>
<bibRefCitation id="AB5E7AFFE875FFC27573FD1FF573FD1F" author="Suarez-Morales E &amp; Morales-Ramirez A" box="[350,842,732,757]" pageId="8" pageNumber="3049" pagination="203 - 214" refId="ref5249" refString="Suarez-Morales E, Morales-Ramirez A. 2003. A new species of Cymbasoma (Crustacea: Copepoda: Monstrilloida) from the Pacific coast of Costa Rica, Central America. Proc Biol Soc Wash. 116 (1): 203 - 214." type="journal article" year="2003">Suárez-Morales and Morales-Ramírez, 2003</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="08CF7C8DE875FFC277A4FD1FF21FFD1F" authorityName="Suárez-Morales &amp; Morales-Ramírez" authorityYear="2009" box="[905,1062,732,757]" class="Hexanauplia" family="Monstrillidae" genus="Cymbasoma" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="guerrerense">
<emphasis id="FDBBDB1CE875FFC277A4FD1FF21FFD1F" box="[905,1062,732,757]" italics="true" pageId="8" pageNumber="3049">C. guerrerense</emphasis>
</taxonomicName>
SuárezMorales, 2009, and the Floridian 
<taxonomicName id="08CF7C8DE875FFC27608FD3DF5D0FCFD" authority="Suarez-Morales and Pilz, 2008" authorityName="Suarez-Morales and Pilz" authorityYear="2008" box="[549,1001,766,791]" class="Hexanauplia" family="Monstrillidae" genus="Cymbasoma" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="davisi">
<emphasis id="FDBBDB1CE875FFC27608FD3DF4BDFCFD" box="[549,644,766,791]" italics="true" pageId="8" pageNumber="3049">C. davisi</emphasis>
<bibRefCitation id="AB5E7AFFE875FFC276A2FD3DF5D0FCFD" author="Suarez-Morales E &amp; Pilz D" box="[655,1001,766,791]" pageId="8" pageNumber="3049" pagination="527 - 533" refId="ref5337" refString="Suarez-Morales E, Pilz D. 2008. A new species of Cymbasoma (Copepoda: Monstrilloida) from Florida with a redescription of C. quadridens. J Mar Biol Assoc U K. 88 (3): 527 - 533. doi: 10.1017 / S 0025315408000623" type="journal article" year="2008">Suárez-Morales and Pilz, 2008</bibRefCitation>
</taxonomicName>
show partial striation patterns covering either the lateral or ventral cephalothorax surfaces, but not completing a cuticular belt (
<bibRefCitation id="AB5E7AFFE875FFC2762CFC87F5BCFCB7" author="Suarez-Morales E &amp; Gasca R" box="[513,901,836,861]" pageId="8" pageNumber="3049" pagination="433 - 439" refId="ref5110" refString="Suarez-Morales E, Gasca R. 1998. Cymbasoma bowmani n. sp., a new monstrilloid (Copepoda: Monstrilloida) from a Caribbean reef, with notes on species variation. J Mar Syst. 15 (1 - 4): 433 - 439. doi: 10.1016 / S 0924 - 7963 (97) 00092 - 4" type="journal article" year="1998">Suárez-Morales and Gasca 1998</bibRefCitation>
; 
<bibRefCitation id="AB5E7AFFE875FFC277B4FC87F798FC95" author="Suarez-Morales E &amp; Morales-Ramirez A" pageId="8" pageNumber="3049" pagination="203 - 214" refId="ref5249" refString="Suarez-Morales E, Morales-Ramirez A. 2003. A new species of Cymbasoma (Crustacea: Copepoda: Monstrilloida) from the Pacific coast of Costa Rica, Central America. Proc Biol Soc Wash. 116 (1): 203 - 214." type="journal article" year="2003">Suárez-Morales and Morales-Ramírez 2003</bibRefCitation>
; 
<bibRefCitation id="AB5E7AFFE875FFC27599FCA5F528FC95" author="Suarez-Morales E &amp; Pilz D" box="[436,785,870,895]" pageId="8" pageNumber="3049" pagination="527 - 533" refId="ref5337" refString="Suarez-Morales E, Pilz D. 2008. A new species of Cymbasoma (Copepoda: Monstrilloida) from Florida with a redescription of C. quadridens. J Mar Biol Assoc U K. 88 (3): 527 - 533. doi: 10.1017 / S 0025315408000623" type="journal article" year="2008">Suárez-Morales and Pilz 2008</bibRefCitation>
