102 lines
14 KiB
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102 lines
14 KiB
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<document id="44E3A00DB738EA79C5E6F10FB474D510" ID-DOI="10.11646/zootaxa.3925.2.8" ID-GBIF-Dataset="66953964-a8da-435c-9fdf-98e4c093482e" ID-ISSN="1175-5326" ID-Zenodo-Dep="236789" ID-ZooBank="EDA5F834-F90A-46E7-A987-448A02F77AC9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461002353221" checkinUser="plazi" docAuthor="Channing, Alan" docDate="2015" docId="3F11878FDC3DFFAB7AC3CFF1B7EBBCAA" docLanguage="en" docName="zt03925p280.pdf" docOrigin="Zootaxa 3925 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rana vertebralis Hewitt 1927" docType="treatment" docVersion="7" lastPageNumber="276" masterDocId="C328FFF7DC38FFAD7A54C94FB354BD5F" masterDocTitle="The Maluti Mystery revisited: Taxonomy of African River Frogs (Pyxicephalidae, Amietia) on the Drakensberg Mountains in southern Africa" masterLastPageNumber="280" masterPageNumber="271" pageNumber="276" updateTime="1698564978940" updateUser="plazi">
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<mods:titleInfo id="1967CFB7F1C645B50669C502498262CF">
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<mods:title id="C619C46162B58B40498661F65CE9A3FA">The Maluti Mystery revisited: Taxonomy of African River Frogs (Pyxicephalidae, Amietia) on the Drakensberg Mountains in southern Africa</mods:title>
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<mods:name id="56DBADB519B9A671274284B4EBABC487" type="personal">
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<mods:namePart id="3154F3DB61CA59D01CE381D33E78EBE1">Channing, Alan</mods:namePart>
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<mods:title id="711C8FC35233960E226FFA2464DA6D31">Zootaxa</mods:title>
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<mods:date id="B71F303EFBBCEF21F127795E60C8AB4E">2015</mods:date>
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<mods:number id="E8E8BCCDD99F4500899357655A0DE2C0">3925</mods:number>
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<treatment id="3F11878FDC3DFFAB7AC3CFF1B7EBBCAA" ID-DOI="http://doi.org/10.5281/zenodo.6102085" ID-GBIF-Taxon="119598735" ID-Zenodo-Dep="6102085" LSID="urn:lsid:plazi:treatment:3F11878FDC3DFFAB7AC3CFF1B7EBBCAA" httpUri="http://treatment.plazi.org/id/3F11878FDC3DFFAB7AC3CFF1B7EBBCAA" lastPageId="6" lastPageNumber="276" pageId="5" pageNumber="276">
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<subSubSection id="FFA26512DC3DFFA87AC3CFF1B180BB87" box="[151,724,1725,1752]" pageId="5" pageNumber="276" type="nomenclature">
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<paragraph id="B7073699DC3DFFA87AC3CFF1B180BB87" blockId="5.[151,724,1725,1752]" box="[151,724,1725,1752]" pageId="5" pageNumber="276">
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<heading id="EC4F81F5DC3DFFA87AC3CFF1B180BB87" bold="true" box="[151,724,1725,1752]" fontSize="11" level="1" pageId="5" pageNumber="276" reason="1">
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<emphasis id="85CCEA8BDC3DFFA87AC3CFF1B180BB87" bold="true" box="[151,724,1725,1752]" pageId="5" pageNumber="276">
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Identification of
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<taxonomicName id="70B84D1ADC3DFFA87B33CFF2B180BB87" authority="Hewitt, 1927" authorityName="Hewitt" authorityYear="1927" box="[359,724,1725,1752]" class="Amphibia" family="Ranidae" genus="Rana" kingdom="Animalia" order="Anura" pageId="5" pageNumber="276" phylum="Chordata" rank="species" species="vertebralis">
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<emphasis id="85CCEA8BDC3DFFA87B33CFF2B17BBB88" bold="true" box="[359,559,1725,1751]" italics="true" pageId="5" pageNumber="276">Rana vertebralis</emphasis>
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<bibRefCitation id="D3294B68DC3DFFA87862CFF1B180BB87" author="Hewitt" box="[566,724,1726,1752]" pageId="5" pageNumber="276" refString="Hewitt, J. (1927) Further descriptions of reptiles and batrachians from South Africa. Records of the Albany Museum, 3, 371 - 415, plates XX - XXIV." type="journal article" year="1927">Hewitt, 1927</bibRefCitation>
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</paragraph>
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<subSubSection id="FFA26512DC3DFFAB7AC3CE4BB7EBBCAA" lastPageId="6" lastPageNumber="277" pageId="5" pageNumber="276" type="description">
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<paragraph id="B7073699DC3DFFAB7AC3CE4BB7EBBCAA" blockId="5.[151,1437,1796,2038]" lastBlockId="6.[151,1437,151,501]" lastPageId="6" lastPageNumber="277" pageId="5" pageNumber="276">
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The two river frogs in question can be easily distinguished on the extensive webbing reaching the last phalanx of the fourth toe in the Maluti River Frog, but with up to three phalanges free in the Phofung River Frog, and small tympanum, less than the eye-tympanum distance in the Maluti River Frog, larger in the Phofung River Frog. The description (
