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<document id="E6BDB2198991214C20A50EEC6324EDDA" ID-CLB-Dataset="27101" ID-DOI="10.11646/zootaxa.4586.2.9" ID-GBIF-Dataset="3c6556e1-0719-4007-b220-15ba5ee33612" ID-ISSN="1175-5326" ID-Zenodo-Dep="2644554" ID-ZooBank="56B383B3-9F25-4B19-991B-8BEE402B0196" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1555539723141" checkinUser="plazi" docAuthor="Salini, S." docDate="2019" docId="0C3D87C22224E37EFF0F276AB760FE1B" docLanguage="en" docName="zootaxa.4586.2.9.pdf" docOrigin="Zootaxa 4586 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Halys shaista Ghauri 1988" docType="treatment" docVersion="9" lastPageNumber="367" masterDocId="F004FFBA2229E36EFF98222FB236FF95" masterDocTitle="Revision of the genus Halys (Hemiptera: Pentatomidae) with description of a new species from India" masterLastPageNumber="375" masterPageNumber="351" pageNumber="364" updateTime="1698722288033" updateUser="ExternalLinkService">
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<mods:title id="4E0C351748DA2E30D7B03B17B0A3BBD4">Revision of the genus Halys (Hemiptera: Pentatomidae) with description of a new species from India</mods:title>
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<subSubSection id="CC8E655F2224E363FF0F276AB3D3FACA" box="[151,485,1349,1375]" pageId="13" pageNumber="364" type="nomenclature">
<paragraph id="842B36D42224E363FF0F276AB3D3FACA" blockId="13.[151,485,1349,1409]" box="[151,485,1349,1375]" pageId="13" pageNumber="364">
<heading id="DF6381B82224E363FF0F276AB3D3FACA" bold="true" box="[151,485,1349,1375]" fontSize="11" level="1" pageId="13" pageNumber="364" reason="1">
<taxonomicName id="43944D572224E363FF0F276AB3D3FACA" authority="Ghauri, 1988" authorityName="Ghauri" authorityYear="1988" box="[151,485,1349,1375]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="364" phylum="Chordata" rank="species" species="shaista">
<emphasis id="B6E0EAC62224E363FF0F276AB3D3FACA" bold="true" box="[151,485,1349,1375]" pageId="13" pageNumber="364">
<emphasis id="B6E0EAC62224E363FF0F276AB30EFACA" bold="true" box="[151,312,1349,1375]" italics="true" pageId="13" pageNumber="364">Halys shaista</emphasis>
<bibRefCitation id="E0054B252224E363FED8276AB3D3FACA" author="Ghauri, M. S. K." box="[320,485,1349,1375]" pageId="13" pageNumber="364" pagination="77 - 92" refId="ref13220" refString="Ghauri, M. S. K. (1988) A revision of Asian species of the genus Halys Fabricius based on the type material (Insecta, Heteroptera, Pentatomidae, Pentatomidae). Entomologische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 51 (6), 77 - 92." type="journal article" year="1988">Ghauri, 1988</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CC8E655F2224E363FF0F2747B33DFA14" box="[151,267,1384,1409]" pageId="13" pageNumber="364" type="description">
<paragraph id="842B36D42224E363FF0F2747B33DFA14" blockId="13.[151,485,1349,1409]" box="[151,267,1384,1409]" pageId="13" pageNumber="364">
(
<figureCitation id="1CAF2A512224E363FF072747B334FA14" box="[159,258,1384,1409]" captionStart="FIGURES 19" captionStartId="4.[151,264,1939,1962]" captionTargetBox="[246,1377,251,1915]" captionTargetId="figure@4.[183,1409,193,1927]" captionTargetPageId="4" captionText="FIGURES 19. Halys Fabricius, 1803 (dorsal and ventral habitus). 12, Halys mudigerensis sp. nov. (male); 3, H. mudigerensis sp. nov. (female without abdomen); 45, H. serrigera Westwood, 1837; 67, H. shaista Ghauri, 1988; 89, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644556/files/figure.png" pageId="13" pageNumber="364">Figs 6, 7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CC8E655F2224E363FF0F279FB3CEF971" pageId="13" pageNumber="364" type="reference_group">
<paragraph id="842B36D42224E363FF0F279FB010FA53" blockId="13.[151,550,1456,1478]" box="[151,550,1456,1478]" pageId="13" pageNumber="364">
<treatmentCitationGroup id="A48411FA2224E363FF0F279FB010FA53" box="[151,550,1456,1478]" pageId="13" pageNumber="364">
<taxonomicName id="43944D572224E363FF0F279FB3E2FA53" authority="Ghauri, 1988: 78" authorityName="Ghauri" authorityPageNumber="78" authorityYear="1988" box="[151,468,1456,1478]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="13" pageNumber="364" phylum="Chordata" rank="species" species="shaista">
<emphasis id="B6E0EAC62224E363FF0F279FB316FA53" box="[151,288,1456,1478]" italics="true" pageId="13" pageNumber="364">Halys shaista</emphasis>
<treatmentCitation id="053510C52224E363FEBE279FB3E2FA53" author="Ghauri, M. S. K." box="[294,468,1456,1478]" page="78" pageId="13" pageNumber="364" year="1988">
<bibRefCitation id="E0054B252224E363FEBE279FB399FA53" author="Ghauri, M. S. K." box="[294,431,1456,1478]" pageId="13" pageNumber="364" pagination="77 - 92" refId="ref13220" refString="Ghauri, M. S. K. (1988) A revision of Asian species of the genus Halys Fabricius based on the type material (Insecta, Heteroptera, Pentatomidae, Pentatomidae). Entomologische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 51 (6), 77 - 92." type="journal article" year="1988">Ghauri, 1988</bibRefCitation>
: 78
</treatmentCitation>
</taxonomicName>
, 8892.
</treatmentCitationGroup>
</paragraph>
<paragraph id="842B36D42224E363FF0F27DCB3CEF971" blockId="13.[151,1437,1523,2015]" pageId="13" pageNumber="364">
<emphasis id="B6E0EAC62224E363FF0F27DCB315F999" bold="true" box="[151,291,1523,1548]" italics="true" pageId="13" pageNumber="364">Colouration</emphasis>
. Dorsum ochraceous with spots or streaks formed by clusters of dark brown or black punctures; antennae nearly concolourous with dorsum, apical three segments darker, apex of segment II and III and base of segment IV and V, ochraceous; scutellum with 1+1 spots at basal angles, black; connexivum concolourous with dorsum except for light to dark brown anterior and posterior fourths of connexival segments; hemelytral membrane light brown with black veins; ventral surface including labium and legs nearly concolourous or paler than dorsum except for black apical segment of labium, apices of tarsal segments and claws and anterior and posterior angles of ventrites and spiracular outline.
</paragraph>
</subSubSection>
<subSubSection id="CC8E655F2224E360FF5F24C0B2D9F85A" lastPageId="14" lastPageNumber="365" pageId="13" pageNumber="364" type="description">
<paragraph id="842B36D42224E363FF5F24C0B06CF8DA" blockId="13.[151,1437,1523,2015]" pageId="13" pageNumber="364">
<emphasis id="B6E0EAC62224E363FF5F24C0B3C9F892" bold="true" box="[199,511,1774,1799]" italics="true" pageId="13" pageNumber="364">Integument and vestiture.</emphasis>
Dorsum including head covered with dense, irregular dark brown to black punctures, sometimes forming irregular patches or spots. Ventral surface including legs with dark brown to black, coarse, dense punctures. Body glabrous.