; 
<bibRefCitation id="AB5E7AFFE875FFC27709FCA5F22CFC95" author="Suarez-Morales E &amp; Morales-Ramirez A" box="[804,1045,870,895]" pageId="8" pageNumber="3049" pagination="1257 - 1271" refId="ref5294" refString="Suarez-Morales E, Morales-Ramirez A. 2009. New species of Monstrilloida (Crustacea: Copepoda) from the Eastern Tropical Pacific. J Nat Hist. 43 (21 - 22): 1257 - 1271. doi: 10.1080 / 00222930902894419" type="journal article" year="2009">Suárez-Morales 2009</bibRefCitation>
). Among the known species of 
<taxonomicName id="08CF7C8DE875FFC2759DFC4AF427FC48" authorityName="Dana" authorityYear="1849" box="[432,542,905,930]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="genus">
<emphasis id="FDBBDB1CE875FFC2759DFC4AF427FC48" box="[432,542,905,930]" italics="true" pageId="8" pageNumber="3049">Monstrilla</emphasis>
</taxonomicName>
, comparable cephalothoracic striation patterns are present only in the Caribbean 
<taxonomicName id="08CF7C8DE875FFC2763DFC6FF547FC2F" authority="Suarez-Morales, 2003" authorityName="Suarez-Morales" authorityYear="2003" box="[528,894,940,965]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="marioi">
<emphasis id="FDBBDB1CE875FFC2763DFC6FF447FC2F" box="[528,638,940,965]" italics="true" pageId="8" pageNumber="3049">M. marioi</emphasis>
Suárez-Morales, 2003
</taxonomicName>
and 
<taxonomicName id="08CF7C8DE875FFC277ECFC6FF4A2FC0D" authority="Suarez-Morales and Castellanos-Osorio, 2019" authorityName="Suarez-Morales and Castellanos-Osorio" authorityYear="2019" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="chetumalensis">
<emphasis id="FDBBDB1CE875FFC277ECFC6FF2BAFC2F" box="[961,1155,940,965]" italics="true" pageId="8" pageNumber="3049">M. chetumalensis</emphasis>
<bibRefCitation id="AB5E7AFFE875FFC2748DFC0DF4A2FC0D" author="Suarez-Morales E &amp; Castellanos-Osorio I" box="[160,667,974,999]" pageId="8" pageNumber="3049" pagination="111 - 123" refId="ref5016" refString="Suarez-Morales E, Castellanos-Osorio I. 2019. A new species of Monstrilla (Copepoda, Monstrilloida, Monstrillidae) from the plankton of a bay system of the northwestern Caribbean, with a key to species. ZooKeys. 876: 111 - 123. doi: 10.3897 / zookeys. 876.38400" type="journal article" year="2019">Suárez-Morales and Castellanos-Osorio, 2019</bibRefCitation>
</taxonomicName>
; a well-developed cuticular ornamentation has been found in the male of 
<taxonomicName id="08CF7C8DE875FFC2760CFC32F6D9FBC7" authority="(Dias and Suarez-Morales 2000)" baseAuthorityName="Dias and Suarez-Morales" baseAuthorityYear="2000" class="Hexanauplia" family="Monstrillidae" genus="Caromiobenella" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="FDBBDB1CE875FFC2760CFC32F569FBE0" box="[545,848,1009,1034]" italics="true" pageId="8" pageNumber="3049">Caromiobenella brasiliensis</emphasis>
(
<bibRefCitation id="AB5E7AFFE875FFC2774FFC32F6EEFBC7" author="Dias C &amp; Suarez-Morales E" pageId="8" pageNumber="3049" pagination="1031 - 1039" refId="ref4110" refString="Dias C, and Suarez-Morales E. 2000. Two new species of Monstrilla (Copepoda: Monstrilloida) from Brazil. J Mar Biol Ass UK. 80: 1031 - 1039." type="journal article" year="2000">Dias and Suárez-Morales 2000</bibRefCitation>
