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Genus
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<emphasis bold="true" box="[397,521,1324,1351]" italics="true" pageId="3" pageNumber="2">Butheolus</emphasis>
<bibRefCitation author="Simon" box="[528,679,1325,1351]" firstAuthor="Simon" pageId="3" pageNumber="2" pagination="207 - 260" refId="ref35698" refString="SIMON, E. 1882. Etude sur les Arachnides de l'Yemen meridional. Viaggio ad Assab nel Mar Rosso, dei Signori G. Doria ed O. Beccari con il R. Avviso ' Esploratore' dal 16 nov. 1879 al 26 feb. 1880. Annali del Museo Civico di Storia Naturale di Genova, 18: 207 - 260." type="journal article" year="1882">Simon, 1882</bibRefCitation>
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<treatmentCitation author="SIMON" box="[192,499,1392,1415]" page="248" pageId="3" pageNumber="2" year="1882">
<taxonomicName ID-CoL="62HVH" authority="Simon, 1882: 248" authorityName="Simon" authorityPageNumber="248" authorityYear="1882" box="[192,499,1392,1415]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[192,299,1392,1415]" italics="true" pageId="3" pageNumber="2">Butheolus</emphasis>
<bibRefCitation author="Simon" box="[305,499,1392,1415]" firstAuthor="Simon" page="248" pageId="3" pageNumber="2" pagination="207 - 260" refId="ref35698" refString="SIMON, E. 1882. Etude sur les Arachnides de l'Yemen meridional. Viaggio ad Assab nel Mar Rosso, dei Signori G. Doria ed O. Beccari con il R. Avviso ' Esploratore' dal 16 nov. 1879 al 26 feb. 1880. Annali del Museo Civico di Storia Naturale di Genova, 18: 207 - 260." type="journal article" year="1882">Simon, 1882: 248</bibRefCitation>
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<treatmentCitation author="KARSCH" box="[192,518,1422,1445]" page="76" pageId="3" pageNumber="2" year="1886">
<taxonomicName ID-CoL="62HVH" authority=": Karsch, 1886: 76" authorityName=": Karsch" authorityPageNumber="76" authorityYear="1886" box="[192,518,1422,1445]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[192,299,1422,1445]" italics="true" pageId="3" pageNumber="2">Butheolus</emphasis>
:
<bibRefCitation author="Karsch" box="[319,518,1422,1445]" firstAuthor="Karsch" page="76" pageId="3" pageNumber="2" pagination="75 - 79" refId="ref33518" refString="KARSCH, F. 1886. Skorpionologische Beitrage (I. Uebereinen sizilianischen Skorpion. - II. Uebersicht der Gruppe Buthina (Androctonina). - III. Ueber einen neuen Opisthacanthus (Peters) Thor). Berliner Entomologische Zeitschrift, 30 (1): 75 - 79." type="journal article" year="1886">Karsch, 1886: 76</bibRefCitation>
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;
<treatmentCitation author="POCOCK" box="[536,801,1422,1445]" page="121" pageId="3" pageNumber="2" year="1890">
<bibRefCitation author="Pocock" box="[536,801,1422,1445]" firstAuthor="Pocock" page="121 - 122" pageId="3" pageNumber="2" pagination="114 - 141" refId="ref35337" refString="POCOCK, R. I. 1890. A revision of the genera of scorpions of the family Buthidae, with descriptions of some South-African species. Proceedings of the Zoological Society, 1890: 114 - 141." type="journal article" year="1890">Pocock, 1890: 121122</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="POCOCK" box="[316,511,1453,1476]" page="47" pageId="3" pageNumber="2" year="1892">
<bibRefCitation author="Pocock" box="[316,511,1453,1476]" firstAuthor="Pocock" page="47" pageId="3" pageNumber="2" pagination="38 - 49" refId="ref35376" refString="POCOCK, R. I. 1892. Descriptions of two new genera of scorpions, with notes on some species of Palamnaeus. Annals and Magazine of Natural History, (6), 9: 38 - 49." type="journal article" year="1892">Pocock, 1892: 47</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="LAURIE" box="[524,723,1453,1476]" page="131" pageId="3" pageNumber="2" year="1896">
<bibRefCitation author="Laurie" box="[524,723,1453,1476]" firstAuthor="Laurie" page="131" pageId="3" pageNumber="2" pagination="121 - 133" refId="ref34505" refString="LAURIE, M. 1896. Further notes on the anatomy and development of scorpions, and their bearing on the classification of the order. Annals and Magazine of Natural History (6), 18: 121 - 133." type="journal article" year="1896">Laurie, 1896: 131</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Lonnberg" page="" pageId="3" pageNumber="2" year="1897">
<bibRefCitation author="Lonnberg" firstAuthor="Lonnberg" pageId="3" pageNumber="2" pagination="193 - 211" refId="ref34578" refString="LONNBERG, E. 1897. Om Skorpionernas och Pedipalpernas geografiska Utbredning. Entomologisk Tidskrift, 18: 193 - 211." type="journal article" year="1897">Lönnberg, 1897</bibRefCitation>
b; 194
</treatmentCitation>
(part);
<treatmentCitation author="KRAEPELIN" box="[510,727,1484,1507]" page="34" pageId="3" pageNumber="2" year="1899">
<bibRefCitation author="Kraepelin" box="[510,727,1484,1507]" firstAuthor="Kraepelin" page="34" pageId="3" pageNumber="2" pagination="1 - 265" refId="ref34297" refString="KRAEPELIN, K. 1899. Scorpiones und Pedipalpi. In: Das Tierreich, R. Friedlander und Sohn Verl Berlin, 8: 1 - 265." type="journal article" year="1899">Kraepelin, 1899: 34</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="POCOCK" box="[240,429,1514,1537]" page="28" pageId="3" pageNumber="2" year="1900">
<bibRefCitation author="Pocock" box="[240,429,1514,1537]" firstAuthor="Pocock" page="28" pageId="3" pageNumber="2" refId="ref35479" refString="POCOCK, R. I. 1900. Arachnida. The Fauna of British India, including Ceylon and B urma. Published under the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp." type="book" year="1900">Pocock, 1900: 28</bibRefCitation>
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(part);
<treatmentCitation author="KRAEPELIN" box="[511,743,1514,1537]" page="337" pageId="3" pageNumber="2" year="1905">
<bibRefCitation author="Kraepelin" box="[511,743,1514,1537]" firstAuthor="Kraepelin" page="337" pageId="3" pageNumber="2" pagination="321 - 364" refId="ref34362" refString="KRAEPELIN, K. 1905. Die geographische Verbreitung der Skorpione. Zoologische Jahrbucher, Abtheilung fur Systematik, 22 (3): 321 - 364." type="journal article" year="1905">Kraepelin, 1905: 337</bibRefCitation>
</treatmentCitation>
, 353;
<treatmentCitation author="SIMON" box="[240,436,1545,1568]" page="77" pageId="3" pageNumber="2" year="1910">
