375 lines
56 KiB
XML
375 lines
56 KiB
XML
<document id="FFEBFC98547216930CAE514C879FFDA8" ID-CLB-Dataset="24150" ID-DOI="10.5252/zoosystema2020v42a3" ID-GBIF-Dataset="7bdbb02c-ce5d-4df5-846a-b45c54fedc5a" ID-Zenodo-Dep="3652791" ID-ZooBank="urn:lsid:zoobank.org:pub:27C94F0F-BB9B-40A3-B615-4DB19C94F042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581020160253" checkinUser="plazi" docAuthor="Kantor, Yuri, Kosyan, Alisa & Sorokin, Pavel" docDate="2020" docId="03D4B94FFFDE203FFF35FE55FDADC375" docLanguage="en" docName="zoosystema2020v42a3.pdf" docOrigin="Zoosystema 42 (3)" docStyle="DocumentStyle:5CC576F744449C1860B4E72ABD884631.7:Zoosystema.2020-.journal_article" docStyleId="5CC576F744449C1860B4E72ABD884631" docStyleName="Zoosystema.2020-.journal_article" docStyleVersion="7" docTitle="Siphonalia pfefferi G. B. Sowerby III 1900" docType="treatment" docVersion="8" lastPageNumber="52" masterDocId="FFEDC137FFCB2028FFEFFFCDFFA9C206" masterDocTitle="On the taxonomic position of Phaenomenella Fraussen & Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species" masterLastPageNumber="55" masterPageNumber="33" pageNumber="52" updateTime="1698780820290" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8EC4700B70E6AE0558F3FAAFDF7E34D0">On the taxonomic position of Phaenomenella Fraussen & Hadorn, 2006 (Neogastropoda, Buccinoidea) with description of two new species</mods:title>
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<mods:namePart id="2637432EA1A8BF6AB7B384AB4EA26901">Kantor, Yuri</mods:namePart>
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<mods:namePart id="26CC1620D61968AD7D1C06A4FB789E78">Kosyan, Alisa</mods:namePart>
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<mods:name id="D3A137F8E1B3EF5CAD39F15298F51B10" type="personal">
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<mods:namePart id="1FDB9FEC57DDEA2F9CA104F6B3353645">Sorokin, Pavel</mods:namePart>
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<mods:typeOfResource id="60AC47FD7090311FEA7B9B8933798040">text</mods:typeOfResource>
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<mods:title id="85D79B6016B251E7DA1E9F2A705E61A8">Zoosystema</mods:title>
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<mods:part id="43B707FFD9FC71A0C279004446E7EA0A">
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<mods:date id="FB8560FDA35F7BBF906C60062506783F">2020</mods:date>
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<mods:number id="1EEF60A737A22C422615F0A1728DBF15">2020-02-04</mods:number>
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<mods:number id="028359D5DA4EC57EA177C4EADA1D3C25">42</mods:number>
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<treatment id="03D4B94FFFDE203FFF35FE55FDADC375" ID-DOI="http://doi.org/10.5281/zenodo.3665215" ID-GBIF-Taxon="162054094" ID-Zenodo-Dep="3665215" LSID="urn:lsid:plazi:treatment:03D4B94FFFDE203FFF35FE55FDADC375" httpUri="http://treatment.plazi.org/id/03D4B94FFFDE203FFF35FE55FDADC375" lastPageId="23" lastPageNumber="52" pageId="21" pageNumber="52">
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<subSubSection id="C3675BD2FFDE203DFF35FE55FD18C3B5" box="[218,689,408,435]" pageId="21" pageNumber="52" type="nomenclature">
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<paragraph id="8BC20859FFDE203DFF35FE55FD18C3B5" blockId="21.[218,689,408,467]" box="[218,689,408,435]" pageId="21" pageNumber="52">
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<heading id="D08ABF35FFDE203DFF35FE55FD18C3B5" box="[218,689,408,435]" centered="true" fontSize="11" level="4" pageId="21" pageNumber="52" reason="4">
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<taxonomicName id="4C7D73DAFFDE203DFF35FE55FD18C3B5" ID-CoL="6YQSF" authority="G. B. Sowerby III, 1900" authorityName="G. B. Sowerby III" authorityYear="1900" box="[218,689,408,435]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="species" species="pfefferi">
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<emphasis id="B909D44BFFDE203DFF35FE55FE0AC3B4" bold="true" box="[218,419,408,435]" italics="true" pageId="21" pageNumber="52">Siphonalia pfefferi</emphasis>
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G. B.