)
</taxonomicName>
, with a wide band of transverse striae on both the lateral and ventral surfaces of the cephalothorax (see 
<bibRefCitation id="AB5E7AFFE875FFC27594FBF5F514FBA5" author="Cruz Lopes da Rosa J &amp; Dias CO &amp; Suarez-Morales E &amp; Weber LI &amp; Gomes Fischer L" box="[441,813,1078,1103]" pageId="8" pageNumber="3049" pagination="241" refId="ref4052" refString="Cruz Lopes da Rosa J, Dias CO, Suarez-Morales E, Weber LI, Gomes Fischer L. 2021. Record of Caromiobenella (Copepoda, Monstrilloida) in Brazil and discovery of the male of C. brasiliensis: morphological and molecular evidence. Diversity. 13: 241. doi: 10.3390 / d 13060241" type="journal article" year="2021">Cruz Lopes da Rosa et al. 2021</bibRefCitation>
. In 
<taxonomicName id="08CF7C8DE875FFC2774DFBF5F5F4FBA5" authorityName="Suarez-Morales" authorityYear="2003" box="[864,973,1078,1103]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="marioi">
<emphasis id="FDBBDB1CE875FFC2774DFBF5F5F4FBA5" box="[864,973,1078,1103]" italics="true" pageId="8" pageNumber="3049">M. marioi</emphasis>
</taxonomicName>
the striation is relatively sparse and limited to the perioral ventral surface (Suárez-Morales 2003). In 
<taxonomicName id="08CF7C8DE875FFC2748DFBBFF75FFB7F" authorityName="Suarez-Morales and Castellanos-Osorio" authorityYear="2019" box="[160,358,1148,1173]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="chetumalensis">
<emphasis id="FDBBDB1CE875FFC2748DFBBFF75FFB7F" box="[160,358,1148,1173]" italics="true" pageId="8" pageNumber="3049">M. chetumalensis</emphasis>
</taxonomicName>
the striation pattern is denser and concentrated on the anterior cephalic area (
<bibRefCitation id="AB5E7AFFE875FFC27562FB5DF567FB5D" author="Suarez-Morales E &amp; Castellanos-Osorio I" box="[335,862,1182,1207]" pageId="8" pageNumber="3049" pagination="111 - 123" refId="ref5016" refString="Suarez-Morales E, Castellanos-Osorio I. 2019. A new species of Monstrilla (Copepoda, Monstrilloida, Monstrillidae) from the plankton of a bay system of the northwestern Caribbean, with a key to species. ZooKeys. 876: 111 - 123. doi: 10.3897 / zookeys. 876.38400" type="journal article" year="2019">Suárez-Morales and Castellanos-Osorio 2019</bibRefCitation>
). In all these species of monstrilloids the striation comprises only simple transverse striae, thus diverging from the pattern shown by the new species 
<taxonomicName id="08CF7C8DE875FFC27657FB27F532FB17" authorityName="Suárez-Morales" authorityYear="2022" box="[634,779,1252,1277]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="albatrossi">
<emphasis id="FDBBDB1CE875FFC27657FB27F532FB17" box="[634,779,1252,1277]" italics="true" pageId="8" pageNumber="3049">M. albatrossi</emphasis>
</taxonomicName>
, with a more complex striation pattern including axial patches of papillae (
<figureCitation id="57F41B8BE875FFC276A2FAC5F4D4FAF5" box="[655,749,1286,1311]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="8" pageNumber="3049">Figure 2</figureCitation>
(a–c)). This is the first time this 
<typeStatus id="1074B9ACE875FFC2707CFAC5F2BAFAF5" box="[1105,1155,1286,1311]" pageId="8" pageNumber="3049">type</typeStatus>
of cuticular ornamentation is reported in male monstrilloids.