<bibRefCitation author="Simon" box="[240,436,1545,1568]" firstAuthor="Simon" page="77" pageId="3" pageNumber="2" pagination="57 - 87" refId="ref35832" refString="SIMON, E. 1910. Revision des scorpions d'Egypte. Bulletin de la Societe Entomologique d'Egypte (Cairo), 1910: 57 - 87." type="journal article" year="1910">Simon, 1910: 77</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="HIRST" box="[455,648,1545,1568]" page="462" pageId="3" pageNumber="2" year="1911">
<bibRefCitation author="Hirst" box="[455,648,1545,1568]" firstAuthor="Hirst" page="462" pageId="3" pageNumber="2" pagination="462 - 473" refId="ref33448" refString="HIRST, S., 1911. Descriptions of new scorpions. Annals and Magazine of Natural History, ser. 8, 7: 462 - 473." type="journal article" year="1911">Hirst, 1911: 462</bibRefCitation>
</treatmentCitation>
, 464;
<treatmentCitation author="BIRULA" page="164" pageId="3" pageNumber="2" year="1917">
<bibRefCitation author="Birula" firstAuthor="Birula" page="164" pageId="3" pageNumber="2" refId="ref32799" refString="BIRULA, A. A. 1917 a. Arachnoidea Arthrogastra Caucasica. Pars I. Scorpiones. Zapiski Kavkazskogo Muzeya (Memoires du Musee du Caucase), Tiflis, Imprimerie de la Chancellerie du Comite pour la Transcaucasie. ser A, 5, 253 pp. (in Russian; published August 1917). English translation: Byalynitskii-Birulya, A. A. 1964. Arthrogastric Arachnids of Caucasia. 1. Scorpions. Israel Program for Scientific Translations, Jerusalem, 170 pp." type="book" year="1917" yearSuffix="a">Birula, 1917a: 164</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="BIRULA" box="[382,581,1576,1599]" page="55" pageId="3" pageNumber="2" year="1917">
<bibRefCitation author="Birula" box="[382,581,1576,1599]" firstAuthor="Birula" page="55" pageId="3" pageNumber="2" refId="ref32888" refString="BIRULA, A. A. 1917 b. Faune de la Russie et des pays Limitrophes Fondee Principalement sur les Collections du Musee Zoologique de l'Academie des Sciences de Russie. Arachnides (Arachnoidea), 1 (1): xx + 227 pp. (in Russian; preface dated October 1917). English translation: Byalynitskii-Birulya, A. A. 1965. Fauna of Russia and Adjacent Countries. Arachnoidea. Vol. I. Scorpions. Israel Program for Scientific Translations, Jerusalem, xix, 154 pp." type="book" year="1917" yearSuffix="b">Birula, 1917b: 55</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="WERNER" box="[597,798,1576,1599]" page="145" pageId="3" pageNumber="2" year="1934">
<bibRefCitation author="Werner" box="[597,798,1576,1599]" firstAuthor="Werner" page="145" pageId="3" pageNumber="2" pagination="8" refId="ref36674" refString="WERNER, F. 1934. Scorpiones, Pedipalpi. In: H. G. Bronns Klassen und Ordnungen des Tierreichs. Akademische Verlaggesellschaft, Leipzig. 5, IV, 8, Lief. 1 - 2, Scorpiones, pp. 1 - 316." type="journal article" year="1934">Werner, 1934:145</bibRefCitation>
</treatmentCitation>
, 266, 270, 294, 299;
<treatmentCitation author="KASTNER" box="[457,665,1606,1629]" page="231" pageId="3" pageNumber="2" year="1941">
<bibRefCitation author="Kastner" box="[457,665,1606,1629]" firstAuthor="Kastner" page="231" pageId="3" pageNumber="2" pagination="117 - 240" refId="ref33570" refString="KASTNER, A. 1941. 1. Ordnung der Arachnida: Scorpiones. In: Krumbach, T. (Ed.). Handbuch der Zoologie, Walter de Gruyter Verlag, Berlin, 3 (1): 117 - 240." type="journal article" year="1941">Kästner, 1941: 231</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="SREENIVASA-REDDY R. P." page="18" pageId="3" pageNumber="2" year="1970">
<bibRefCitation author="SREENIVASA-REDDY R. P." journalOrPublisher="Journal of Natural History" pageId="3" pageNumber="2" pagination="17 - 23" part="4" refId="ref36068" refString="SREENIVASA-REDDY R. P. 1970. The systematic position of the scorpion genus Charmus. Journal of Natural History, 4: 17 - 23." title="The systematic position of the scorpion genus Charmus" type="journal article" year="1970">Sreenivasa- Reddy, 1970</bibRefCitation>
: 18
</treatmentCitation>
;
<treatmentCitation author="STAHNKE" box="[441,657,1637,1660]" page="127" pageId="3" pageNumber="2" year="1972">
<bibRefCitation author="Stahnke" box="[441,657,1637,1660]" firstAuthor="Stahnke" page="127" pageId="3" pageNumber="2" pagination="121 - 133" refId="ref36117" refString="STAHNKE, H. L. 1972. A key to the genera of Buthidae (Scorpionida). Entomological News, 83 (5): 121 - 133." type="journal article" year="1972">Stahnke, 1972: 127</bibRefCitation>
</treatmentCitation>
, 129, fig. 12 (part);
<treatmentCitation author="VACHON" box="[322,547,1668,1691]" page="906" pageId="3" pageNumber="2" year="1974">
<bibRefCitation author="Vachon" box="[322,547,1668,1691]" firstAuthor="Vachon" page="906" pageId="3" pageNumber="2" pagination="857 - 958" refId="ref36318" refString="VACHON, M. 1974. Etude des caracteres utilises pour classe les familles et les genre de Scorpiones (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturalle Paris, Zoologie, (3) 104 (140): 857 - 958." type="journal article" year="1974">Vachon, 1974: 906</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="VACHON" box="[567,801,1668,1691]" page="1598" pageId="3" pageNumber="2" year="1975">
<bibRefCitation author="Vachon" box="[567,801,1668,1691]" firstAuthor="Vachon" page="1598" pageId="3" pageNumber="2" pagination="1597 - 1599" refId="ref36380" refString="VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalps des Scorpions (Arachnides) dans le classement des genres de famille des Buthidae Simon. Compte rendus hebdomadaires des seances de l'Academie des Sciences, Paris Ser. D Sciences Naturelles, 281 (21): 1597 - 1599." type="journal article" year="1975">Vachon, 1975: 1598</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="VACHON" box="[316,525,1698,1721]" page="253" pageId="3" pageNumber="2" year="1980">
<bibRefCitation author="Vachon" box="[316,525,1698,1721]" firstAuthor="Vachon" page="253" pageId="3" pageNumber="2" pagination="251 - 263" refId="ref36437" refString="VACHON, M. 1980. Scorpions du Dhofar. The Scientific Results of the Oman Flora and Fauna Survey 1977 (Dhofar). Journal of Oman Studies. Special Report, 2: 251 - 263." type="journal article" year="1980">Vachon, 1980: 253</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="FRANCKE" box="[610,797,1698,1721]" page="6" pageId="3" pageNumber="2" year="1985">
<bibRefCitation author="Francke" box="[610,797,1698,1721]" firstAuthor="Francke" page="6" pageId="3" pageNumber="2" pagination="1 - 32" refId="ref33339" refString="FRANCKE, O. F. 1985. Conspectus genericus scorpionorum 1758 - 1982 (Arachnida: Scorpiones). Occasional Papers of the Museum, Texas Tech University, 98: 1 - 32." type="journal article" year="1985">Francke, 1985: 6</bibRefCitation>
</treatmentCitation>
, 15;