</paragraph>
<paragraph id="842B36D42224E363FF5F2574B094F829" blockId="13.[151,1437,1523,2015]" pageId="13" pageNumber="364">
<emphasis id="B6E0EAC62224E363FF5F2574B30FF8E1" bold="true" box="[199,313,1883,1908]" italics="true" pageId="13" pageNumber="364">Structure.</emphasis>
Labium elongate reaching anterior margin of abdominal segment VII. Anterior and posterior margins of pronotum nearly straight. Humeri angulate, not elevated, disc of pronotum and scutellum more or less flat. Other characters as in generic description.
</paragraph>
<paragraph id="842B36D42224E360FF5F25E9B3A0FD58" blockId="13.[151,1437,1523,2015]" lastBlockId="14.[151,1437,151,717]" lastPageId="14" lastPageNumber="365" pageId="13" pageNumber="364">
<emphasis id="B6E0EAC62224E363FF5F25E9B0A3F84A" bold="true" box="[199,661,1990,2015]" pageId="13" pageNumber="364">
External male genitalia (
<figureCitation id="1CAF2A512224E363FE6625E9B0B1F84A" box="[510,647,1990,2015]" captionStart-0="FIGURES 6168" captionStart-1="FIGURES 6976" captionStartId-0="14.[151,264,1635,1658]" captionStartId-1="15.[151,264,1121,1144]" captionTargetBox-0="[205,1394,783,1545]" captionTargetBox-1="[181,1411,227,1050]" captionTargetId-0="figure@14.[181,1407,756,1623]" captionTargetId-1="figure@15.[151,1435,193,1101]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURES 6168. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." captionText-1="FIGURES 6976. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri-0="https://zenodo.org/record/2644574/files/figure.png" httpUri-1="https://zenodo.org/record/2644576/files/figure.png" pageId="13" pageNumber="364">Figs 6176</figureCitation>
).
</emphasis>
<emphasis id="B6E0EAC62224E363FD3A25E9B157F84A" box="[674,865,1990,2015]" italics="true" pageId="13" pageNumber="364">Genital capsule.</emphasis>
Quadrangular. Dorsal rim outlined with mat of minute, fine hairs, widely and shallowly excavated concave, with median short region, black and glabrous (
<figureCitation id="1CAF2A512227E360FAF722B8B280FF40" captionStart="FIGURES 6168" captionStartId="14.[151,264,1635,1658]" captionTargetBox="[205,1394,783,1545]" captionTargetId="figure@14.[181,1407,756,1623]" captionTargetPageId="14" captionText="FIGURES 6168. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="14" pageNumber="365">Fig. 61</figureCitation>
); slightly notched adjacent to caudal lobes; infoldings with sublateral region sclerotized black with tooth reduced to mere sclerotized, minute serrations towards end of sclerotized region on either side of median black region (
<figureCitation id="1CAF2A512227E360FF69232BB37FFE89" box="[241,329,260,285]" captionStart="FIGURES 9499" captionStartId="22.[151,264,1273,1296]" captionTargetBox="[170,1415,467,1204]" captionTargetId="figure@22.[151,1435,409,1253]" captionTargetPageId="22" captionText="FIGURES 9499. FIGURES 9495. Genital capsule (caudal). 94, H. shaista Ghauri, 1988; 95, H. sulcata (Thunberg, 1783). Lettering: t—tooth; ts—tooth like serrations. FIGURES 9699. Capitate processes. 96, H. mudigerensis sp. nov.; 97, H. serrigera Westwood, 1837; 98, H. shaista Ghauri, 1988; 99, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644584/files/figure.png" pageId="14" pageNumber="365">Fig. 94</figureCitation>
); ventral rim together with infoldings broadly and widely excavated concave, lack the median, inverted U-shaped process unlike in other species and with a median tumescence, notched medially (
<figureCitation id="1CAF2A512227E360FA9F2308B76AFED5" box="[1287,1372,295,320]" captionStart="FIGURES 6168" captionStartId="14.[151,264,1635,1658]" captionTargetBox="[205,1394,783,1545]" captionTargetId="figure@14.[181,1407,756,1623]" captionTargetPageId="14" captionText="FIGURES 6168. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="14" pageNumber="365">Fig. 62</figureCitation>
); 1+1 sclerotized, obliquely straight narrow ridge on sides of median tumescence on inner side; ridges reaching caudolateral margin of infoldings; infoldings well developed, elevated, laterally joined to caudal lobes by a minute V-notch.
<emphasis id="B6E0EAC62227E360FF6723BAB340FE39" box="[255,374,405,428]" italics="true" pageId="14" pageNumber="365">Paramere.</emphasis>
Roughly C-shaped (
<figureCitation id="1CAF2A512227E360FDF923BBB0D2FE38" box="[609,740,404,429]" captionStart="FIGURES 6168" captionStartId="14.[151,264,1635,1658]" captionTargetBox="[205,1394,783,1545]" captionTargetId="figure@14.[181,1407,756,1623]" captionTargetPageId="14" captionText="FIGURES 6168. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="14" pageNumber="365">Figs 6466</figureCitation>
), with apically angulate and expanded crown, short stem and expanded plate-like apodeme. Crown with short, blunt, black tooth medially on dorsolateral margin.
<emphasis id="B6E0EAC62227E360FAA72398B7AAFE45" box="[1343,1436,439,464]" italics="true" pageId="14" pageNumber="365">Phallus.</emphasis>
Articulatory apparatus with basal plates and support bridge complex, dorsal connectives widened into 1+1 oval capitate processes (
<figureCitation id="1CAF2A512227E360FEEF23D0B3E4FD8D" box="[375,466,511,536]" captionStart="FIGURES 6168" captionStartId="14.[151,264,1635,1658]" captionTargetBox="[205,1394,783,1545]" captionTargetId="figure@14.[181,1407,756,1623]" captionTargetPageId="14" captionText="FIGURES 6168. H. shaista Ghauri, 1988—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral)." httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="14" pageNumber="365">Figs 67</figureCitation>
,
<figureCitation id="1CAF2A512227E360FE79202FB037FD8D" box="[481,513,512,536]" captionStart="FIGURES 9499" captionStartId="22.[151,264,1273,1296]" captionTargetBox="[170,1415,467,1204]" captionTargetId="figure@22.[151,1435,409,1253]" captionTargetPageId="22" captionText="FIGURES 9499. FIGURES 9495. Genital capsule (caudal). 94, H. shaista Ghauri, 1988; 95, H. sulcata (Thunberg, 1783). Lettering: t—tooth; ts—tooth like serrations. FIGURES 9699. Capitate processes. 96, H. mudigerensis sp. nov.; 97, H. serrigera Westwood, 1837; 98, H. shaista Ghauri, 1988; 99, H. sulcata (Thunberg, 1783)" httpUri="https://zenodo.org/record/2644584/files/figure.png" pageId="14" pageNumber="365">98</figureCitation>
). Phallosoma sclerotized, constricted towards both ends, broadest medially and concave dorsally; two pairs of membranous conjunctival processes, dorsal pair elongate with bifid apex (
<figureCitation id="1CAF2A512227E360FAA7200BB280FDF5" captionStart="FIGURES 6976" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 6976. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="14" pageNumber="365">Figs 69, 70</figureCitation>
), ventral pair shorter than dorsal pair (
<figureCitation id="1CAF2A512227E360FDFA2068B0D2FDF5" box="[610,740,583,608]" captionStart="FIGURES 6976" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 6976. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="14" pageNumber="365">Figs 71, 72</figureCitation>
). Processes of aedeagus hard, sclerotized, fused along midline at proximal half encompassing short, sclerotized aedeagus and separated at distal half; distal half expanded into a subrectangular plate with rounded apical margin (
<figureCitation id="1CAF2A512227E360FD4720A0B10EFD3D" box="[735,824,655,680]" captionStart="FIGURES 6976" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 6976. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="14" pageNumber="365">Fig. 72</figureCitation>
); aedeagus obliquely bent upwards and shorter than processes of aedeagus.
