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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.727.20222" ID-GBIF-Dataset="13853cb6-522e-4f94-b633-c4a51ba63624" ID-PMC="PMC5799789" ID-Pensoft-Pub="1313-2970-727-1" ID-PubMed="29416388" ID-ZBK="12B663F7190040788E1EEF8BAC4DF81B" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-727-1" ModsDocOrigin="ZooKeys 727" ModsDocTitle="A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)" checkinTime="1515696595036" checkinUser="pensoft" docAuthor="Wood, Hannah M. &amp; Scharff, Nikolaj" docDate="2017" docId="0C178F9C420EEC6A9E2DD6EEE954BF6D" docLanguage="en" docName="ZooKeys 727: 1-96" docOrigin="ZooKeys 727" docSource="http://dx.doi.org/10.3897/zookeys.727.20222" docTitle="Madagascarchaea Wood &amp; Scharff, 2017, gen. n." docType="treatment" docUuid="041CF907-A633-440F-A0DE-EA875E1265DF" docUuidSource="ZooBank" docVersion="5" lastPageNumber="1" masterDocId="FFEBD959ED2FFFAA9450FF8B1957B348" masterDocTitle="A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="1" updateTime="1668165276717" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A review of the Madagascan pelican spiders of the genera Eriauchenius O. Pickard-Cambridge, 1881 and Madagascarchaea gen. n. (Araneae, Archaeidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wood, Hannah M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Scharff, Nikolaj</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>727</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>96</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.727.20222</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.727.20222</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-727-1</mods:identifier>
<mods:identifier type="ZBK">12B663F7190040788E1EEF8BAC4DF81B</mods:identifier>
<mods:identifier type="ZooBank">12B663F7190040788E1EEF8BAC4DF81B</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="140245904" LSID="urn:lsid:zoobank.org:act:041CF907-A633-440F-A0DE-EA875E1265DF" httpUri="http://treatment.plazi.org/id/0C178F9C420EEC6A9E2DD6EEE954BF6D" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
Genus
<taxonomicName LSID="http://zoobank.org/041CF907-A633-440F-A0DE-EA875E1265DF" class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Madagascarchaea</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">gen. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type species">
<paragraph pageId="0" pageNumber="1">Type species.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName family="Archaeidae" genus="Archaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Archaea gracilicollis" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="gracilicollis">Archaea gracilicollis</taxonomicName>
Millot, 1948
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">The name refers to the Madagascan distribution and is feminine in gender.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Distinguished from all other archaeids by the presence of six protrusions, each with a small spine, on the crown of the cephalon, and by the presence of a retrolateral apophysis on the male pedipalpal patella.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
See
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood (2008)</bibRefCitation>
for a complete description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="included species">
<paragraph pageId="0" pageNumber="1">Included species.</paragraph>
<paragraph pageId="0" pageNumber="1">
14 described species
<taxonomicName lsidName="M. ambre" pageId="0" pageNumber="1" rank="species" species="ambre">M. ambre</taxonomicName>
(Wood, 2008),
<taxonomicName lsidName="M. anabohazo" pageId="0" pageNumber="1" rank="species" species="anabohazo">M. anabohazo</taxonomicName>
(Wood, 2008),
<taxonomicName lsidName="M. borimontsina" pageId="0" pageNumber="1" rank="species" species="borimontsina">M. borimontsina</taxonomicName>
(Wood, 2008),
<taxonomicName lsidName="M. gracilicollis" pageId="0" pageNumber="1" rank="species" species="gracilicollis">M. gracilicollis</taxonomicName>
(Millot, 1948),
<taxonomicName lsidName="M. griswoldi" pageId="0" pageNumber="1" rank="species" species="griswoldi">M. griswoldi</taxonomicName>
(Wood, 2008),
<taxonomicName lsidName="M. halambohitra" pageId="0" pageNumber="1" rank="species" species="halambohitra">M. halambohitra</taxonomicName>
(Wood, 2008),
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
(Millot, 1948),
<taxonomicName lsidName="M. lavatenda" pageId="0" pageNumber="1" rank="species" species="lavatenda">M. lavatenda</taxonomicName>