<treatmentCitation author="VACHON, M. &amp; R. KINZELBACH" box="[282,635,1729,1752]" page="92" pageId="3" pageNumber="2" year="1987">
<bibRefCitation author="VACHON, M. &amp; R. KINZELBACH" box="[282,635,1729,1752]" firstAuthor="Vachon" journalOrPublisher="Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East, Mainz, 1985. Beihefte zum Tubinger Atlas des Vorderen Orients, Reihe A (Naturwissenschaften)" page="92" pageId="3" pageNumber="2" pagination="91 - 103" part="28" refId="ref36476" refString="VACHON, M. &amp; R. KINZELBACH. 1987. On the taxonomy and distribution of the scorpions of the Middle East. i n Krupp, F., W. Schneider &amp; R. Kinzelbach (eds.). Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East, Mainz, 1985. Beihefte zum Tubinger Atlas des Vorderen Orients, Reihe A (Naturwissenschaften), 28: 91 - 103." title="On the taxonomy and distribution of the scorpions of the Middle East. i n Krupp, F., W. Schneider &amp; R. Kinzelbach (eds.)" type="journal article" year="1987">Vachon &amp; Kinzelbach, 1987: 92</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="SISSOM, W. D." page="97" pageId="3" pageNumber="2" year="1990">
<bibRefCitation author="Sissom" editor="Polis, G. A." firstAuthor="Sissom" journalOrPublisher="Stanford University Press, Stanford, California" page="97" pageId="3" pageNumber="2" pagination="64 - 160" refId="ref35859" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160 in Polis, G. A. (ed.). The Biology of Scorpions. Stanford University Press, Stanford, California." title="Systematics, biogeography and paleontology" type="book chapter" volumeTitle="The Biology of Scorpions" year="1990">Sissom, 1990: 97</bibRefCitation>
</treatmentCitation>
, 101 (part);
<treatmentCitation author="EL-HENNAWY, H. K." page="97" pageId="3" pageNumber="2" year="1992">
<bibRefCitation author="EL-HENNAWY, H. K." firstAuthor="El-Hennawy" journalOrPublisher="Serket" pageId="3" pageNumber="2" pagination="95 - 153" part="2" refId="ref33103" refString="EL-HENNAWY, H. K. 1992. A catalogue of the scorpions described from the Arab Countries (1758 - 1990) (Arachnida: Scorpionida). Serket, 2 (4): 95 - 153." title="A catalogue of the scorpions described from the Arab Countries (1758 - 1990) (Arachnida: Scorpionida)" type="journal article" year="1992">El-Hennawy, 1992: 97, 101, 104, 114</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="NENILIN, A. B. &amp; V. FET" box="[419,685,1790,1813]" page="17" pageId="3" pageNumber="2" year="1992">
<bibRefCitation author="NENILIN, A. B. &amp; V. FET" box="[419,685,1790,1813]" firstAuthor="Nenilin" journalOrPublisher="Arthropoda Selecta" page="17" pageId="3" pageNumber="2" pagination="3 - 31" part="1" refId="ref35238" refString="NENILIN, A. B. &amp; V. FET. 1992. Zoogeographical analysis of the world scorpion fauna (Arachnida: Scorpiones). Arthropoda Selecta, 1 (2): 3 - 31 (in Russian)." title="Zoogeographical analysis of the world scorpion fauna (Arachnida: Scorpiones)" type="journal article" year="1992">Nenilin &amp; Fet, 1992: 17</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="LOURENCO, W. R. &amp; M. VACHON" page="297" pageId="3" pageNumber="2" year="1995">
<bibRefCitation author="Lourenco &amp; Vachon" firstAuthor="Lourenco" journalOrPublisher="Bulletin du Museum National d'Histoire Naturelle de Paris" page="297" pageId="3" pageNumber="2" pagination="297 - 305" part="17" refId="ref34968" refString="LOURENCO, W. R. &amp; M. VACHON. 1995. Un nouveau genre et deux nouvelles especes de scorpions Buthidae d'Iran. Bulletin du Museum National d'Histoire Naturelle de Paris, 17 (3 - 4): 297 - 305." title="Un nouveau genre et deux nouvelles especes de scorpions Buthidae d'Iran" type="journal article" year="1995">Lourenço &amp; Vachon, 1995: 297</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="KOVARIK" box="[517,798,1821,1844]" page="177" pageId="3" pageNumber="2" year="1996">
<bibRefCitation author="Kovarik" box="[517,798,1821,1844]" firstAuthor="Kovarik" page="177 - 178" pageId="3" pageNumber="2" pagination="177 - 181" refId="ref33615" refString="KOVARIK, F. 1996. Baloorthochirus becvari gen. et sp. n. from Pakistan, and taxonomic position of Orthochirus luteipes (Scorpiones: Buthidae). Acta Societatis Zoologicae Bohemicae, 60: 177 - 181." type="journal article" year="1996">Kovařík, 1996: 177178</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="KOVARIK" box="[240,474,1852,1875]" page="117" pageId="3" pageNumber="2" year="1998">
<bibRefCitation author="Kovarik" box="[240,474,1852,1875]" firstAuthor="Kovarik" page="117" pageId="3" pageNumber="2" pagination="115 - 124" refId="ref33656" refString="KOVARIK, F. 1998 a. Three new genera and species of Scorpiones (Buthidae) from Somalia. Acta Societatis Zoologicae Bohemicae, 62: 115 - 124." type="journal article" year="1998" yearSuffix="a">Kovařík, 1998a: 117</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="KOVARIK" box="[491,723,1852,1875]" page="105" pageId="3" pageNumber="2" year="1998">
<bibRefCitation author="Kovarik" box="[491,723,1852,1875]" firstAuthor="Kovarik" page="105" pageId="3" pageNumber="2" refId="ref33687" refString="KOVARIK, F. 1998 b. Stiri. Madagaskar, Jihlava, 175 pp." type="book" year="1998" yearSuffix="b">Kovařík, 1998b: 105</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="FET" box="[240,484,1882,1905]" page="88" pageId="3" pageNumber="2" year="2000">
<bibRefCitation author="Fet &amp; Lowe" box="[240,484,1882,1905]" firstAuthor="Fet" page="88" pageId="3" pageNumber="2" refId="ref33203" refString="FET, V. &amp; G. LOWE. 2000. Family Buthidae. Pp. 54 - 286 in FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. Catalog of the Scorpions of the World (1758 - 1998). The New York Entomological Society." type="book" year="2000">Fet &amp; Lowe, 2000: 88</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="LOURENCO" box="[567,797,1882,1905]" page="179" pageId="3" pageNumber="2" year="2001">
<bibRefCitation author="Lourenco" box="[567,797,1882,1905]" firstAuthor="Lourenco" page="179" pageId="3" pageNumber="2" pagination="171 - 183" refId="ref34688" refString="LOURENCO, W. R. 2001. Taxonomic considerations on the genera Butheolus Simon, Nanobuthus Pocock and Neobuthus Hirst (Scorpions, Buthidae) with the description of a new species of Neobuthus from Ethiopia. Ecology of Desert Environments, 2001: 171 - 183." type="journal article" year="2001">Lourenço, 2001: 179</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="KOVARIK" box="[878,1157,199,222]" page="137" pageId="3" pageNumber="2" year="2003">