</paragraph>
<caption id="D0EB665C2227E360FF0F244CB097F922" httpUri="https://zenodo.org/record/2644574/files/figure.png" pageId="14" pageNumber="365" startId="14.[151,264,1635,1658]" targetBox="[205,1394,783,1545]" targetPageId="14">
<paragraph id="842B36D42227E360FF0F244CB097F922" blockId="14.[151,1436,1635,1719]" pageId="14" pageNumber="365">
<emphasis id="B6E0EAC62227E360FF0F244CB361F9ED" bold="true" box="[151,343,1635,1658]" pageId="14" pageNumber="365">FIGURES 6168.</emphasis>
<taxonomicName id="43944D572227E360FEF8244BB06AF9EC" authority="Ghauri, 1988" authorityName="Ghauri" authorityYear="1988" box="[352,604,1635,1657]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="14" pageNumber="365" phylum="Chordata" rank="species" species="shaista">
<emphasis id="B6E0EAC62227E360FEF8244BB3FDF9EC" box="[352,459,1635,1657]" italics="true" pageId="14" pageNumber="365">H. shaista</emphasis>
Ghauri, 1988
</taxonomicName>
—male genitalia. 61, genital capsule (dorsal); 62, genital capsule (ventral); 63, genital capsule (caudal); 64, paramere (dorsal); 65, paramere (dorsolateral); 66, paramere (ventral); 67, articulatory apparatus; 68, phallus without articulatory apparatus (ventral).
</paragraph>
</caption>
<paragraph id="842B36D42227E360FF5F24F1B2D9F85A" blockId="14.[151,1437,1758,1999]" pageId="14" pageNumber="365">
<emphasis id="B6E0EAC62227E360FF5F24F1B094F962" bold="true" box="[199,674,1758,1783]" pageId="14" pageNumber="365">
External female genitalia (
<figureCitation id="1CAF2A512227E360FD9524F1B0A2F962" box="[525,660,1758,1783]" captionStart="FIGURES 6976" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 6976. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="14" pageNumber="365">Figs 7376</figureCitation>
).
</emphasis>
<emphasis id="B6E0EAC62227E360FD3524F1B119F962" box="[685,815,1758,1783]" italics="true" pageId="14" pageNumber="365">Terminalia.</emphasis>
Valvifers VIII roughly triangular, with mesal margin straight, inner posterior angles rounded; posterior margin nearly straight or slightly convex. Valvulae VIII with apical (= caudal) margin not uniformly rounded, but distinctly indented medially (roughly broad W-shaped) forming two lobes (
<figureCitation id="1CAF2A512227E360FEEE2565B3FDF8F6" box="[374,459,1866,1891]" captionStart="FIGURES 6976" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 6976. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="14" pageNumber="365">Fig. 76</figureCitation>
). Valvifers IX single, transverse, narrow, inverted trapezoidal sclerite. Laterotergite IX obliquely placed, not reaching apex of abdomen. Laterotergite VIII broad, elongate, triangular with nearly smooth caudal margin.
<emphasis id="B6E0EAC62227E360FEDE25BDB3EEF83E" box="[326,472,1938,1963]" italics="true" pageId="14" pageNumber="365">Spermatheca</emphasis>
(
<figureCitation id="1CAF2A512227E360FE7025BDB00DF83E" box="[488,571,1938,1963]" captionStart="FIGURES 6976" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 6976. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="14" pageNumber="365">Fig. 73</figureCitation>
) apical receptacle orbicular with two finger-like processes, directed towards flanges.
</paragraph>
</subSubSection>
<caption id="D0EB665C2226E361FF0F264EB2FCFB20" httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="15" pageNumber="366" startId="15.[151,264,1121,1144]" targetBox="[181,1411,227,1050]" targetPageId="15">
<paragraph id="842B36D42226E361FF0F264EB2FCFB20" blockId="15.[151,1436,1121,1205]" pageId="15" pageNumber="366">
<emphasis id="B6E0EAC62226E361FF0F264EB361FBE2" bold="true" box="[151,343,1121,1144]" pageId="15" pageNumber="366">FIGURES 6976.</emphasis>
<taxonomicName id="43944D572226E361FEF8264CB06AFBED" authority="Ghauri, 1988" authorityName="Ghauri" authorityYear="1988" box="[352,604,1122,1144]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="15" pageNumber="366" phylum="Chordata" rank="species" species="shaista">
<emphasis id="B6E0EAC62226E361FEF8264CB34CFBED" box="[352,378,1123,1144]" italics="true" pageId="15" pageNumber="366">H.</emphasis>
<emphasis id="B6E0EAC62226E361FE1B264DB3FDFBED" box="[387,459,1122,1144]" italics="true" pageId="15" pageNumber="366">shaista</emphasis>
Ghauri, 1988
</taxonomicName>
—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII.
</paragraph>
</caption>
<subSubSection id="CC8E655F2226E361FF5F26CDB608FA1F" pageId="15" pageNumber="366" type="diagnosis">
<paragraph id="842B36D42226E361FF5F26CDB608FA1F" blockId="15.[151,1437,1250,2031]" pageId="15" pageNumber="366">
<emphasis id="B6E0EAC62226E361FF5F26CDB3F8FB6E" bold="true" box="[199,462,1250,1275]" pageId="15" pageNumber="366">Differential diagnosis.</emphasis>
This species can be differentiated from other species of
<taxonomicName id="43944D572226E361FBDF26CDB6BEFB6E" authorityName="Fabricius" authorityYear="1803" box="[1095,1160,1250,1275]" class="Squamata" family="Viperidae" genus="Halys" higherTaxonomySource="CoL" kingdom="Animalia" pageId="15" pageNumber="366" phylum="Chordata" rank="genus">
<emphasis id="B6E0EAC62226E361FBDF26CDB6BEFB6E" box="[1095,1160,1250,1275]" italics="true" pageId="15" pageNumber="366">Halys</emphasis>
</taxonomicName>
by the peculiar shape of paramere and absence of inverted U-shaped process on ventral rim of genital capsule but with a median tumescence. The parameral crown is apically angulate, possessing a single tooth around the middle of its dorsolateral magin, triangular valvifers VIII with posterior margin nearly straight or slightly convex (
<figureCitation id="1CAF2A512226E361FA8A2761B7A2FAF2" box="[1298,1428,1358,1383]" captionStart="FIGURES 6976" captionStartId="15.[151,264,1121,1144]" captionTargetBox="[181,1411,227,1050]" captionTargetId="figure@15.[151,1435,193,1101]" captionTargetPageId="15" captionText="FIGURES 6976. H. shaista Ghauri, 1988—male and female genitalia. 69, phallus (dorsal); 70, phallus (ventral); 7172, phallus (lateral); 73, spermatheca; 74, female terminalia (before digestion); 75, female terminalia (after digestion); 76, valvulae VIII." httpUri="https://zenodo.org/record/2644576/files/figure.png" pageId="15" pageNumber="366">Figs 74, 75</figureCitation>
) and valvulae VIII distinctly indented medially (roughly broad W-shaped) (
<tableCitation id="C916036F2226E361FC41275DB61BFA1F" box="[985,1069,1394,1419]" captionStart="TABLE 1" captionStartId="20.[151,239,185,208]" captionTargetPageId="20" captionText="TABLE 1. Diagnostic characters of species of Halys from India" httpUri="http://table.plazi.org/id/D0EB665C223DE37AFF0F2296B177FF45" pageId="15" pageNumber="366" tableUuid="D0EB665C223DE37AFF0F2296B177FF45">Table 1</tableCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CC8E655F2226E361FF5F27B9B706F966" pageId="15" pageNumber="366" type="description">
<paragraph id="842B36D42226E361FF5F27B9B706F966" blockId="15.[151,1437,1250,2031]" pageId="15" pageNumber="366">
<emphasis id="B6E0EAC62226E361FF5F27B9B3DBFA3A" bold="true" box="[199,493,1430,1455]" pageId="15" pageNumber="366">Measurements (in mm).</emphasis>
Median (minimummaximum).