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<bibRefCitation id="D3294B68DC3DFFA87B75CE3EB2FBBAD6" author="Hewitt" box="[289,431,1905,1930]" pageId="5" pageNumber="276" refString="Hewitt, J. (1927) Further descriptions of reptiles and batrachians from South Africa. Records of the Albany Museum, 3, 371 - 415, plates XX - XXIV." type="journal article" year="1927">Hewitt 1927</bibRefCitation>
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) is brief and incomplete, with an important error. Only four statements are diagnostic, and all four identify the Maluti River Frog—Tympanum width less than eye-tympanum distance; toes completely webbed in five
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<typeStatus id="6803883BDC3DFFA87B10CEF5B22BBA8D" box="[324,383,1978,2002]" pageId="5" pageNumber="276">types</typeStatus>
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(the description refers to half-webbed toes, but examination of the
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<typeStatus id="6803883BDC3DFFA87E24CEF6B786BA8D" box="[1136,1234,1977,2002]" pageId="5" pageNumber="276" type="holotype">holotype</typeStatus>
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<bibRefCitation id="D3294B68DC3DFFA87E8DCEF5B63DBA8D" author="PEM" box="[1241,1385,1977,2002]" pageId="5" pageNumber="276" refString="Holotype PEM A 1550 Mont-aux-Sources" type="book" year="1550">PEM A1550</bibRefCitation>
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, and the
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<typeStatus id="6803883BDC3DFFA87A96CE91B279BAA9" box="[194,301,2014,2038]" pageId="5" pageNumber="276" type="paratype">paratypes</typeStatus>
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PEM A1551, A1552, A1562 and A10562 shows this to be in error, only PEM A1555 has webbing that does not reach the tip of the fourth toe).
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<bibRefCitation id="D3294B68DC3EFFAB78FAC9D8B004BDEF" author="Hewitt" box="[686,848,151,176]" pageId="6" pageNumber="277" refString="Hewitt, J. (1927) Further descriptions of reptiles and batrachians from South Africa. Records of the Albany Museum, 3, 371 - 415, plates XX - XXIV." type="journal article" year="1927">Hewitt (1927)</bibRefCitation>
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illustrates the
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<typeStatus id="6803883BDC3EFFAB79ABC9D7B77BBDEF" box="[1023,1071,152,176]" pageId="6" pageNumber="277">type</typeStatus>
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in Plate 24. The quality of the reproduction is too poor to make out useful features. The left foot of the
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<typeStatus id="6803883BDC3EFFAB799CC9F2B0ACBD8A" box="[968,1016,189,213]" pageId="6" pageNumber="277">type</typeStatus>
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appears to show that three phalanges of the fourth toe are free of webbing (
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<figureCitation id="2F832A1CDC3EFFAB7839C990B1E1BDA7" box="[621,693,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,250,1300,1322]" captionTargetBox="[278,1309,545,1278]" captionTargetId="figure@6.[278,1309,545,1278]" captionTargetPageId="6" captionText="FIGURE 3. Type of Rana vertebralis Hewitt, 1927, PEM A 1550. Comparison of Plate 24, Figure 2 of Hewitt (1927) (A) with the specimen today (B) showing the extent of webbing." httpUri="https://zenodo.org/record/236792/files/figure.png" pageId="6" pageNumber="277">Fig. 3</figureCitation>
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A). This would identify the
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<typeStatus id="6803883BDC3EFFAB7E5FC9AFB76FBDA7" box="[1035,1083,224,248]" pageId="6" pageNumber="277">type</typeStatus>
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as a Phofung River Frog, in contradiction to the small tympanum. Unfortunately the
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<typeStatus id="6803883BDC3EFFAB7958C84AB068BC42" box="[780,828,261,285]" pageId="6" pageNumber="277">type</typeStatus>
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is now in very poor condition, but it clearly shows the fourth toe to be completely webbed. It appears that
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<figureCitation id="2F832A1CDC3EFFAB788BC868B075BC1F" box="[735,801,295,320]" captionStart="FIGURE 2" captionStartId="5.[151,250,1562,1584]" captionTargetBox="[153,1433,1021,1540]" captionTargetId="figure@5.[153,1433,1021,1541]" captionTargetPageId="5" captionText="FIGURE 2. (A) The type of Rana hymenopus Boulenger, 1920. NHMUK 1947.2. 28.65, (B) the type of Rana draconensis FitzSimons, 1948. NMSA 734 and (C) the type of Phrynobatrachus lawrencei FitzSimons, 1947. NMSA 669. All are Phofung River frogs." httpUri="https://zenodo.org/record/236791/files/figure.png" pageId="6" pageNumber="277">Fig. 2</figureCitation>
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of Plate 24 of
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<bibRefCitation id="D3294B68DC3EFFAB799EC868B73DBC1F" author="Hewitt" box="[970,1129,295,320]" pageId="6" pageNumber="277" refString="Hewitt, J. (1927) Further descriptions of reptiles and batrachians from South Africa. Records of the Albany Museum, 3, 371 - 415, plates XX - XXIV." type="journal article" year="1927">Hewitt (1927)</bibRefCitation>