<bibRefCitation author="Kovarik" box="[878,1157,199,222]" firstAuthor="Kovarik" page="137 - 138" pageId="3" pageNumber="2" pagination="133 - 159" refId="ref33703" refString="KOVARIK, F. 2003. Scorpions of Djibouti, Eritrea, Ethiopia, and Somalia (Arachnida: Scorpiones), with a key and descriptions of three new species. Acta Societatis Zoologicae Bohemicae, 67: 133 - 159." type="journal article" year="2003">Kovařík, 2003: 137138</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="SOLEGLAD, M. E. &amp; V. FET" page="88" pageId="3" pageNumber="2" year="2003">
<bibRefCitation author="SOLEGLAD, M. E. &amp; V. FET" firstAuthor="Soleglad" journalOrPublisher="Euscorpius" page="88" pageId="3" pageNumber="2" pagination="1 - 34" part="5" refId="ref35997" refString="SOLEGLAD, M. E. &amp; V. FET. 2003 a. The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni). Euscorpius, 5: 1 - 34." title="The scorpion sternum: structure and phylogeny (Scorpiones: Orthosterni)" type="journal article" year="2003">Soleglad &amp; Fet, 2003: 88</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="KOVARIK" box="[1003,1229,230,253]" httpUri="http://treatment.plazi.org/id/038B20403C335B2EFC6EF8CBFDCEF9E5" page="2" pageId="3" pageNumber="2" year="2004">
<bibRefCitation author="Kovarik" box="[1003,1229,230,253]" firstAuthor="Kovarik" page="2 - 3" pageId="3" pageNumber="2" pagination="1 - 33" refId="ref33746" refString="KOVARIK, F. 2004. Revision and taxonomic position of genera Afghanorthochirus Lourenco &amp; Vachon, Baloorthochirus Kovarik, Butheolus Simon, Nanobuthus Pocock, Orthochiroides Kovarik, Pakistanorthochirus Lourenco, and Asian Orthochirus Karsch, with descriptions of twelve new species (Scorpiones, Buthidae). Euscorpius, 16: 1 - 33." type="journal article" year="2004">Kovařík, 2004: 23</bibRefCitation>
</treatmentCitation>
, 25 (part);
<treatmentCitation author="FET, V. &amp; M. E. SOLEGLAD" page="11" pageId="3" pageNumber="2" year="2005">
<bibRefCitation author="FET, V. &amp; M. E. SOLEGLAD" journalOrPublisher="Euscorpius" pageId="3" pageNumber="2" pagination="1 - 13" part="31" refId="ref33263" refString="FET, V. &amp; M. E. SOLEGLAD. 2005. Contributions to Scorpion Systematics. I. On recent changes in highlevel taxonomy. Euscorpius, 31: 1 - 13." title="Contributions to Scorpion Systematics. I. On recent changes in highlevel taxonomy." type="journal article" year="2005">Fet &amp; Soleglad, 2005: 11</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="FET, V. &amp; M. E. SOLEGLAD &amp; G. LOWE" page="3" pageId="3" pageNumber="2" year="2005">
<bibRefCitation author="FET, V. &amp; M. E. SOLEGLAD &amp; G. LOWE" box="[1097,1435,260,283]" firstAuthor="Fet" journalOrPublisher="Euscorpius" page="3" pageId="3" pageNumber="2" pagination="1 - 40" part="23" refId="ref33298" refString="FET, V., M. E. SOLEGLAD &amp; G. LOWE. 2005. A new trichobothrial character for the high-level systematics of Buthoidea (Scorpiones: Buthida). Euscorpius, 23: 1 - 40." title="A new trichobothrial character for the high-level systematics of Buthoidea (Scorpiones: Buthida)" type="journal article" year="2005">Fet, Soleglad &amp; Lowe, 2005: 3</bibRefCitation>
, 11, 22
</treatmentCitation>
, fig. 23 (part);
<treatmentCitation author="LOURENCO" box="[1162,1436,291,314]" page="26" pageId="3" pageNumber="2" year="2005">
<bibRefCitation author="Lourenco" box="[1162,1436,291,314]" firstAuthor="Lourenco" page="26 - 28" pageId="3" pageNumber="2" pagination="21 - 28" refId="ref34737" refString="LOURENCO, W. R. 2005. Description of three new species of scorpion from Sudan (Scorpiones, Buthidae). Boletin Sociedad Entomologica Aragonesa, 3 6: 21 - 28." type="journal article" year="2005">Lourenço, 2005: 2628</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="PRENDINI, L. &amp; W. WHEELER" box="[878,1226,322,345]" page="481" pageId="3" pageNumber="2" year="2005">
<bibRefCitation author="PRENDINI, L. &amp; W. WHEELER" box="[878,1226,322,345]" firstAuthor="Prendini" journalOrPublisher="Cladistics" page="481" pageId="3" pageNumber="2" pagination="446 - 494" part="21" refId="ref35565" refString="PRENDINI, L. &amp; W. WHEELER. 2005. Scorpion high- er phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing. Cladistics, 21: 446 - 494." title="Scorpion high- er phylogeny and classification, taxonomic anarchy, and standards for peer review in online publishing" type="journal article" year="2005">Prendini &amp; Wheeler, 2005: 481</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="HENDRIXSON" page="56" pageId="3" pageNumber="2" year="2006">
<bibRefCitation author="Hendrixson" firstAuthor="Hendrixson" page="56" pageId="3" pageNumber="2" pagination="33 - 120" refId="ref33409" refString="HENDRIXSON, B. E. 2006. Buthid scorpions of Saudi Arabia, with notes on other families (Scorpiones: Buthidae, Liochelidae, Scorpionidae). Fauna of Arabia, 21: 33 - 120." type="journal article" year="2006">Hendrixson, 2006: 56</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="LOURENCO, W. R. &amp; J. - X. QI" box="[1005,1356,352,375]" page="91" pageId="3" pageNumber="2" year="2006">
<bibRefCitation author="LOURENCO, W. R. &amp; J. - X. QI" box="[1005,1356,352,375]" firstAuthor="Lourenco" journalOrPublisher="Zoology in the Middle East" page="91 - 93" pageId="3" pageNumber="2" pagination="91 - 97" part="37" refId="ref34866" refString="LOURENCO, W. R. &amp; J. - X. QI. 2006. Further considerations on the genus Butheolus Simon, 1882 and description of one new species from Saudi Arabia (Scorpiones: Buthidae). Zoology in the Middle East, 37: 91 - 97." title="Further considerations on the genus Butheolus Simon, 1882 and description of one new species from Saudi Arabia (Scorpiones: Buthidae)" type="journal article" year="2006">Lourenço &amp; Qi, 2006: 9193</bibRefCitation>
</treatmentCitation>
(part);
<treatmentCitation author="DUPRE" box="[878,1052,383,406]" page="4" pageId="3" pageNumber="2" year="2007">
<bibRefCitation author="Dupre" box="[878,1052,383,406]" firstAuthor="Dupre" page="4" pageId="3" pageNumber="2" pagination="1 - 31" refId="ref33077" refString="DUPRE, G. 2007. Conspectus genericus scorpionorum 1758 - 2006 (Arachnida: Scorpiones). Euscorpius, 50: 1 - 31." type="journal article" year="2007">Dupré, 2007: 4</bibRefCitation>
</treatmentCitation>
, 12, 17 (part);
<treatmentCitation author="KOVARIK" box="[1229,1436,383,406]" page="23" pageId="3" pageNumber="2" year="2009">
<bibRefCitation author="Kovarik" box="[1229,1436,383,406]" firstAuthor="Kovarik" page="23" pageId="3" pageNumber="2" refId="ref33839" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Jakub Rolcik - Clairon Production, Prague, 170 pp." type="book" year="2009">Kovařík, 2009: 23</bibRefCitation>