<emphasis id="B6E0EAC62226E361FCF427B9B185FA3A" bold="true" box="[876,947,1430,1455]" pageId="15" pageNumber="366">Males</emphasis>
(n=5). Body length 18.39 (17.2019.52); head length 4.56 (4.184.83), head width 3.23 (3.053.44), interocular width 1.77 (1.581.97), lengths of antennal segments (I) 1.11 (1.041.19): (II) 2.35 (2.062.52): (III) 2.50 (2.362.70): (IV) 2.36 (2.352.36): (V) 1.90 (1.89 1.91); lengths of labial segments (I) 2.31 (2.152.38): (II) 4.17 (3.844.51): (III) 4.75 (4.424.95): (IV) 3.00 (2.79 3.17); pronotum length 3.93 (3.724.13); pronotum width 8.71 (8.039.34); scutellum length 6.66 (6.057.00); scutellum width 4.88 (4.365.23).
<emphasis id="B6E0EAC62226E361FD812465B04AF9F6" bold="true" box="[537,636,1610,1635]" pageId="15" pageNumber="366">Females</emphasis>
(n=5). Body length 21.07 (19.2722.96); head length 4.89 (4.475.20), head width 3.44 (3.343.61), interocular width 1.97 (1.832.12), lengths of antennal segments (I) 1.13 (1.041.33): (II) 2.38 (2.252.61): (III) 2.39 (2.312.57): (IV) 2.25 (2.152.34): (V) 1.78; lengths of labial segments (I) 2.45 (2.262.67): (II) 4.32 (4.214.51): (III) 4.75 (4.515.00): (IV) 3.00 (2.793.21); pronotum length 4.25 (3.824.53); pronotum width 9.28 (8.2710.24); scutellum length 7.26 (6.487.86); scutellum width 5.25 (4.645.78).
</paragraph>
</subSubSection>
<subSubSection id="CC8E655F2226E37EFF5F24D1B6A1FED3" lastPageId="16" lastPageNumber="367" pageId="15" pageNumber="366" type="materials_examined">
<paragraph id="842B36D42226E37EFF5F24D1B6A1FED3" blockId="15.[151,1437,1250,2031]" lastBlockId="16.[151,1437,151,398]" lastPageId="16" lastPageNumber="367" pageId="15" pageNumber="366">
<emphasis id="B6E0EAC62226E361FF5F24D1B387F882" bold="true" box="[199,433,1790,1815]" pageId="15" pageNumber="366">Material examined.</emphasis>
<materialsCitation id="34FC3C892226E361FE2024D0B187F882" ID-GBIF-Occurrence="2236308472" box="[440,945,1790,1815]" collectingDate="1976-04-08" country="India" location="Hebbal" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<collectingCountry id="FC8376442226E361FE2024D0B03AF882" box="[440,524,1791,1815]" name="India" pageId="15" pageNumber="366">INDIA</collectingCountry>
:
<collectingRegion id="4650F8362226E361FD8E24D1B0AFF882" box="[534,665,1790,1815]" country="India" name="Karnataka" pageId="15" pageNumber="366">
<emphasis id="B6E0EAC62226E361FD8E24D1B0AFF882" bold="true" box="[534,665,1790,1815]" pageId="15" pageNumber="366">Karnataka</emphasis>
</collectingRegion>
:
<specimenCount id="9292FD5D2226E361FD3024D0B0FDF882" box="[680,715,1791,1815]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
,
<location id="814B600F2226E361FD4124D1B11BF882" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FD4124D1B11BF882" box="[729,813,1790,1815]" country="India" name="Hebbal" pageId="15" pageNumber="366" stateProvince="Karnataka">Hebbal</location>
,
<collectingDate id="E06EE9FC2226E361FCA024D1B187F882" box="[824,945,1790,1815]" pageId="15" pageNumber="366" value="1976-04-08">
<date id="F02A10142226E361FCA024D1B187F882" box="[824,945,1790,1815]" pageId="15" pageNumber="366" value="1976-04-08">08.iv.1976</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="34FC3C892226E361FC2624D0B754F883" ID-GBIF-Occurrence="2236308487" box="[958,1378,1790,1815]" collectingDate="1979-05-13" collectorName="Kumar, A. R. V" country="India" location="Sirsi" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FC2624D0B1D6F882" box="[958,992,1791,1815]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
,
<location id="814B600F2226E361FC7724D1B613F882" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FC7724D1B613F882" box="[1007,1061,1790,1815]" country="India" name="Sirsi" pageId="15" pageNumber="366" stateProvince="Karnataka">Sirsi</location>
,
<collectingDate id="E06EE9FC2226E361FBA824D0B697F882" box="[1072,1185,1791,1815]" pageId="15" pageNumber="366" value="1979-05-13">
<date id="F02A10142226E361FBA824D0B697F882" box="[1072,1185,1791,1815]" pageId="15" pageNumber="366" value="1979-05-13">13.v.1979</date>
</collectingDate>
,
<collectorName id="296153022226E361FB3524D0B754F883" box="[1197,1378,1791,1815]" pageId="15" pageNumber="366">Kumar, A. R. V</collectorName>
</materialsCitation>
.;
<materialsCitation id="34FC3C892226E361FAEA24D0B3D4F8AE" ID-GBIF-Occurrence="2236308479" collectingDate="1979-09-03" country="India" elevation="916" location="Bangalore" pageId="15" pageNumber="366" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FAEA24D0B7A0F882" box="[1394,1430,1791,1815]" pageId="15" pageNumber="366" type="male">1♂</specimenCount>
,
<location id="814B600F2226E361FF0F250DB33BF8AE" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FF0F250DB33BF8AE" box="[151,269,1826,1851]" country="India" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
,
<elevation id="0FB9D1E72226E361FE81250DB36DF8AE" box="[281,347,1826,1851]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="366" unit="m" value="916.0">
<quantity id="436C9B312226E361FE81250DB36DF8AE" box="[281,347,1826,1851]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="366" unit="m" value="916.0">916m</quantity>
</elevation>
,
<collectingDate id="E06EE9FC2226E361FEFF250DB3D4F8AE" box="[359,482,1826,1851]" pageId="15" pageNumber="366" value="1979-09-03">
<date id="F02A10142226E361FEFF250DB3D4F8AE" box="[359,482,1826,1851]" pageId="15" pageNumber="366" value="1979-09-03">03.ix.1979</date>
</collectingDate>
</materialsCitation>
;
<materialsCitation id="34FC3C892226E361FE77250DB1AEF8AE" ID-GBIF-Occurrence="2236308474" box="[495,920,1826,1851]" collectingDate="1982-08-15" country="India" location="Bangalore" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FE77250DB024F8AF" box="[495,530,1826,1850]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
,
<location id="814B600F2226E361FDB9250DB0A1F8AE" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FDB9250DB0A1F8AE" box="[545,663,1826,1851]" country="India" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
,
<collectingDate id="E06EE9FC2226E361FD3B250DB11AF8AE" box="[675,812,1826,1851]" pageId="15" pageNumber="366" value="1982-08-15">
<date id="F02A10142226E361FD3B250DB11AF8AE" box="[675,812,1826,1851]" pageId="15" pageNumber="366" value="1982-08-15">15.viii.1982</date>
</collectingDate>
, students
</materialsCitation>
;
<specimenCount id="9292FD5D2226E361FC3C250DB1F1F8AF" box="[932,967,1826,1850]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
, Hebbal, students;
<materialsCitation id="34FC3C892226E361FB3A250DB3F7F8CB" ID-GBIF-Occurrence="2236308476" collectingDate="1988-08-08" collectorName="Hegde, S. K" country="India" elevation="916" location="Bangalore" pageId="15" pageNumber="366" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FB3A250DB6F0F8AF" box="[1186,1222,1826,1850]" pageId="15" pageNumber="366" type="male">1♂</specimenCount>