(Wood, 2008),
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
(Platnick, 1991),
<taxonomicName lsidName="M. namoroka" pageId="0" pageNumber="1" rank="species" species="namoroka">M. namoroka</taxonomicName>
(Wood, 2008),
<taxonomicName lsidName="M. spiceri" pageId="0" pageNumber="1" rank="species" species="spiceri">M. spiceri</taxonomicName>
(Wood, 2008),
<taxonomicName lsidName="M. tsingyensis" pageId="0" pageNumber="1" rank="species" species="tsingyensis">M. tsingyensis</taxonomicName>
(Lotz, 2003),
<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
(Millot, 1948),
<taxonomicName lsidName="M. voronakely" pageId="0" pageNumber="1" rank="species" species="voronakely">M. voronakely</taxonomicName>
(Wood, 2008), and 4 new species described here:
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea fohy" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="fohy">Madagascarchaea fohy</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea lotzi" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="lotzi">Madagascarchaea lotzi</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea moramora" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="moramora">Madagascarchaea moramora</taxonomicName>
sp. n.,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea rabesahala" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="rabesahala">Madagascarchaea rabesahala</taxonomicName>
sp. n
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Madagascar</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
We elevate the &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea gracilicollis" pageId="0" pageNumber="1" rank="species" species="gracilicollis">gracilicollis</taxonomicName>
group&quot; (
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
) to genus level based on phylogenetic analysis that strongly supports two monophyletic clades,
<taxonomicName class="Arachnida" family="Archaeidae" genus="Eriauchenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriauchenius" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Eriauchenius</taxonomicName>
and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Madagascarchaea</taxonomicName>
gen. n., that have diversified on Madagascar (Fig. 1) (
<bibRefCitation author="Wood, HM" journalOrPublisher="Evolution" pageId="0" pageNumber="1" pagination="462 - 481" title="Why is Madagascar special? The extraordinarily slow evolution of pelican spiders (Araneae, Archaeidae)." url="https://doi.org/10.1111/evo.12578" volume="69" year="2015">Wood et al. 2015</bibRefCitation>
;
<bibRefCitation author="Wood, HM" journalOrPublisher="Cladistics" pageId="0" pageNumber="1" pagination="598 - 626" title="Phylogenetic placement of pelican spiders (Archaeidae, Araneae), with insight into evolution of the &quot; neck &quot; and predatory behaviours of the superfamily Palpimanoidea." url="https://doi.org/10.1111/j.1096-0031.2012.00411.x" volume="28" year="2012">Wood et al. 2012</bibRefCitation>
). These two genera are not sister clades, but instead
<taxonomicName class="Arachnida" family="Archaeidae" genus="Afrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afrarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Afrarchaea</taxonomicName>
is sister to
<taxonomicName class="Arachnida" family="Archaeidae" genus="Eriauchenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriauchenius" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Eriauchenius</taxonomicName>
in
<bibRefCitation author="Wood, HM" journalOrPublisher="Evolution" pageId="0" pageNumber="1" pagination="462 - 481" title="Why is Madagascar special? The extraordinarily slow evolution of pelican spiders (Araneae, Archaeidae)." url="https://doi.org/10.1111/evo.12578" volume="69" year="2015">Wood et al (2015)</bibRefCitation>
, and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Afrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afrarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Afrarchaea</taxonomicName>
is sister to
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Madagascarchaea</taxonomicName>
gen. n. in
<bibRefCitation author="Wood, HM" journalOrPublisher="Cladistics" pageId="0" pageNumber="1" pagination="598 - 626" title="Phylogenetic placement of pelican spiders (Archaeidae, Araneae), with insight into evolution of the &quot; neck &quot; and predatory behaviours of the superfamily Palpimanoidea." url="https://doi.org/10.1111/j.1096-0031.2012.00411.x" volume="28" year="2012">Wood et al (2012)</bibRefCitation>
.