</treatmentCitation>
, 31;
<treatmentCitation author="EL-HENNAWY, H. K." box="[924,1156,414,437]" page="126" pageId="3" pageNumber="2" year="2009">
<bibRefCitation author="EL-HENNAWY, H. K." box="[924,1105,414,437]" journalOrPublisher="Serket" pageId="3" pageNumber="2" pagination="119 - 128" part="11" refId="ref33143" refString="EL-HENNAWY, H. K. 2009. Scorpions of Saudi Arabia. Serket 11 (3 / 4): 119 - 128." title="Scorpions of Saudi Arabia" type="journal article" year="2009">Hennawy, 2009:</bibRefCitation>
126
</treatmentCitation>
(part);
<treatmentCitation author="LOWE" box="[1245,1436,414,437]" page="37" pageId="3" pageNumber="2" year="2010">
<bibRefCitation author="Lowe" box="[1245,1436,414,437]" firstAuthor="Lowe" page="37" pageId="3" pageNumber="2" pagination="1 - 53" refId="ref35042" refString="LOWE, G. 2010 a. New picobuthoid scorpions (Scorpiones: Buthidae) from Oman. Euscorpius, 93: 1 - 53." type="journal article" year="2010" yearSuffix="a">Lowe, 2010a: 37</bibRefCitation>
</treatmentCitation>
, 44;
<treatmentCitation author="LOURENCO, W. R. &amp; E. - A. LEGUIN" box="[921,1233,444,467]" page="1" pageId="3" pageNumber="2" year="2011">
<bibRefCitation author="LOURENCO, W. R. &amp; E. - A. LEGUIN" box="[921,1233,444,467]" firstAuthor="Lourenco" journalOrPublisher="Euscorpius" page="1" pageId="3" pageNumber="2" pagination="1 - 19" part="123" refId="ref34815" refString="LOURENCO, W. R. &amp; E. - A. LEGUIN. 2011. Further considerations on the species of the genus Orthochirus Karsch, 1891 from Africa, with description of three new species (Scorpiones: Buthidae). Euscorpius, 123: 1 - 19." title="Further considerations on the species of the genus Orthochirus Karsch, 1891 from Africa, with description of three new species (Scorpiones: Buthidae)" type="journal article" year="2011">Lourenço &amp; Leguin, 2011: 1</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="KOVARIK, F. &amp; G. LOWE" page="1-2" pageId="3" pageNumber="2" year="2012">
<bibRefCitation author="KOVARIK, F. &amp; G. LOWE" firstAuthor="Kovarik" journalOrPublisher="Euscorpius" page="1 - 2" pageId="3" pageNumber="2" pagination="1 - 25" part="139" refId="ref33864" refString="KOVARIK, F. &amp; G. LOWE. 2012. Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae). Euscorpius, 139: 1 - 25." title="Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae)" type="journal article" year="2012">Kovařík &amp; Lowe, 2012: 12</bibRefCitation>
, 1823
</treatmentCitation>
(part);
<treatmentCitation author="KOVARIK" page="4" pageId="3" pageNumber="2" year="2013">
<bibRefCitation author="Kovarik" etAl="et al." firstAuthor="Kovarik" page="4" pageId="3" pageNumber="2" pagination="1 - 19" refId="ref33987" refString="KOVARIK, F., G. LOWE, J. PLISKOVA &amp; F. STAHLAVSKY. 2013. A new scorpion genus, Gint gen. n., from the Horn of Africa (Scorpiones, Buthidae). Euscorpius, 173: 1 - 19." type="journal article" year="2013">Kovařík et al., 2013: 4</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="YANG &amp; LOURENCO &amp; ZHU" box="[907,1125,506,529]" page="5" pageId="3" pageNumber="2" year="2013">
<bibRefCitation author="Yang" box="[907,1125,506,529]" etAl="et al." firstAuthor="Yang" page="5" pageId="3" pageNumber="2" pagination="55125" refId="ref36772" refString="YANG, X., Y. NORMA-RASHID, W. R. LOURENCO &amp; M. ZHU. 2013. True lateral eye numbers for extant buthids: a new discovery on an old character. PLoS ONE, 8 (1): e 55125. doi: 10.1371 / journal. pone. 0055125." type="journal article" year="2013">Yang et al., 2013: 5</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="LORIA &amp; PRENDINI" box="[1138,1436,506,529]" page="25" pageId="3" pageNumber="2" year="2014">
<bibRefCitation author="Loria &amp; Prendini" box="[1138,1436,506,529]" firstAuthor="Loria" page="25" pageId="3" pageNumber="2" pagination="112913" refId="ref34600" refString="LORIA, S. F. &amp; L. PRENDINI. 2014. Homology of the lateral eyes of Scorpiones: a six-ocellus model. PLoS ONE 9 (12): e 112913. doi: 10.1371 / journal. pone. 0112913." type="journal article" year="2014">Loria &amp; Prendini, 2014: 25</bibRefCitation>
</treatmentCitation>
, tab. 5;
<treatmentCitation author="LOWE" box="[972,1254,536,559]" page="120" pageId="3" pageNumber="2" year="2014">
<bibRefCitation author="Lowe" box="[972,1254,536,559]" etAl="et al." firstAuthor="Lowe" page="120" pageId="3" pageNumber="2" pagination="1 - 129" refId="ref35151" refString="LOWE, G., E. A. YAGMUR &amp; F. KOVARIK. 2014. A review of the genus Leiurus Ehrenberg, 1828 (Scorpiones: Buthidae) with description of four new species from the Arabian Peninsula. Euscorpius, 191: 1 - 129." type="journal article" year="2014">Lowe et al., 2014: 120</bibRefCitation>
</treatmentCitation>
, fig. 98;
<treatmentCitation author="LOWE" page="1" pageId="3" pageNumber="2" year="2016">
<bibRefCitation author="Lowe &amp; Kovarik" firstAuthor="Lowe" page="1" pageId="3" pageNumber="2" pagination="1 - 25" refId="ref35100" refString="LOWE, G. &amp; F. KOVARIK. 2016. Scorpions of the Horn of Africa (Arachnida, Scorpiones). Part V. Two new Species of Neobuthus Hirst, 1911 (Buthidae), from Ethiopia and Eritrea. Euscorpius, 138: 1 - 25." type="journal article" year="2016">Lowe &amp; Kovařík, 2016: 1</bibRefCitation>
, 43
</treatmentCitation>
;
<treatmentCitation author="LORIA, S. F. &amp; L. PRENDINI" box="[1168,1437,567,590]" page="" pageId="3" pageNumber="2" year="2018">
<bibRefCitation author="Loria &amp; Prendini" box="[1168,1437,567,590]" firstAuthor="Loria" journalOrPublisher="Zoologischer Anzeiger" pageId="3" pageNumber="2" pagination="183 - 191" part="273" refId="ref34645" refString="LORIA, S. F. &amp; L. PRENDINI. 2018. Ultrastructural comparison of the eyespot and ocelli of scorpions, and implications for the systematics of Chaerilidae Pocock, 1893. Zoologischer Anzeiger, 273: 183 - 191." title="Ultrastructural comparison of the eyespot and ocelli of scorpions, and implications for the systematics of Chaerilidae Pocock, 1893" type="journal article" year="2018">Loria &amp; Prendini, 2018</bibRefCitation>
</treatmentCitation>
: tab. 1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="2" type="type_taxon">
<paragraph blockId="3.[830,1440,659,743]" pageId="3" pageNumber="2">
<collectionCode box="[830,1011,659,682]" pageId="3" pageNumber="2">
<typeStatus box="[830,897,659,682]" pageId="3" pageNumber="2">TYPE</typeStatus>
SPECIES
</collectionCode>
.