,
<location id="814B600F2226E361FB4C250DB77CF8AE" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FB4C250DB77CF8AE" box="[1236,1354,1826,1851]" country="India" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
,
<elevation id="0FB9D1E72226E361FACE250DB7AEF8AE" box="[1366,1432,1826,1851]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="366" unit="m" value="916.0">
<quantity id="436C9B312226E361FACE250DB7AEF8AE" box="[1366,1432,1826,1851]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="366" unit="m" value="916.0">916m</quantity>
</elevation>
,
<collectingDate id="E06EE9FC2226E361FF0F2569B317F8CA" box="[151,289,1862,1887]" pageId="15" pageNumber="366" value="1988-08-08">
<date id="F02A10142226E361FF0F2569B317F8CA" box="[151,289,1862,1887]" pageId="15" pageNumber="366" value="1988-08-08">08.viii.1988</date>
</collectingDate>
,
<collectorName id="296153022226E361FEB62569B3F7F8CB" box="[302,449,1862,1887]" pageId="15" pageNumber="366">Hegde, S. K</collectorName>
</materialsCitation>
.;
<specimenCount id="9292FD5D2226E361FE4B2568B3C0F8CA" box="[467,502,1863,1887]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
, Mudigere, students;
<materialsCitation id="34FC3C892226E361FD772568B622F8CA" ID-GBIF-Occurrence="2236308480" box="[751,1044,1862,1887]" collectingDate="1993-02-06" country="India" location="Mudigere" pageId="15" pageNumber="366" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FD772568B122F8CA" box="[751,788,1863,1887]" pageId="15" pageNumber="366" type="male">1♂</specimenCount>
,
<location id="814B600F2226E361FCBA2569B1A4F8CA" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FCBA2569B1A4F8CA" box="[802,914,1862,1887]" country="India" name="Mudigere" pageId="15" pageNumber="366" stateProvince="Karnataka">Mudigere</location>
,
<collectingDate id="E06EE9FC2226E361FC072569B622F8CA" box="[927,1044,1862,1887]" pageId="15" pageNumber="366" value="1993-02-06">
<date id="F02A10142226E361FC072569B622F8CA" box="[927,1044,1862,1887]" pageId="15" pageNumber="366" value="1993-02-06">06.ii.1993</date>
</collectingDate>
</materialsCitation>
;
<specimenCount id="9292FD5D2226E361FBBA2568B673F8CA" box="[1058,1093,1863,1887]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
, Bangalore, GKVK, students;
<materialsCitation id="34FC3C892226E361FF0F2545B052F816" ID-GBIF-Occurrence="2236308481" box="[151,612,1898,1923]" collectingDate="2004-04-11" collectorName="Hari" country="India" elevation="916" location="Bangalore" pageId="15" pageNumber="366" specimenCount="3" specimenCount-female="3" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FF0F2545B28DF817" box="[151,187,1898,1922]" pageId="15" pageNumber="366" type="female">3♀</specimenCount>
,
<location id="814B600F2226E361FF552545B375F816" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FF552545B375F816" box="[205,323,1898,1923]" country="India" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
,
<elevation id="0FB9D1E72226E361FECC2545B3A0F816" box="[340,406,1898,1923]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="366" unit="m" value="916.0">
<quantity id="436C9B312226E361FECC2545B3A0F816" box="[340,406,1898,1923]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="366" unit="m" value="916.0">916m</quantity>
</elevation>
,
<collectingDate id="E06EE9FC2226E361FE3F2545B028F816" box="[423,542,1898,1923]" pageId="15" pageNumber="366" value="2004-04-11">
<date id="F02A10142226E361FE3F2545B028F816" box="[423,542,1898,1923]" pageId="15" pageNumber="366" value="2004-04-11">11.iv.2004</date>
</collectingDate>
,
<collectorName id="296153022226E361FDB72545B052F816" box="[559,612,1898,1923]" pageId="15" pageNumber="366">Hari</collectorName>
</materialsCitation>
;
<materialsCitation id="34FC3C892226E361FDEE2545B1C3F816" ID-GBIF-Occurrence="2236308457" box="[630,1013,1898,1923]" collectingDate="2005-02-14" country="India" location="Hebbal" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FDEE2545B0AEF817" box="[630,664,1898,1922]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
,
<location id="814B600F2226E361FD342545B136F816" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FD342545B136F816" box="[684,768,1898,1923]" country="India" name="Hebbal" pageId="15" pageNumber="366" stateProvince="Karnataka">Hebbal</location>
,
<collectingDate id="E06EE9FC2226E361FC892545B1B2F816" box="[785,900,1898,1923]" pageId="15" pageNumber="366" value="2005-02-14">
<date id="F02A10142226E361FC892545B1B2F816" box="[785,900,1898,1923]" pageId="15" pageNumber="366" value="2005-02-14">14.ii.2005</date>
</collectingDate>
, students
</materialsCitation>
;
<materialsCitation id="34FC3C892226E361FB9E2545B2C0F832" ID-GBIF-Occurrence="2236308495" collectingDate="2005-05-05" country="India" location="Bangalore" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FB9E2545B61FF817" box="[1030,1065,1898,1922]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
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<location id="814B600F2226E361FBA42545B684F816" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FBA42545B684F816" box="[1084,1202,1898,1923]" country="India" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
, GKVK,
<collectingDate id="E06EE9FC2226E361FABF2545B7A1F816" box="[1319,1431,1898,1923]" pageId="15" pageNumber="366" value="2005-05-05">
<date id="F02A10142226E361FABF2545B7A1F816" box="[1319,1431,1898,1923]" pageId="15" pageNumber="366" value="2005-05-05">05.v.2005</date>
</collectingDate>
, students
</materialsCitation>
;
<specimenCount id="9292FD5D2226E361FE9225A0B318F832" box="[266,302,1935,1959]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
;
<materialsCitation id="34FC3C892226E361FEDC25A0B10AF832" ID-GBIF-Occurrence="2236308475" box="[324,828,1934,1959]" collectingDate="2005-05-21" country="India" elevation="980" location="Mudigere" pageId="15" pageNumber="366" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FEDC25A0B35FF832" box="[324,361,1935,1959]" pageId="15" pageNumber="366" type="male">1♂</specimenCount>
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<location id="814B600F2226E361FEE625A1B3D8F832" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FEE625A1B3D8F832" box="[382,494,1934,1959]" country="India" name="Mudigere" pageId="15" pageNumber="366" stateProvince="Karnataka">Mudigere</location>
,
<quantity id="436C9B312226E361FD9925A0B072F832" box="[513,580,1935,1959]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="15" pageNumber="366" unit="m" value="980.0">