</paragraph>
<paragraph id="CF70070EE875FFC274EDFA8FF202F990" blockId="8.[160,1156,142,1658]" pageId="8" pageNumber="3049">
The structure of the genital complex is among the most important taxonomic characters for male monstrilloids (
<bibRefCitation id="AB5E7AFFE875FFC275DEFAADF4D2FA6D" author="Suarez-Morales E" box="[499,747,1390,1415]" pageId="8" pageNumber="3049" pagination="203 - 210" refId="ref4759" refString="Suarez-Morales E. 2000 c. The male of Cymbasoma quintanarooense (Suarez-Morales) (Copepoda, Monstrilloida) from the Caribbean with an identification key for the known males of Cymbasoma. Mar Biol Res. 85 (3): 203 - 210." type="journal article" year="2000">Suárez-Morales 2000c</bibRefCitation>
, 2003). Two 
<typeStatus id="1074B9ACE875FFC27753FAADF583FA6D" box="[894,954,1390,1415]" pageId="8" pageNumber="3049">types</typeStatus>
of male genitalia have been reported to occur in 
<taxonomicName id="08CF7C8DE875FFC2762CFA52F454FA40" authorityName="Dana" authorityYear="1849" box="[513,621,1425,1450]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="genus">
<emphasis id="FDBBDB1CE875FFC2762CFA52F454FA40" box="[513,621,1425,1450]" italics="true" pageId="8" pageNumber="3049">Monstrilla</emphasis>
</taxonomicName>
(
<bibRefCitation id="AB5E7AFFE875FFC27654FA52F529FA40" author="McAlice BJ" box="[633,784,1425,1450]" pageId="8" pageNumber="3049" pagination="627 - 634" refId="ref4499" refString="McAlice BJ. 1985. On the male of Monstrilla helgolandica Claus (Copepoda, Monstrilloida). J Crustacean Biol. 5: 627 - 634. doi: 10.2307 / 1548240" type="journal article" year="1985">McAlice 1985</bibRefCitation>
; Suárez-Morales 2000, 2003): one (
<typeStatus id="1074B9ACE875FFC2748BFA77F6E0FA27" box="[166,217,1460,1485]" pageId="8" pageNumber="3049">type</typeStatus>
I) having a relatively long, slender genital shaft with short, globular distal, nondivergent lappets, as in 
<taxonomicName id="08CF7C8DE875FFC27593FA15F595FA05" authority="Suarez-Morales and Dias, 2001" authorityName="Suarez-Morales and Dias" authorityYear="2001" box="[446,940,1494,1519]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="bahiana">
<emphasis id="FDBBDB1CE875FFC27593FA15F407FA05" box="[446,574,1494,1519]" italics="true" pageId="8" pageNumber="3049">M. bahiana</emphasis>
<bibRefCitation id="AB5E7AFFE875FFC27664FA15F595FA05" author="Suarez-Morales E &amp; Dias CO" box="[585,940,1494,1519]" pageId="8" pageNumber="3049" pagination="65 - 81" refId="ref5069" refString="Suarez-Morales E, Dias CO. 2001. Taxonomic report of some monstrilloids (Copepoda: Monstrilloida) from Brazil with description of four new species. Bull Inst R Sci Nat Belg Biol. 71: 65 - 81." type="journal article" year="2001">Suárez-Morales and Dias, 2001</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="08CF7C8DE875FFC27790FA15F21FFA05" authorityName="Suarez-Morales" authorityYear="1993" box="[957,1062,1494,1519]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="reidae">