<taxonomicName authority="Simon, 1882" authorityName="Simon" authorityYear="1882" box="[1022,1402,659,682]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis box="[1022,1256,659,682]" italics="true" pageId="3" pageNumber="2">Butheolus thalassinus</emphasis>
<bibRefCitation author="Simon" box="[1263,1402,659,682]" firstAuthor="Simon" pageId="3" pageNumber="2" pagination="207 - 260" refId="ref35698" refString="SIMON, E. 1882. Etude sur les Arachnides de l'Yemen meridional. Viaggio ad Assab nel Mar Rosso, dei Signori G. Doria ed O. Beccari con il R. Avviso ' Esploratore' dal 16 nov. 1879 al 26 feb. 1880. Annali del Museo Civico di Storia Naturale di Genova, 18: 207 - 260." type="journal article" year="1882">Simon, 1882</bibRefCitation>
</taxonomicName>
, by subsequent designation (
<bibRefCitation author="Kraepelin" box="[1112,1333,690,713]" firstAuthor="Kraepelin" page="5" pageId="3" pageNumber="2" pagination="1 - 24" refId="ref34268" refString="KRAEPELIN, K. 1895. Nachtrag zu Theil I der Revision der Scorpione. Beiheft zum Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 12: 1 - 24." type="journal article" year="1895">Kraepelin, 1895: 5</bibRefCitation>
;
<bibRefCitation author="Pocock" firstAuthor="Pocock" page="295" pageId="3" pageNumber="2" pagination="292 - 316" refId="ref35417" refString="POCOCK, R. I. 1895. On the Arachnida and Myriopoda obtained by Dr. Anderson's collector during Mr T. Brent's expedition to the Hadramaut, South Arabia, with a supplement upon the scorpions obtained by Dr. Anderson in Egypt and the Eastern Soudan. Journal of the Linnean Society, 25: 292 - 316." type="journal article" year="1895">Pocock, 1895: 295</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="2" type="discussion">
<paragraph blockId="3.[830,1441,782,1234]" pageId="3" pageNumber="2">
<bibRefCitation author="Simon" box="[878,1033,782,805]" firstAuthor="Simon" pageId="3" pageNumber="2" pagination="207 - 260" refId="ref35698" refString="SIMON, E. 1882. Etude sur les Arachnides de l'Yemen meridional. Viaggio ad Assab nel Mar Rosso, dei Signori G. Doria ed O. Beccari con il R. Avviso ' Esploratore' dal 16 nov. 1879 al 26 feb. 1880. Annali del Museo Civico di Storia Naturale di Genova, 18: 207 - 260." type="journal article" year="1882">Simon (1882)</bibRefCitation>
described
<taxonomicName authorityName="Simon" authorityYear="1882" box="[1163,1320,782,805]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis box="[1163,1179,782,805]" italics="true" pageId="3" pageNumber="2">B</emphasis>
.
<emphasis box="[1200,1320,782,805]" italics="true" pageId="3" pageNumber="2">thalassinus</emphasis>
</taxonomicName>
as a new species under
<taxonomicName authorityName=": Karsch" authorityYear="1886" box="[984,1091,812,835]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[984,1091,812,835]" italics="true" pageId="3" pageNumber="2">Butheolus</emphasis>
</taxonomicName>
, but did not explicitly designate it as the
<typeStatus box="[945,991,843,866]" pageId="3" pageNumber="2">type</typeStatus>
species. In the appendix to the same publication, he described a second new species under the same genus (
<taxonomicName authorityName="Simon" authorityYear="1882" box="[970,1087,904,927]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="aristidis">
<emphasis box="[970,986,904,927]" italics="true" pageId="3" pageNumber="2">B</emphasis>
.
<emphasis box="[999,1087,904,927]" italics="true" pageId="3" pageNumber="2">aristidis</emphasis>
</taxonomicName>
), leaving ambiguity about which was the
<typeStatus box="[929,975,935,958]" pageId="3" pageNumber="2">type</typeStatus>
species. Later,
<bibRefCitation author="Birula" box="[1157,1367,935,958]" firstAuthor="Birula" page="292" pageId="3" pageNumber="2" pagination="295 - 299" refId="ref32727" refString="BIRULA, A. A. 1898. Miscellanea scorpiologica. III. Zur Synonymie der russischen Skorpione (Forsetzung). Annuaire du Musee Zoologique de I' Academie Imperiale des Sciences de St. - Petersbourg, 3: 295 - 299." type="journal article" year="1898">Birula (1898: 292)</bibRefCitation>
transferred
<taxonomicName authorityName="Simon" authorityYear="1882" box="[905,1027,966,989]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="aristidis">
<emphasis box="[905,921,966,989]" italics="true" pageId="3" pageNumber="2">B</emphasis>
.
<emphasis box="[940,1027,966,989]" italics="true" pageId="3" pageNumber="2">aristidis</emphasis>
</taxonomicName>
to
<taxonomicName authorityName="Karsch" authorityYear="1892" box="[1070,1200,966,989]" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[1070,1200,966,989]" italics="true" pageId="3" pageNumber="2">Orthochirus</emphasis>
</taxonomicName>
. According to
<bibRefCitation author="Fet &amp; Lowe" firstAuthor="Fet" page="88" pageId="3" pageNumber="2" refId="ref33203" refString="FET, V. &amp; G. LOWE. 2000. Family Buthidae. Pp. 54 - 286 in FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. Catalog of the Scorpions of the World (1758 - 1998). The New York Entomological Society." type="book" year="2000">Fet &amp; Lowe (2000: 88)</bibRefCitation>
,
<taxonomicName authorityName="Simon" authorityYear="1882" box="[1033,1186,996,1019]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis box="[1033,1049,996,1019]" italics="true" pageId="3" pageNumber="2">B</emphasis>
.