<elevation id="0FB9D1E72226E361FD9925A0B072F832" box="[513,580,1935,1959]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="15" pageNumber="366" unit="m" value="980.0">980m</elevation>
</quantity>
,
<collectingDate id="E06EE9FC2226E361FDC025A1B0FEF832" box="[600,712,1934,1959]" pageId="15" pageNumber="366" value="2005-05-21">
<date id="F02A10142226E361FDC025A1B0FEF832" box="[600,712,1934,1959]" pageId="15" pageNumber="366" value="2005-05-21">21.v.2005</date>
</collectingDate>
, students
</materialsCitation>
;
<materialsCitation id="34FC3C892226E361FCC825A0B768F832" ID-GBIF-Occurrence="2236308489" box="[848,1374,1934,1959]" collectingDate="2005-05-22" country="India" location="Bangalore" pageId="15" pageNumber="366" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FCC825A0B142F832" box="[848,884,1935,1959]" pageId="15" pageNumber="366" type="male">1♂</specimenCount>
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<location id="814B600F2226E361FC1125A1B1C9F832" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FC1125A1B1C9F832" box="[905,1023,1934,1959]" country="India" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
, GKVK,
<collectingDate id="E06EE9FC2226E361FBE225A1B6DDF832" box="[1146,1259,1934,1959]" pageId="15" pageNumber="366" value="2005-05-22">
<date id="F02A10142226E361FBE225A1B6DDF832" box="[1146,1259,1934,1959]" pageId="15" pageNumber="366" value="2005-05-22">22.v.2005</date>
</collectingDate>
, students
</materialsCitation>
;
<materialsCitation id="34FC3C892226E361FAEA25A0B051F85E" ID-GBIF-Occurrence="2236308456" collectingDate="2005-06-07" country="India" location="Bangalore" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FAEA25A0B7A3F832" box="[1394,1429,1935,1959]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
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<location id="814B600F2226E361FF0F259DB33BF85E" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FF0F259DB33BF85E" box="[151,269,1970,1995]" country="India" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
, GKVK,
<collectingDate id="E06EE9FC2226E361FEE6259DB3CFF85E" box="[382,505,1970,1995]" pageId="15" pageNumber="366" value="2005-06-07">
<date id="F02A10142226E361FEE6259DB3CFF85E" box="[382,505,1970,1995]" pageId="15" pageNumber="366" value="2005-06-07">07.vi.2005</date>
</collectingDate>
, students
</materialsCitation>
;
<materialsCitation id="34FC3C892226E361FDEF259DB6BAF85F" ID-GBIF-Occurrence="2236308478" box="[631,1164,1970,1995]" collectingDate="2005-07-16" collectorName="Salini, S" country="India" location="Bangalore" municipality="Hebbal" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FDEF259DB0ACF85F" box="[631,666,1970,1994]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
,
<collectingMunicipality id="644FACAE2226E361FD34259DB136F85E" box="[684,768,1970,1995]" pageId="15" pageNumber="366">Hebbal</collectingMunicipality>
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<location id="814B600F2226E361FC97259DB1B3F85E" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FC97259DB1B3F85E" box="[783,901,1970,1995]" country="India" municipality="Hebbal" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
,
<collectingDate id="E06EE9FC2226E361FC0C259DB623F85E" box="[916,1045,1970,1995]" pageId="15" pageNumber="366" value="2005-07-16">
<date id="F02A10142226E361FC0C259DB623F85E" box="[916,1045,1970,1995]" pageId="15" pageNumber="366" value="2005-07-16">16.vii.2005</date>
</collectingDate>
,
<collectorName id="296153022226E361FBBD259DB6BAF85F" box="[1061,1164,1970,1995]" pageId="15" pageNumber="366">Salini, S</collectorName>
</materialsCitation>
.;
<specimenCount id="9292FD5D2226E361FB3A259DB6F3F85F" box="[1186,1221,1970,1994]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
, Mudigere,
<quantity id="436C9B312226E361FACD259DB7AEF85F" box="[1365,1432,1970,1994]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="15" pageNumber="366" unit="m" value="980.0">980m</quantity>
, Rekha, M.;
<materialsCitation id="34FC3C892226E361FEB125F8B08BF87A" ID-GBIF-Occurrence="2236308462" box="[297,701,2006,2031]" collectingDate="2005-11-07" country="India" location="Mandya" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FEB125F8B37AF87A" box="[297,332,2007,2031]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
,
<location id="814B600F2226E361FEF825F9B389F87A" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FEF825F9B389F87A" box="[352,447,2006,2031]" country="India" name="Mandya" pageId="15" pageNumber="366" stateProvince="Karnataka">Mandya</location>
,
<collectingDate id="E06EE9FC2226E361FE4925F9B07AF87A" box="[465,588,2006,2031]" pageId="15" pageNumber="366" value="2005-11-07">
<date id="F02A10142226E361FE4925F9B07AF87A" box="[465,588,2006,2031]" pageId="15" pageNumber="366" value="2005-11-07">07.xi.2005</date>
</collectingDate>
, students
</materialsCitation>
;
<materialsCitation id="34FC3C892226E361FD5725F8B6EEF87A" ID-GBIF-Occurrence="2236308460" box="[719,1240,2006,2031]" collectingDate="2006-04-28" collectorName="Salini, S" country="India" elevation="916" location="Bangalore" pageId="15" pageNumber="366" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2226E361FD5725F8B0C4F87A" box="[719,754,2007,2031]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
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<location id="814B600F2226E361FC9F25F9B14BF87A" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2226E361FC9F25F9B14BF87A" box="[775,893,2006,2031]" country="India" name="Bangalore" pageId="15" pageNumber="366" stateProvince="Karnataka">Bangalore</location>
,
<elevation id="0FB9D1E72226E361FC1725F9B1E4F87A" box="[911,978,2006,2031]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="366" unit="m" value="916.0">
<quantity id="436C9B312226E361FC1725F9B1E4F87A" box="[911,978,2006,2031]" metricMagnitude="2" metricUnit="m" metricValue="9.16" pageId="15" pageNumber="366" unit="m" value="916.0">916m</quantity>
</elevation>
,
<collectingDate id="E06EE9FC2226E361FC7B25F9B66DF87A" box="[995,1115,2006,2031]" pageId="15" pageNumber="366" value="2006-04-28">
<date id="F02A10142226E361FC7B25F9B66DF87A" box="[995,1115,2006,2031]" pageId="15" pageNumber="366" value="2006-04-28">28.iv.2006</date>
</collectingDate>
,
<collectorName id="296153022226E361FBF625F9B6EEF87A" box="[1134,1240,2006,2031]" pageId="15" pageNumber="366">Salini, S</collectorName>
</materialsCitation>
.;
<specimenCount id="9292FD5D2226E361FB6925F8B722F87A" box="[1265,1300,2007,2031]" pageId="15" pageNumber="366" type="female">1♀</specimenCount>
, Mudigere, Naveen, K. S.;