<emphasis id="FDBBDB1CE875FFC27790FA15F21FFA05" box="[957,1062,1494,1519]" italics="true" pageId="8" pageNumber="3049">M. reidae</emphasis>
</taxonomicName>
SuárezMorales, 1993 and 
<taxonomicName id="08CF7C8DE875FFC2755AFA3AF4DFF9F8" authority="Suarez-Morales, 2003" authorityName="Suarez-Morales" authorityYear="2003" box="[375,742,1529,1554]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="globosa">
<emphasis id="FDBBDB1CE875FFC2755AFA3AF7D6F9F8" box="[375,495,1529,1554]" italics="true" pageId="8" pageNumber="3049">M. globosa</emphasis>
Suárez-Morales, 2003
</taxonomicName>
; and a second one (
<typeStatus id="1074B9ACE875FFC277E1FA3AF5C6F9F8" box="[972,1023,1529,1554]" pageId="8" pageNumber="3049">type</typeStatus>
II) showing a short, robust shaft with non-globose, separate distal lappets, as in the widespread 
<taxonomicName id="08CF7C8DE875FFC2748DF9FDF7C8F9BD" authority="Giesbrecht, 1891" authorityName="Giesbrecht" authorityYear="1891" box="[160,497,1598,1623]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="FDBBDB1CE875FFC2748DF9FDF725F9BD" box="[160,284,1598,1623]" italics="true" pageId="8" pageNumber="3049">M. grandis</emphasis>
Giesbrecht, 1891
</taxonomicName>
(see 
<bibRefCitation id="AB5E7AFFE875FFC27617F9FDF543F9BD" author="Suarez-Morales E &amp; Carrillo A &amp; Morales-Ramirez A" box="[570,890,1598,1623]" pageId="8" pageNumber="3049" pagination="619 - 638" refId="ref4953" refString="Suarez-Morales E, Carrillo A, Morales-Ramirez A. 2013. Report on some monstrilloids (Crustacea: Copepoda) from a reef area off the Caribbean coast of Costa Rica, Central America with description of two new species. J Nat Hist. 47 (5 - 12): 619 - 638. doi: 10.1080 / 00222933.2012. 742933" type="journal article" year="2013">Suárez-Morales et al. 2013</bibRefCitation>
), 
<taxonomicName id="08CF7C8DE875FFC277B8F9FDF218F9BD" authorityName="Suarez-Morales" authorityYear="1994" box="[917,1057,1598,1623]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="FDBBDB1CE875FFC277B8F9FDF218F9BD" box="[917,1057,1598,1623]" italics="true" pageId="8" pageNumber="3049">M. elongata</emphasis>
</taxonomicName>
SuárezMorales, 1994, 
<taxonomicName id="08CF7C8DE875FFC27564F9A2F4A6F990" authority="Giesbrecht, 1892" authorityName="Giesbrecht" authorityYear="1892" box="[329,671,1633,1658]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="longiremis">
<emphasis id="FDBBDB1CE875FFC27564F9A2F7E5F990" box="[329,476,1633,1658]" italics="true" pageId="8" pageNumber="3049">M. longiremis</emphasis>
Giesbrecht, 1892
</taxonomicName>
and 
<taxonomicName id="08CF7C8DE875FFC276F5F9A2F20FF990" authority="Suarez-Morales, 2003" authorityName="Suarez-Morales" authorityYear="2003" box="[728,1078,1633,1658]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="8" pageNumber="3049" phylum="Arthropoda" rank="species" species="marioi">
<emphasis id="FDBBDB1CE875FFC276F5F9A2F579F990" box="[728,832,1633,1658]" italics="true" pageId="8" pageNumber="3049">M. marioi</emphasis>
Suárez-Morales, 2003
</taxonomicName>
.