<emphasis box="[1066,1186,996,1019]" italics="true" pageId="3" pageNumber="2">thalassinus</emphasis>
</taxonomicName>
was established as the
<typeStatus box="[830,876,1027,1050]" pageId="3" pageNumber="2">type</typeStatus>
species of
<taxonomicName authorityName=": Karsch" authorityYear="1886" box="[1004,1111,1027,1050]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[1004,1111,1027,1050]" italics="true" pageId="3" pageNumber="2">Butheolus</emphasis>
</taxonomicName>
by subsequent designation of
<bibRefCitation author="Simon" box="[830,1031,1058,1081]" firstAuthor="Simon" page="77" pageId="3" pageNumber="2" pagination="57 - 87" refId="ref35832" refString="SIMON, E. 1910. Revision des scorpions d'Egypte. Bulletin de la Societe Entomologique d'Egypte (Cairo), 1910: 57 - 87." type="journal article" year="1910">Simon (1910: 77)</bibRefCitation>
. However, a subsequent designation according to
<collectionCode box="[980,1042,1088,1111]" pageId="3" pageNumber="2">ICZN</collectionCode>
<collectingRegion box="[1054,1090,1088,1111]" country="Djibouti" name="Arta" pageId="3" pageNumber="2">Art</collectingRegion>
69.1.1 had already been made earlier by
<bibRefCitation author="Kraepelin" box="[945,1163,1119,1142]" firstAuthor="Kraepelin" page="5" pageId="3" pageNumber="2" pagination="1 - 24" refId="ref34268" refString="KRAEPELIN, K. 1895. Nachtrag zu Theil I der Revision der Scorpione. Beiheft zum Jahrbuch der Hamburgischen Wissenschaftlichen Anstalten, 12: 1 - 24." type="journal article" year="1895">Kraepelin (1895: 5)</bibRefCitation>
and Pocock (1985: 295) when they referenced
<taxonomicName authorityName="Simon" authorityYear="1882" box="[1074,1226,1150,1173]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="thalassinus">
<emphasis box="[1074,1090,1150,1173]" italics="true" pageId="3" pageNumber="2">B</emphasis>
.
<emphasis box="[1106,1226,1150,1173]" italics="true" pageId="3" pageNumber="2">thalassinus</emphasis>
</taxonomicName>
as the
<typeStatus box="[1308,1354,1150,1173]" pageId="3" pageNumber="2">type</typeStatus>
species of
<taxonomicName authorityName=": Karsch" authorityYear="1886" box="[860,967,1180,1203]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis box="[860,967,1180,1203]" italics="true" pageId="3" pageNumber="2">Butheolus</emphasis>
</taxonomicName>
, and it was also cited as the
<typeStatus box="[1276,1322,1180,1203]" pageId="3" pageNumber="2">type</typeStatus>
species by
<bibRefCitation author="Pocock" box="[830,1030,1211,1234]" firstAuthor="Pocock" page="28" pageId="3" pageNumber="2" refId="ref35479" refString="POCOCK, R. I. 1900. Arachnida. The Fauna of British India, including Ceylon and B urma. Published under the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp." type="book" year="1900">Pocock (1900: 28)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="diagnosis">
<paragraph blockId="3.[830,1441,1269,1905]" lastBlockId="5.[830,1440,1739,1793]" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
DIAGNOSIS. Small buthids (
<bibRefCitation author="Kovarik" box="[1171,1338,1269,1292]" firstAuthor="Kovarik" pageId="3" pageNumber="2" refId="ref33839" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Jakub Rolcik - Clairon Production, Prague, 170 pp." type="book" year="2009">Kovařík, 2009</bibRefCitation>
;
<bibRefCitation author="SISSOM, W. D." editor="Polis, G. A." journalOrPublisher="Stanford University Press, Stanford, California" pageId="3" pageNumber="2" pagination="64 - 160" refId="ref35859" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160 in Polis, G. A. (ed.). The Biology of Scorpions. Stanford University Press, Stanford, California." title="Systematics, biogeography and paleontology" type="book chapter" volumeTitle="The Biology of Scorpions" year="1990">Sissom, 1990</bibRefCitation>
), adults
<quantity box="[989,1111,1300,1323]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.6" metricValueMin="2.4" pageId="3" pageNumber="2" unit="mm" value="30.0" valueMax="36.0" valueMin="24.0">2436 mm</quantity>
in length; carapace strongly trapezoidal, ratio of posterior W/ anterior
<geoCoordinate box="[1315,1386,1330,1353]" degrees="1.9" direction="west" orientation="longitude" pageId="3" pageNumber="2" precision="5555" value="-1.9">W 1.9</geoCoordinate>
2.5, preocular area inclined downwards towards anterior margin, surface densely granular, without distinct carinae; tergites densely granular, tergites IIII lacking carinae or with weak median carina, IVVI weakly tricarinate with median and paired lateral carinae; tergite VII with broad median carina, two pairs of lateral carinae; posterior margins of sternites IIIVI armed with fringe of non-contiguous digitate denticles that are larger in males; metasomal segments nearly uniform in width and depth, robust with granulated carinae, segments IIV densely granular on lateral and ventral intercarinal surfaces; metasoma IIII with 10 carinae, IV with 4 or 6 complete carinae (ventrosubmedian, ventrolateral pairs complete, dorsosubmedian pairs may be complete, dorsolateral pairs weak or incomplete), V with 2 complete carinae (ventrolateral pairs); ventrolateral carinae on metasoma V without enlarged lobate dentition; telson with slightly elongated ovoid vesicle, with or without a blunt subaculear tubercle, aculeus shorter than vesicle; pectines with fulcra; hemispermatophore with flagellum separated from a 3-lobed sperm hemiduct, basal lobe a small, narrow, hook-like process; chelicerae with characteristic buthid pattern of dentition (
<bibRefCitation author="Vachon" box="[635,790,1720,1743]" firstAuthor="Vachon" pageId="4" pageNumber="3" pagination="161 - 166" refId="ref36237" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents de cheliceres chez les scorpions. Bulletin du Museum National d'Histoire Naturelle, Paris, (2), 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
), two denticles on ventral aspect of fixed finger; pedipalps orthobothriotaxic,
<typeStatus box="[397,443,1784,1807]" pageId="4" pageNumber="3">type</typeStatus>
Aβ (
<bibRefCitation author="Vachon" box="[511,666,1784,1807]" firstAuthor="Vachon" pageId="4" pageNumber="3" pagination="857 - 958" refId="ref36318" refString="VACHON, M. 1974. Etude des caracteres utilises pour classe les familles et les genre de Scorpiones (Arachnides). 1. La trichobothriotaxie en Arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum National d'Histoire Naturalle Paris, Zoologie, (3) 104 (140): 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
,
<bibRefCitation author="Vachon" box="[686,739,1784,1807]" firstAuthor="Vachon" pageId="4" pageNumber="3" pagination="1597 - 1599" refId="ref36380" refString="VACHON, M. 1975. Sur l'utilisation de la trichobothriotaxie du bras des pedipalps des Scorpions (Arachnides) dans le classement des genres de famille des Buthidae Simon. Compte rendus hebdomadaires des seances de l'Academie des Sciences, Paris Ser. D Sciences Naturelles, 281 (21): 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
), patella with