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was retouched, as the dark line along the margin of the webbing is not present in the
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<typeStatus id="6803883BDC3EFFAB794FC802B01FBC3A" box="[795,843,333,357]" pageId="6" pageNumber="277">type</typeStatus>
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(
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<figureCitation id="2F832A1CDC3EFFAB790EC803B0C8BC3B" box="[858,924,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,250,1300,1322]" captionTargetBox="[278,1309,545,1278]" captionTargetId="figure@6.[278,1309,545,1278]" captionTargetPageId="6" captionText="FIGURE 3. Type of Rana vertebralis Hewitt, 1927, PEM A 1550. Comparison of Plate 24, Figure 2 of Hewitt (1927) (A) with the specimen today (B) showing the extent of webbing." httpUri="https://zenodo.org/record/236792/files/figure.png" pageId="6" pageNumber="277">Fig. 3</figureCitation>
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B). The webbing might have been invisible in the photograph if this retouching had not been done. Whoever did it, however, got it wrong. It also appears that Hewitt's photograph was flipped horizontally, as the right foot and right hand of the
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<typeStatus id="6803883BDC3EFFAB7E15C8DAB725BCF2" box="[1089,1137,405,429]" pageId="6" pageNumber="277">type</typeStatus>
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show the same orientation as the left foot and left arm in the photograph (
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<figureCitation id="2F832A1CDC3EFFAB78FEC8F8B1BBBC8F" box="[682,751,439,464]" captionStart="FIGURE 3" captionStartId="6.[151,250,1300,1322]" captionTargetBox="[278,1309,545,1278]" captionTargetId="figure@6.[278,1309,545,1278]" captionTargetPageId="6" captionText="FIGURE 3. Type of Rana vertebralis Hewitt, 1927, PEM A 1550. Comparison of Plate 24, Figure 2 of Hewitt (1927) (A) with the specimen today (B) showing the extent of webbing." httpUri="https://zenodo.org/record/236792/files/figure.png" pageId="6" pageNumber="277">Fig. 3</figureCitation>
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). The same specimen was illustrated by
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<bibRefCitation id="D3294B68DC3EFFAB7EEEC8F7B6CCBC8F" author="Tarrant" box="[1210,1432,440,464]" pageId="6" pageNumber="277" refString="Tarrant, J., Cunningham, M. J. & Du Preez, L. H. (2008) Maluti Mystery: A systematic review of Amietia vertebralis (Hewitt, 1927) and Strongylopus hymenopus (Boulenger, 1920) (Anura: Pyxicephalidae). Zootaxa, 1962, 33 - 48." type="journal article" year="2008">
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Tarrant
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<emphasis id="85CCEA8BDC3EFFAB7F46C8F6B610BC8F" box="[1298,1348,440,464]" italics="true" pageId="6" pageNumber="277">et al</emphasis>
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. (2008
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</bibRefCitation>
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:
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<figureCitation id="2F832A1CDC3EFFAB7AC3C893B38CBCAB" box="[151,216,476,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1562,1584]" captionTargetBox="[153,1433,1021,1540]" captionTargetId="figure@5.[153,1433,1021,1541]" captionTargetPageId="5" captionText="FIGURE 2. (A) The type of Rana hymenopus Boulenger, 1920. NHMUK 1947.2. 28.65, (B) the type of Rana draconensis FitzSimons, 1948. NMSA 734 and (C) the type of Phrynobatrachus lawrencei FitzSimons, 1947. NMSA 669. All are Phofung River frogs." httpUri="https://zenodo.org/record/236791/files/figure.png" pageId="6" pageNumber="277">Fig. 2</figureCitation>
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A) to show the poor condition. Note that the Tarrant
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<emphasis id="85CCEA8BDC3EFFAB7972C892B00ABCAB" box="[806,862,476,500]" italics="true" pageId="6" pageNumber="277">et al.</emphasis>
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<figureCitation id="2F832A1CDC3EFFAB7931C893B0F1BCAB" box="[869,933,476,501]" captionStart="FIGURE 2" captionStartId="5.[151,250,1562,1584]" captionTargetBox="[153,1433,1021,1540]" captionTargetId="figure@5.[153,1433,1021,1541]" captionTargetPageId="5" captionText="FIGURE 2. (A) The type of Rana hymenopus Boulenger, 1920. NHMUK 1947.2. 28.65, (B) the type of Rana draconensis FitzSimons, 1948. NMSA 734 and (C) the type of Phrynobatrachus lawrencei FitzSimons, 1947. NMSA 669. All are Phofung River frogs." httpUri="https://zenodo.org/record/236791/files/figure.png" pageId="6" pageNumber="277">Fig. 2</figureCitation>
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A is flipped horizontally.
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</paragraph>
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</treatment>
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