</paragraph>
<paragraph id="CF70070EE874FFC374EDFF4DF79EFCB7" blockId="9.[160,1156,142,861]" pageId="9" pageNumber="3050">
Overall, the new species 
<taxonomicName id="08CF7C8DE874FFC375CEFF4DF448FF4D" authorityName="Suárez-Morales" authorityYear="2022" box="[483,625,142,167]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="albatrossi">
<emphasis id="FDBBDB1CE874FFC375CEFF4DF448FF4D" box="[483,625,142,167]" italics="true" pageId="9" pageNumber="3050">M. albatrossi</emphasis>
</taxonomicName>
most closely resembles 
<taxonomicName id="08CF7C8DE874FFC377A2FF4DF276FF4D" authorityName="Suarez-Morales and Castellanos-Osorio" authorityYear="2019" box="[911,1103,142,167]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="chetumalensis">
<emphasis id="FDBBDB1CE874FFC377A2FF4DF276FF4D" box="[911,1103,142,167]" italics="true" pageId="9" pageNumber="3050">M. chetumalensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="08CF7C8DE874FFC3748DFF72F731FF20" authorityName="Suarez-Morales" authorityYear="2003" box="[160,264,177,202]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="marioi">
<emphasis id="FDBBDB1CE874FFC3748DFF72F731FF20" box="[160,264,177,202]" italics="true" pageId="9" pageNumber="3050">M. marioi</emphasis>
</taxonomicName>
; both congeners share with the new species several relevant characters: (1) the same 
<typeStatus id="1074B9ACE874FFC374CEFF17F72FFF07" box="[227,278,212,237]" pageId="9" pageNumber="3050">type</typeStatus>
of genital complex (
<typeStatus id="1074B9ACE874FFC375D0FF17F408FF07" box="[509,561,212,237]" pageId="9" pageNumber="3050">type</typeStatus>
II) and genital lappet structure, (2) a well-developed cephalothoracic cuticular ornamentation, (3) similar body proportions, (4) the presence of paired protuberances representing the fifth legs on the fifth pedigerous somite, and (5) a corrugate forehead. However, the new species shows a unique combination of characters, the most distinctive being: (1) cephalothorax with a belt-like striation pattern covering 1/3 of the post-oral cephalothoracic surface, (2) cuticular striation pattern including intercalate fields of small papillae, (3) 
<typeStatus id="1074B9ACE874FFC376E4FE67F4C5FE57" box="[713,764,420,445]" pageId="9" pageNumber="3050">type</typeStatus>
II genital complex with corrugate margin connecting the two lappets, (4) fifth antennulary segment with rounded proximal protuberance. The cephalic cuticular striation of both 
<taxonomicName id="08CF7C8DE874FFC377E5FE2AF27BFDE8" authorityName="Suarez-Morales" authorityYear="2003" box="[968,1090,489,514]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="marioi">
<emphasis id="FDBBDB1CE874FFC377E5FE2AF27BFDE8" box="[968,1090,489,514]" italics="true" pageId="9" pageNumber="3050">M. marioi</emphasis>
</taxonomicName>
and 
<taxonomicName id="08CF7C8DE874FFC3748CFDCFF758FDCF" authorityName="Suarez-Morales and Castellanos-Osorio" authorityYear="2019" box="[161,353,524,549]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="chetumalensis">
<emphasis id="FDBBDB1CE874FFC3748CFDCFF758FDCF" box="[161,353,524,549]" italics="true" pageId="9" pageNumber="3050">M. chetumalensis</emphasis>
</taxonomicName>
differs widely from the pattern observed in the new species. In both species, the cephalic striation is well developed but limited to the ventral perioral area and forehead. Also, in 
<taxonomicName id="08CF7C8DE874FFC37580FD92F449FD80" authorityName="Suarez-Morales and Castellanos-Osorio" authorityYear="2019" box="[429,624,593,618]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="chetumalensis">
<emphasis id="FDBBDB1CE874FFC37580FD92F449FD80" box="[429,624,593,618]" italics="true" pageId="9" pageNumber="3050">M. chetumalensis</emphasis>
</taxonomicName>
the fifth pedigerous somite has (1) a pair of posterolateral expansions and (2) a dorsal field of striae (