<materialsCitation id="34FC3C892239E37EFED222B7B16BFF25" ID-GBIF-Occurrence="2236308452" box="[330,861,151,176]" collectingDate="2008-02-28" collectorName="Kalleshwaraswamy" country="India" location="Mudigere" pageId="16" pageNumber="367" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2239E37EFED222B7B358FF25" box="[330,366,152,176]" pageId="16" pageNumber="367" type="male">1♂</specimenCount>
,
<location id="814B600F2239E37EFEE622B8B3D8FF25" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2239E37EFEE622B8B3D8FF25" box="[382,494,151,176]" country="India" name="Mudigere" pageId="16" pageNumber="367" stateProvince="Karnataka">Mudigere</location>
,
<collectingDate id="E06EE9FC2239E37EFE6322B8B059FF25" box="[507,623,151,176]" pageId="16" pageNumber="367" value="2008-02-28">
<date id="F02A10142239E37EFE6322B8B059FF25" box="[507,623,151,176]" pageId="16" pageNumber="367" value="2008-02-28">28.ii.2008</date>
</collectingDate>
,
<collectorName id="296153022239E37EFDE622B8B16BFF25" box="[638,861,151,176]" pageId="16" pageNumber="367">Kalleshwaraswamy</collectorName>
</materialsCitation>
;
<materialsCitation id="34FC3C892239E37EFCF222B7B71CFF25" ID-GBIF-Occurrence="2236308471" box="[874,1322,151,176]" collectingDate="2008-09-11" collectingDateMax="2008-09-12" collectingDateMin="2008-09-11" collectorName="Murthy, S" country="India" location="Raichur" pageId="16" pageNumber="367" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2239E37EFCF222B7B1BBFF25" box="[874,909,152,176]" pageId="16" pageNumber="367" type="female">1♀</specimenCount>
,
<location id="814B600F2239E37EFC0622B8B1CEFF25" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2239E37EFC0622B8B1CEFF25" box="[926,1016,151,176]" country="India" name="Raichur" pageId="16" pageNumber="367" stateProvince="Karnataka">Raichur</location>
,
<collectingDate id="E06EE9FC2239E37EFB9E22B8B693FF25" box="[1030,1189,151,176]" pageId="16" pageNumber="367" value="2008-09-11" valueMax="2008-09-12" valueMin="2008-09-11">
<date id="F02A10142239E37EFB9E22B8B693FF25" box="[1030,1189,151,176]" pageId="16" pageNumber="367" value="2008-09-11" valueMax="2008-09-12" valueMin="2008-09-11">11-12.ix.2008</date>
</collectingDate>
,
<collectorName id="296153022239E37EFB2B22B8B71CFF25" box="[1203,1322,151,176]" pageId="16" pageNumber="367">Murthy, S</collectorName>
</materialsCitation>
.;
<specimenCount id="9292FD5D2239E37EFAA622B7B755FF25" box="[1342,1379,152,176]" pageId="16" pageNumber="367" type="male">9♂</specimenCount>
;
<materialsCitation id="34FC3C892239E37EFAEA22B7B162FF42" ID-GBIF-Occurrence="2236308461" collectingDate="2012-05-01" country="India" location="Hebbal" municipality="Bangalore" pageId="16" pageNumber="367" specimenCount="2" specimenCount-female="2" stateProvince="Karnataka">
<specimenCount id="9292FD5D2239E37EFAEA22B7B7A1FF25" box="[1394,1431,152,176]" pageId="16" pageNumber="367" type="female">2♀</specimenCount>
,
<collectingMunicipality id="644FACAE2239E37EFF0F2291B33BFF42" box="[151,269,190,215]" pageId="16" pageNumber="367">Bangalore</collectingMunicipality>
,
<location id="814B600F2239E37EFE842291B346FF42" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2239E37EFE842291B346FF42" box="[284,368,190,215]" country="India" municipality="Bangalore" name="Hebbal" pageId="16" pageNumber="367" stateProvince="Karnataka">Hebbal</location>
,
<collectingDate id="E06EE9FC2239E37EFEE72290B3C6FF42" box="[383,496,191,215]" pageId="16" pageNumber="367" value="2012-05-01">
<date id="F02A10142239E37EFEE72290B3C6FF42" box="[383,496,191,215]" pageId="16" pageNumber="367" value="2012-05-01">01.v.2012</date>
</collectingDate>
, ex.
<taxonomicName id="43944D572239E37EFDB32291B162FF42" authority=", Manjunath, B" authorityName="Manjunath, B" box="[555,852,190,215]" class="Magnoliopsida" family="Bignoniaceae" genus="Spathodea" kingdom="Plantae" order="Lamiales" pageId="16" pageNumber="367" phylum="Tracheophyta" rank="genus">
<emphasis id="B6E0EAC62239E37EFDB32291B097FF42" box="[555,673,190,215]" italics="true" pageId="16" pageNumber="367">Spathodea</emphasis>
, Manjunath, B
</taxonomicName>
</materialsCitation>
;
<materialsCitation id="34FC3C892239E37EFCFC2290B3D4FF6F" ID-GBIF-Occurrence="2236308467" collectingDate="2012-02-04" collectorName="Yeshwanth, H. M" country="India" elevation="930" latitude="12.966666" location="Bangalore" pageId="16" pageNumber="367" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2239E37EFCFC2290B1B1FF42" box="[868,903,191,215]" pageId="16" pageNumber="367" type="female">1♀</specimenCount>
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<location id="814B600F2239E37EFC012291B639FF42" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2239E37EFC012291B639FF42" box="[921,1039,190,215]" country="India" latitude="12.966666" name="Bangalore" pageId="16" pageNumber="367" stateProvince="Karnataka">Bangalore</location>
, GKVK,
<elevation id="0FB9D1E72239E37EFB192290B6F2FF42" box="[1153,1220,191,215]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="16" pageNumber="367" unit="m" value="930.0">
<quantity id="436C9B312239E37EFB192290B6F2FF42" box="[1153,1220,191,215]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="16" pageNumber="367" unit="m" value="930.0">930m</quantity>
</elevation>
,
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58N
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77
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85E,
<collectingDate id="E06EE9FC2239E37EFF0F22CDB33DFF6E" box="[151,267,226,251]" pageId="16" pageNumber="367" value="2012-02-04">
<date id="F02A10142239E37EFF0F22CDB33DFF6E" box="[151,267,226,251]" pageId="16" pageNumber="367" value="2012-02-04">04.ii.2012</date>
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,
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</materialsCitation>
.;
<materialsCitation id="34FC3C892239E37EFE6A22CCB679FF6F" ID-GBIF-Occurrence="2236308483" box="[498,1103,226,251]" collectingDate="2014-02-28" collectorName="Yeshwanth, H. M" country="India" location="Bangalore" pageId="16" pageNumber="367" specimenCount="1" specimenCount-female="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2239E37EFE6A22CCB023FF6E" box="[498,533,227,251]" pageId="16" pageNumber="367" type="female">1♀</specimenCount>
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<location id="814B600F2239E37EFDBB22CDB0AFFF6E" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2239E37EFDBB22CDB0AFFF6E" box="[547,665,226,251]" country="India" name="Bangalore" pageId="16" pageNumber="367" stateProvince="Karnataka">Bangalore</location>
, GKVK,
<collectingDate id="E06EE9FC2239E37EFC9C22CDB14EFF6E" box="[772,888,226,251]" pageId="16" pageNumber="367" value="2014-02-28">
<date id="F02A10142239E37EFC9C22CDB14EFF6E" box="[772,888,226,251]" pageId="16" pageNumber="367" value="2014-02-28">28.ii.2014</date>
</collectingDate>
,