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<bibRefCitation id="EFEC75A8FFDE203DFE03FE55FD18C3B5" author="SOWERBY III G. B." box="[492,689,408,435]" journalOrPublisher="Annals and Magazine of Natural History" pageId="21" pageNumber="52" pagination="439 - 441" part="5" refId="ref13378" refString="SOWERBY III G. B. 1900. - New species of Mollusca of the genera Voluta, Conus, Siphonalia, and Euthria. Annals and Magazine of Natural History series 7 5: 439 - 441." title="New species of Mollusca of the genera Voluta, Conus, Siphonalia, and Euthria" type="journal article" year="1900">Sowerby III, 1900</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3675BD2FFDE203DFED2FE75FDE7C3D5" box="[317,590,440,467]" pageId="21" pageNumber="52" type="description">
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<paragraph id="8BC20859FFDE203DFED2FE75FDE7C3D5" blockId="21.[218,689,408,467]" box="[317,590,440,467]" pageId="21" pageNumber="52">
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<heading id="D08ABF35FFDE203DFED2FE75FDE7C3D5" box="[317,590,440,467]" centered="true" fontSize="11" level="4" pageId="21" pageNumber="52" reason="1">
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(
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<figureCitation id="134614DCFFDE203DFEABFE75FE13C3D5" box="[324,442,440,467]" captionStart="FIG" captionStartId="18.[132,143,1982,2000]" captionTargetBox="[132,1455,216,1829]" captionTargetId="figure@18.[132,1455,216,1829]" captionTargetPageId="18" captionText="FIG. 9. — Shells of studied Siphonalia spp.: A-C, S. cassidariaeformis (Reeve, 1846): specimen no. 1 (A), specimen no. 2 (B), specimen no. 3 (C); D, E, S. pfefferi G. B. Sowerby III, 1900; D, specimen no. 1; E, specimen no. 2. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.3652831" httpUri="https://zenodo.org/record/3652831/files/figure.png" pageId="21" pageNumber="52">Figs 9D, E</figureCitation>
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;
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<figureCitation id="134614DCFFDE203DFE27FE74FDB2C3D5" box="[456,539,441,467]" captionStart="FIG" captionStartId="20.[132,143,1406,1424]" captionTargetBox="[132,1455,215,1359]" captionTargetId="figure@20.[132,1455,214,1359]" captionTargetPageId="20" captionText="FIG. 11. — Radulae of Siphonalia spp. A, S. cassidariaeformis (Reeve, 1846) no. 1; B, S. cassidariaeformis no. 2; C, S. pfefferi G. B. Sowerby III, 1900 no. 1;" figureDoi="http://doi.org/10.5281/zenodo.3652843" httpUri="https://zenodo.org/record/3652843/files/figure.png" pageId="21" pageNumber="52">11C, D</figureCitation>
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;
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<figureCitation id="134614DCFFDE203DFDC6FE74FDECC3D5" box="[553,581,441,467]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">12</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3675BD2FFDE203DFF6BFE3CFD48C00F" box="[132,737,497,521]" pageId="21" pageNumber="52" type="reference_group">
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<paragraph id="8BC20859FFDE203DFF6BFE3CFD48C00F" blockId="21.[132,737,497,521]" box="[132,737,497,521]" pageId="21" pageNumber="52">
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<taxonomicName id="4C7D73DAFFDE203DFF6BFE3CFDF1C00F" ID-CoL="6YQSF" authority="G. B. Sowerby III, 1900: 440" authorityName="G. B. Sowerby III" authorityYear="1900" box="[132,600,497,521]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="species" species="pfefferi">
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<emphasis id="B909D44BFFDE203DFF6BFE3CFE86C00F" box="[132,303,497,521]" italics="true" pageId="21" pageNumber="52">Siphonalia pfefferi</emphasis>
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G. B.