<bibRefCitation id="AB5E7AFFE874FFC377BCFDB7F78DFD45" author="Suarez-Morales E &amp; Castellanos-Osorio I" pageId="9" pageNumber="3050" pagination="111 - 123" refId="ref5016" refString="Suarez-Morales E, Castellanos-Osorio I. 2019. A new species of Monstrilla (Copepoda, Monstrilloida, Monstrillidae) from the plankton of a bay system of the northwestern Caribbean, with a key to species. ZooKeys. 876: 111 - 123. doi: 10.3897 / zookeys. 876.38400" type="journal article" year="2019">Suárez-Morales and Castellanos-Osorio 2019</bibRefCitation>
; 
<figureCitation id="57F41B8BE874FFC375ECFD55F426FD45" box="[449,543,662,687]" captionStart="Figure 2" captionStartId="4.[159,224,1339,1362]" captionTargetBox="[165,1117,215,1304]" captionTargetId="figure-85@4.[163,1153,142,1307]" captionTargetPageId="4" captionText="Figure 2. Monstrilla albatrossi sp. nov., holotype adult male from off Mindanao, Philippines, female holotype. (a) Anterior half of cephalothorax, ventral view; (b) same, right lateral view; (c) same, left lateral view. Abbreviations: hb, hyaline bodies (sensu Suárez-Morales 2018): if, intercalate field of striae; oc, oral cone; sb, striate cuticular belt. Scale bars: a–f = 100 μm; g = 50 μm" figureDoi="http://doi.org/10.5281/zenodo.6456866" httpUri="https://zenodo.org/record/6456866/files/figure.png" pageId="9" pageNumber="3050">Figure 2</figureCitation>
(d)). In this species the margin connecting the genital lappets is straight. Furthermore, both 
<taxonomicName id="08CF7C8DE874FFC3767BFD7AF4F9FD38" authorityName="Suarez-Morales" authorityYear="2003" box="[598,704,697,722]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="marioi">
<emphasis id="FDBBDB1CE874FFC3767BFD7AF4F9FD38" box="[598,704,697,722]" italics="true" pageId="9" pageNumber="3050">M. marioi</emphasis>
</taxonomicName>
and 
<taxonomicName id="08CF7C8DE874FFC376D6FD7AF582FD38" authorityName="Suarez-Morales and Castellanos-Osorio" authorityYear="2019" box="[763,955,697,722]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="chetumalensis">
<emphasis id="FDBBDB1CE874FFC376D6FD7AF582FD38" box="[763,955,697,722]" italics="true" pageId="9" pageNumber="3050">M. chetumalensis</emphasis>
</taxonomicName>
have four caudal setae (Suárez-Morales 2003; Suárez-Morales and Castellanos-Osorio 2018), thus diverging from the six caudal setae present in the new species 
<taxonomicName id="08CF7C8DE874FFC3771CFD3DF587FCFD" authorityName="Suárez-Morales" authorityYear="2022" box="[817,958,766,791]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="species" species="albatrossi">
<emphasis id="FDBBDB1CE874FFC3771CFD3DF587FCFD" box="[817,958,766,791]" italics="true" pageId="9" pageNumber="3050">M. albatrossi</emphasis>
</taxonomicName>
. Considering this finding, the Philippine diversity of 
<taxonomicName id="08CF7C8DE874FFC3761CFCE2F4A6FCD0" authorityName="Dana" authorityYear="1849" box="[561,671,801,826]" class="Hexanauplia" family="Monstrillidae" genus="Monstrilla" kingdom="Animalia" order="Monstrilloida" pageId="9" pageNumber="3050" phylum="Arthropoda" rank="genus">
<emphasis id="FDBBDB1CE874FFC3761CFCE2F4A6FCD0" box="[561,671,801,826]" italics="true" pageId="9" pageNumber="3050">Monstrilla</emphasis>
</taxonomicName>
now includes seven nominal species (see 
<bibRefCitation id="AB5E7AFFE874FFC3748DFC87F7A3FCB7" author="Suarez-Morales E" box="[160,410,836,861]" pageId="9" pageNumber="3050" pagination="680 - 686" refId="ref4674" refString="Suarez-Morales E. 2000 a. A new species and new geographic records of Monstrilla (Copepoda: Monstrilloida) from the Philippines. J Crustacean Biol. 20 (4): 680 - 686. doi: 10.1163 / 20021975 - 99990092" type="journal article" year="2000">Suárez-Morales 2000a</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>