<collectorName id="296153022239E37EFC1C22CDB679FF6F" box="[900,1103,226,251]" pageId="16" pageNumber="367">Yeshwanth, H. M</collectorName>
</materialsCitation>
.;
<specimenCount id="9292FD5D2239E37EFBC722CCB6B2FF6E" box="[1119,1156,227,251]" pageId="16" pageNumber="367" type="female">1♀</specimenCount>
;
<materialsCitation id="34FC3C892239E37EFB0A22CCB15DFEB7" ID-GBIF-Occurrence="2236308490" collectingDate="2014-03-07" collectorName="Murraya &amp; Salini, S" country="India" elevation="930" latitude="12.966666" location="Bangalore" longLatPrecision="1291" longitude="77.583336" pageId="16" pageNumber="367" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2239E37EFB0A22CCB680FF6E" box="[1170,1206,227,251]" pageId="16" pageNumber="367" type="male">1♂</specimenCount>
,
<location id="814B600F2239E37EFB5B22CDB70FFF6E" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2239E37EFB5B22CDB70FFF6E" box="[1219,1337,226,251]" country="India" latitude="12.966666" longLatPrecision="1291" longitude="77.583336" name="Bangalore" pageId="16" pageNumber="367" stateProvince="Karnataka">Bangalore</location>
, GKVK,
<elevation id="0FB9D1E72239E37EFF0F2325B2EFFEB7" box="[151,217,266,290]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="16" pageNumber="367" unit="m" value="930.0">
<quantity id="436C9B312239E37EFF0F2325B2EFFEB7" box="[151,217,266,290]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="16" pageNumber="367" unit="m" value="930.0">930m</quantity>
</elevation>
,
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58N
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,
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35E
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,
<collectingDate id="E06EE9FC2239E37EFE262326B00CFEB7" box="[446,570,265,290]" pageId="16" pageNumber="367" value="2014-03-07">
<date id="F02A10142239E37EFE262326B00CFEB7" box="[446,570,265,290]" pageId="16" pageNumber="367" value="2014-03-07">07.iii.2014</date>
</collectingDate>
,
<emphasis id="B6E0EAC62239E37EFDDF2325B0FBFEB4" box="[583,717,266,289]" italics="true" pageId="16" pageNumber="367">
ex
<collectorName id="296153022239E37EFDF12325B0FBFEB4" box="[617,717,266,289]" pageId="16" pageNumber="367">Murraya</collectorName>
</emphasis>
sp.,
<collectorName id="296153022239E37EFC9D2326B15DFEB7" box="[773,875,265,290]" pageId="16" pageNumber="367">Salini, S</collectorName>
</materialsCitation>
.;
<materialsCitation id="34FC3C892239E37EFCE62325B7A4FEB7" ID-GBIF-Occurrence="2236308463" box="[894,1426,265,290]" collectingDate="2014-08-16" collectorName="Salini, S" country="India" location="Hebbal" municipality="Bangalore" pageId="16" pageNumber="367" specimenCount="1" specimenCount-male="1" stateProvince="Karnataka">
<specimenCount id="9292FD5D2239E37EFCE62325B194FEB7" box="[894,930,266,290]" pageId="16" pageNumber="367" type="male">1♂</specimenCount>
,
<collectingMunicipality id="644FACAE2239E37EFC292326B611FEB7" box="[945,1063,265,290]" pageId="16" pageNumber="367">Bangalore</collectingMunicipality>
,
<location id="814B600F2239E37EFBAC2326B6BEFEB7" LSID="urn:lsid:plazi:treatment:0C3D87C22224E37EFF0F276AB760FE1B:814B600F2239E37EFBAC2326B6BEFEB7" box="[1076,1160,265,290]" country="India" municipality="Bangalore" name="Hebbal" pageId="16" pageNumber="367" stateProvince="Karnataka">Hebbal</location>
,
<collectingDate id="E06EE9FC2239E37EFB0D2326B729FEB7" box="[1173,1311,265,290]" pageId="16" pageNumber="367" value="2014-08-16">
<date id="F02A10142239E37EFB0D2326B729FEB7" box="[1173,1311,265,290]" pageId="16" pageNumber="367" value="2014-08-16">16.viii.2014</date>
</collectingDate>
,
<collectorName id="296153022239E37EFAB42326B7A4FEB7" box="[1324,1426,265,290]" pageId="16" pageNumber="367">Salini, S</collectorName>
</materialsCitation>
.;
<materialsCitation id="34FC3C892239E37EFF0F2301B01CFED3" ID-GBIF-Occurrence="2236308494" box="[151,554,301,326]" collectorName="Mudigere &amp; Kantha Raj." country="India" location="Kerala" pageId="16" pageNumber="367" specimenCount="1" specimenCount-male="1" stateProvince="Kerala">
<specimenCount id="9292FD5D2239E37EFF0F2301B28DFED3" box="[151,187,302,326]" pageId="16" pageNumber="367" type="male">1♂</specimenCount>
,
<collectorName id="296153022239E37EFF502302B30EFED3" box="[200,312,301,326]" pageId="16" pageNumber="367">Mudigere</collectorName>
,
<collectorName id="296153022239E37EFEDB2302B3FAFED3" box="[323,460,301,326]" pageId="16" pageNumber="367">Kantha Raj.</collectorName>
;
<collectingRegion id="4650F8362239E37EFE4F2302B01CFED3" box="[471,554,301,326]" country="India" name="Kerala" pageId="16" pageNumber="367">
<emphasis id="B6E0EAC62239E37EFE4F2302B01CFED3" bold="true" box="[471,554,301,326]" pageId="16" pageNumber="367">Kerala</emphasis>
</collectingRegion>
</materialsCitation>
:
<specimenCount id="9292FD5D2239E37EFDA02301B06BFED3" box="[568,605,302,326]" pageId="16" pageNumber="367" type="male">1♂</specimenCount>
, Thrissur, KAU, Vellanikkara, Students collection.
</paragraph>
</subSubSection>
<subSubSection id="CC8E655F2239E37EFF5F237EB760FE1B" pageId="16" pageNumber="367" type="discussion">
<paragraph id="842B36D42239E37EFF5F237EB760FE1B" blockId="16.[151,1437,151,398]" pageId="16" pageNumber="367">
<emphasis id="B6E0EAC62239E37EFF5F237EB30DFEFF" bold="true" box="[199,315,337,362]" pageId="16" pageNumber="367">Remarks.</emphasis>
This species was originally described by
<bibRefCitation id="E0054B252239E37EFCB9237EB1F5FEFF" author="Ghauri, M. S. K." box="[801,963,337,362]" pageId="16" pageNumber="367" pagination="77 - 92" refId="ref13220" refString="Ghauri, M. S. K. (1988) A revision of Asian species of the genus Halys Fabricius based on the type material (Insecta, Heteroptera, Pentatomidae, Pentatomidae). Entomologische Abhandlungen Staatliches Museum fur Tierkunde Dresden, 51 (6), 77 - 92." type="journal article" year="1988">Ghauri (1988)</bibRefCitation>
from Coonoor,
<collectingRegion id="4650F8362239E37EFBE7237EB73FFEFF" box="[1151,1289,337,362]" country="India" name="Tamil Nadu" pageId="16" pageNumber="367">Tamil Nadu</collectingRegion>
, and has not been reported outside of
<collectingRegion id="4650F8362239E37EFE35235AB004FE1B" box="[429,562,373,398]" country="India" name="Tamil Nadu" pageId="16" pageNumber="367">Tamil Nadu</collectingRegion>
so far. This species is recorded for the first from
<collectingRegion id="4650F8362239E37EFBC2235AB6FBFE1B" box="[1114,1229,373,398]" country="India" name="Karnataka" pageId="16" pageNumber="367">Karnataka</collectingRegion>
and
<collectingRegion id="4650F8362239E37EFA9B235AB767FE1B" box="[1283,1361,373,398]" country="India" name="Kerala" pageId="16" pageNumber="367">Kerala</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>