<emphasis box="[300,313,1814,1837]" italics="true" pageId="4" pageNumber="3">d</emphasis>
<subScript attach="left" box="[313,322,1824,1839]" fontSize="6" pageId="4" pageNumber="3">3</subScript>
internal to dorsomedian carina, chela manus with
<emphasis box="[254,270,1845,1868]" italics="true" pageId="4" pageNumber="3">V</emphasis>
<subScript attach="left" box="[270,279,1855,1870]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
not strongly displaced internally,
<emphasis box="[681,697,1845,1868]" italics="true" pageId="4" pageNumber="3">V</emphasis>
<subScript attach="left" box="[697,706,1855,1870]" fontSize="6" pageId="4" pageNumber="3">1</subScript>
<emphasis box="[720,736,1845,1868]" italics="true" pageId="4" pageNumber="3">V</emphasis>
<subScript attach="left" box="[736,745,1855,1870]" fontSize="6" pageId="4" pageNumber="3">2</subScript>
axis nearly collinear with long axis of chela, chela fixed finger with
<emphasis box="[963,990,1598,1621]" italics="true" pageId="4" pageNumber="3">db</emphasis>
on proximal half of finger between
<emphasis box="[1404,1440,1598,1621]" italics="true" pageId="4" pageNumber="3">esb</emphasis>
and
<emphasis box="[880,910,1628,1651]" italics="true" pageId="4" pageNumber="3">est</emphasis>
,
<emphasis box="[928,943,1628,1651]" italics="true" pageId="4" pageNumber="3">it</emphasis>
near tip; pedipalps short, chelae small with carinae reduced or obsolete, dentate margins of fingers armed with 67 (rarely 8) non-imbricated linear subrows of primary denticles; subrows flanked by mid-row internal and proximal external accessory denticles; movable finger with 3 enlarged subdistal denticles; males without recess or scalloping of dentate margins at base of pedipalp fingers, with chela manus narrower than females; tergites lacking macrosetae; tibial spurs present on legs IIIIV; basitarsi IIII with regular series of long macrosetae on retrosuperior, retroinferior and inferior margins; ventral surfaces of telotarsi with paired rows of macrosetae; prolateral and retrolateral tarsal spurs present on all legs; moderate sexual dimorphism in setation, macrosetae on carapace, pedipalps, legs, sternites and metasoma longer in females than males.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7118818" ID-Zenodo-Dep="7118818" httpUri="https://zenodo.org/record/7118818/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,278,1497,1520]" targetBox="[191,1442,191,1468]" targetPageId="4">
<paragraph blockId="4.[192,1441,1497,1545]" pageId="4" pageNumber="3">
<emphasis bold="true" box="[192,336,1497,1520]" pageId="4" pageNumber="3">Figures 12:</emphasis>
Habitus of male
<taxonomicName authority="Vachon, 1980" authorityName="Vachon" authorityYear="1980" box="[510,862,1499,1520]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="gallagheri">
<emphasis box="[510,714,1499,1520]" italics="true" pageId="4" pageNumber="3">Butheolus gallagheri</emphasis>
<bibRefCitation author="Vachon" box="[722,862,1499,1520]" firstAuthor="Vachon" pageId="4" pageNumber="3" pagination="251 - 263" refId="ref36437" refString="VACHON, M. 1980. Scorpions du Dhofar. The Scientific Results of the Oman Flora and Fauna Survey 1977 (Dhofar). Journal of Oman Studies. Special Report, 2: 251 - 263." type="journal article" year="1980">Vachon, 1980</bibRefCitation>
</taxonomicName>
.
<emphasis bold="true" box="[873,939,1499,1520]" pageId="4" pageNumber="3">Fig. 1.</emphasis>
Dorsal aspect.
<emphasis bold="true" box="[1093,1159,1499,1520]" pageId="4" pageNumber="3">Fig. 2.</emphasis>
Ventral aspect. Oman, wadi above Khor Rori Beach, E. of Taqah, 17°3.2'N 54°25.33'E, 18.X.1993, leg. G. Lowe. Scale bar: 5 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7118820" ID-Zenodo-Dep="7118820" httpUri="https://zenodo.org/record/7118820/files/figure.png" pageId="5" pageNumber="4" startId="5.[192,278,1614,1637]" targetBox="[200,1429,203,1573]" targetPageId="5">
<paragraph blockId="5.[192,1440,1614,1686]" pageId="5" pageNumber="4">
<emphasis bold="true" box="[192,334,1614,1637]" pageId="5" pageNumber="4">Figures 34:</emphasis>
Prosoma and mesosoma of male
<taxonomicName authority="Vachon, 1980" authorityName="Vachon" authorityYear="1980" box="[661,1008,1615,1636]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="gallagheri">
<emphasis box="[661,863,1615,1636]" italics="true" pageId="5" pageNumber="4">Butheolus gallagheri</emphasis>
<bibRefCitation author="Vachon" box="[870,1008,1615,1636]" firstAuthor="Vachon" pageId="5" pageNumber="4" pagination="251 - 263" refId="ref36437" refString="VACHON, M. 1980. Scorpions du Dhofar. The Scientific Results of the Oman Flora and Fauna Survey 1977 (Dhofar). Journal of Oman Studies. Special Report, 2: 251 - 263." type="journal article" year="1980">Vachon, 1980</bibRefCitation>
</taxonomicName>
.
<emphasis bold="true" box="[1018,1082,1615,1636]" pageId="5" pageNumber="4">Fig. 3.</emphasis>
Dorsal aspect, carapace and tergites. Same locality data as male in Figs. 12.
<emphasis bold="true" box="[599,665,1640,1661]" pageId="5" pageNumber="4">Fig. 4.</emphasis>
Ventral aspect, coxosternal area and sternites. Oman, Salalah, leg. A. Ullrich. Scale bar: 1 mm.
</paragraph>
</caption>
<subSubSection lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="discussion">
<paragraph blockId="5.[830,1440,1831,1885]" lastBlockId="6.[192,802,1570,1654]" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
REMARKS. Sexual dimorphism in setation is apparent in
<taxonomicName authorityName=": Karsch" authorityYear="1886" box="[865,972,1862,1885]" class="Arachnida" family="Buthidae" genus="Butheolus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis box="[865,972,1862,1885]" italics="true" pageId="5" pageNumber="4">Butheolus</emphasis>
</taxonomicName>
, but is not as pronounced as it is in
<taxonomicName authority="Hirst, 1911" authorityName="Hirst" authorityYear="1911" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" lastPageId="6" lastPageNumber="5" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis italics="true" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">Neobuthus</emphasis>
<bibRefCitation author="Hirst" box="[274,398,1570,1593]" firstAuthor="Hirst" pageId="6" pageNumber="5" pagination="462 - 473" refId="ref33448" refString="HIRST, S., 1911. Descriptions of new scorpions. Annals and Magazine of Natural History, ser. 8, 7: 462 - 473." type="journal article" year="1911">Hirst, 1911</bibRefCitation>
</taxonomicName>
, in which males usually bear much shorter, stouter macrosetae than females (
<bibRefCitation author="KOVARIK, F. &amp; G. LOWE" journalOrPublisher="Euscorpius" pageId="6" pageNumber="5" pagination="1 - 25" part="139" refId="ref33864" refString="KOVARIK, F. &amp; G. LOWE. 2012. Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae). Euscorpius, 139: 1 - 25." title="Review of the genus Neobuthus Hirst, 1911 with description of a new species from Ethiopia (Scorpiones: Buthidae)" type="journal article" year="2012">Kovařík &amp; Lowe, 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>