<taxonomicName class="Arachnida" family="Archaeidae" genus="Eriauchenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eriauchenius" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Eriauchenius</taxonomicName>
and
<taxonomicName class="Arachnida" family="Archaeidae" genus="Afrarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Afrarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Afrarchaea</taxonomicName>
are never recovered as sister clades.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Madagascarchaea</taxonomicName>
gen. n. contains two main clades, the &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
group&quot; and the &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">jeanneli</taxonomicName>
group&quot; (Fig. 1). The &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
group&quot; contains
<taxonomicName lsidName="M. borimontsina" pageId="0" pageNumber="1" rank="species" species="borimontsina">M. borimontsina</taxonomicName>
,
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
,
<taxonomicName lsidName="M. vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">M. vadoni</taxonomicName>
,
<taxonomicName lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
sp. n., and
<taxonomicName lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
sp. n., and these species share the following traits: a retrolateral apophysis on the male pedipalpal femur (compare fig. 12A with fig. 12B in
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
) and a highly reduced FSGP that lacks
<normalizedToken originalValue="“wings”">&quot;wings&quot;</normalizedToken>
(Figs 23B, 24B, 25B). The female genitalia of &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
group&quot; species do not have interspecific variation. Instead, somatic traits are more useful for identifying &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea vadoni" pageId="0" pageNumber="1" rank="species" species="vadoni">vadoni</taxonomicName>
group&quot; species. We split what was previously considered
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
into two species:
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
and
<taxonomicName lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
sp. n. For clarity, we describe both
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
and
<taxonomicName lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
sp. n. here. In addition, there is a single female specimen (CASENT9046596), denoted as &quot;
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Madagascarchaea</taxonomicName>
sp. 1&quot; in Fig. 1 that may be a new species, however, since this is the only known specimen and females are difficult to diagnose, we chose not to describe it (collection information: Madagascar, Fianarantsoa,
<normalizedToken originalValue="Réserve">Reserve</normalizedToken>
<normalizedToken originalValue="Forestiére">Forestiere</normalizedToken>
<normalizedToken originalValue="d’Agnalazaha">d'Agnalazaha</normalizedToken>
, Mahabo, 42.9 km 215° SW Farafangana,
<geoCoordinate direction="south" orientation="latitude" precision="15" value="-23.193888">23°11'38&quot;S</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="15" value="47.723057">47°43'23&quot;E</geoCoordinate>
, 20 m, 19 Apr 2006, littoral rainforest, maxi winkler litter extraction, B.L. Fisher et al.).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
,
<taxonomicName lsidName="M. ambre" pageId="0" pageNumber="1" rank="species" species="ambre">M. ambre</taxonomicName>
,
<taxonomicName lsidName="M. lotzi" pageId="0" pageNumber="1" rank="species" species="lotzi">M. lotzi</taxonomicName>
sp. n., and
<taxonomicName lsidName="M. moramora" pageId="0" pageNumber="1" rank="species" species="moramora">M. moramora</taxonomicName>
sp. n. belong to the &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">jeanneli</taxonomicName>
group&quot; and are part of a species complex distinguished from other
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Madagascarchaea</taxonomicName>
by having an elongated and pointy head (Figs 26A, 27A, 28A), a triangular abdomen that is either straight across the posterior (Fig. 28A) or invaginated (Figs 26A, 27A), and a conductor that is a large triangular piece in the prolateral view (Figs 26D, 27D). The current study splits the
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
of
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood (2008)</bibRefCitation>
into several different species:
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
,
<taxonomicName lsidName="M. lotzi" pageId="0" pageNumber="1" rank="species" species="lotzi">M. lotzi</taxonomicName>
sp. n., and
<taxonomicName lsidName="M. moramora" pageId="0" pageNumber="1" rank="species" species="moramora">M. moramora</taxonomicName>
sp. n. For clarity, we also redescribe
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
here.
<taxonomicName lsidName="M. ambre" pageId="0" pageNumber="1" rank="species" species="ambre">M. ambre</taxonomicName>
, is considered a part of the &quot;
<taxonomicName genus="Palpimanoidea" lsidName="Palpimanoidea jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">jeanneli</taxonomicName>
-complex&quot; but is not redescribed here, see
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood (2008)</bibRefCitation>
for a complete description. These four species form a monophyletic group (Fig. 1), although
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
is not included in the phylogeny. Females cannot be distinguished. For the males, embolus shape is the main morphological difference among species.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="madagascarchaea gen. n.: to supplement “gracilicollis group” key">
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Arachnida" family="Archaeidae" genus="Madagascarchaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Madagascarchaea" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Madagascarchaea</taxonomicName>
gen. n.: to supplement &quot;Gracilicollis group&quot; key
</paragraph>
<paragraph pageId="0" pageNumber="1">
When using the identification key from
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood (2008)</bibRefCitation>
, if a specimen is identified as either
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
or
<taxonomicName lsidName="M. ambre" pageId="0" pageNumber="1" rank="species" species="ambre">M. ambre</taxonomicName>
, then use the following key to separate
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
,
<taxonomicName lsidName="M. ambre" pageId="0" pageNumber="1" rank="species" species="ambre">M. ambre</taxonomicName>
,
<taxonomicName lsidName="M. lotzi" pageId="0" pageNumber="1" rank="species" species="lotzi">M. lotzi</taxonomicName>
sp. n., and
<taxonomicName lsidName="M. moramora" pageId="0" pageNumber="1" rank="species" species="moramora">M. moramora</taxonomicName>
sp. n.; females of these four species are indistinguishable:
</paragraph>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
27
<normalizedToken originalValue="E–F">E-F</normalizedToken>
,
<normalizedToken originalValue="G–H">G-H</normalizedToken>
27H
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="M. lotzi" pageId="0" pageNumber="1" rank="species" species="lotzi">M. lotzi</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
26F, I, L28
<normalizedToken originalValue="E–F">E-F</normalizedToken>
28A
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
28A
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood 2008</bibRefCitation>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="M. ambre" pageId="0" pageNumber="1" rank="species" species="ambre">M. ambre</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
26
<normalizedToken originalValue="E–F,H–I">E-F,H-I</normalizedToken>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
28
<normalizedToken originalValue="D–F28D–F">D-F28D-F</normalizedToken>
28A
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="M. moramora" pageId="0" pageNumber="1" rank="species" species="moramora">M. moramora</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
26
<normalizedToken originalValue="D–L26E–F">D-L26E-F</normalizedToken>
,
<normalizedToken originalValue="H–I">H-I</normalizedToken>
26A
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="M. jeanneli" pageId="0" pageNumber="1" rank="species" species="jeanneli">M. jeanneli</taxonomicName>
</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="1">
When using the identification key from
<bibRefCitation author="Wood, H" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="255 - 296" title="A revision of the assassin spiders of the Eriaucheniusgracilicollis group, a clade of spiders endemic to Madagascar (Araneae: Archaeidae)." url="https://doi.org/10.1111/j.1096-3642.2007.00359.x" volume="152" year="2008">Wood (2008)</bibRefCitation>
, if a specimen is identified as
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
, then use the following key to separate species of
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
,
<taxonomicName lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
sp. n., and
<taxonomicName lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
sp. n.:
</paragraph>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">23A24A</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">25A</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="M. rabesahala" pageId="0" pageNumber="1" rank="species" species="rabesahala">M. rabesahala</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
23H, J23C
<taxonomicName lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="M. fohy" pageId="0" pageNumber="1" rank="species" species="fohy">M. fohy</taxonomicName>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
24
<normalizedToken originalValue="G–I25G–I">G-I25G-I</normalizedToken>
24C
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName lsidName="M. legendrei" pageId="0" pageNumber="1" rank="species" species="legendrei">M. legendrei</taxonomicName>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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