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<bibRefCitation id="EFEC75A8FFDE203DFE9DFE3CFDF1C00F" author="SOWERBY III G. B." box="[370,600,497,521]" journalOrPublisher="Annals and Magazine of Natural History" pageId="21" pageNumber="52" pagination="439 - 441" part="5" refId="ref13378" refString="SOWERBY III G. B. 1900. - New species of Mollusca of the genera Voluta, Conus, Siphonalia, and Euthria. Annals and Magazine of Natural History series 7 5: 439 - 441." title="New species of Mollusca of the genera Voluta, Conus, Siphonalia, and Euthria" type="journal article" year="1900">Sowerby III, 1900: 440</bibRefCitation>
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</taxonomicName>
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, pl. 11, fig. 3.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3675BD2FFDE203DFF6BFDEDFE66C096" pageId="21" pageNumber="52" type="materials_examined">
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<paragraph id="8BC20859FFDE203DFF6BFDEDFD5BC055" blockId="21.[132,775,544,595]" pageId="21" pageNumber="52">
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MATERIAL EXAMINED. —
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<materialsCitation id="3B150204FFDE203DFE95FDEDFD5BC055" ID-GBIF-Occurrence="2573220329" collectingDate="1996-05-10" country="Japan" location="Kyushu" municipality="Kyushu" pageId="21" pageNumber="52" specimenCount="2" stateProvince="Miyazaki Prefecture">
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<emphasis id="B909D44BFFDE203DFE95FDEDFE1BC03E" bold="true" box="[378,434,544,568]" pageId="21" pageNumber="52">
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<collectingCountry id="F36A48C9FFDE203DFE95FDEDFE1BC03E" box="[378,434,544,568]" name="Japan" pageId="21" pageNumber="52">Japan</collectingCountry>
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</emphasis>
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• 1 lot,
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<specimenCount id="9D7BC3D0FFDE203DFE10FDECFDDEC03F" box="[511,631,545,569]" pageId="21" pageNumber="52" type="generic">2 specimens</specimenCount>
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; Off Hashima,
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<collectingRegion id="49B9C6BBFFDE203DFF6BFDF6FEE0C055" box="[132,329,571,595]" country="Japan" name="Miyazaki" pageId="21" pageNumber="52">Miyazaki Prefecture</collectingRegion>
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,
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<collectingMunicipality id="6BA69223FFDE203DFEBDFDF6FE36C055" box="[338,415,571,595]" pageId="21" pageNumber="52">Kyushu</collectingMunicipality>
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;
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<date id="FFC32E99FFDE203DFE45FDF6FDB8C055" box="[426,529,571,595]" pageId="21" pageNumber="52" value="1996-05-10">
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<collectingDate id="EF87D771FFDE203DFE45FDF6FDB8C055" box="[426,529,571,595]" pageId="21" pageNumber="52" value="1996-05-10">10.V.1996</collectingDate>
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</date>
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(nos. 1, 2, figs 8D, E).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC20859FFDE203DFF6BFDBAFE66C096" blockId="21.[132,775,631,1041]" box="[132,463,631,657]" pageId="21" pageNumber="52">COMPLEMENT TO DESCRIPTION</paragraph>
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</subSubSection>
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<subSubSection id="C3675BD2FFDE203DFF6BFD55FE99C5EB" pageId="21" pageNumber="52" type="description">
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<paragraph id="8BC20859FFDE203DFF6BFD55FF66C0B4" blockId="21.[132,775,631,1041]" box="[132,207,664,690]" pageId="21" pageNumber="52">
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<emphasis id="B909D44BFFDE203DFF6BFD55FF66C0B4" box="[132,207,664,690]" italics="true" pageId="21" pageNumber="52">Radula</emphasis>
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</paragraph>
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<paragraph id="8BC20859FFDE203DFF6BFD7AFE75C617" blockId="21.[132,775,631,1041]" pageId="21" pageNumber="52">
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Radula rather similar in both specimens (
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<figureCitation id="134614DCFFDE203DFDC1FD7AFD02C0D4" box="[558,683,695,722]" captionStart="FIG" captionStartId="20.[132,143,1406,1424]" captionTargetBox="[132,1455,215,1359]" captionTargetId="figure@20.[132,1455,214,1359]" captionTargetPageId="20" captionText="FIG. 11. — Radulae of Siphonalia spp. A, S. cassidariaeformis (Reeve, 1846) no. 1; B, S. cassidariaeformis no. 2; C, S. pfefferi G. B. Sowerby III, 1900 no. 1;" figureDoi="http://doi.org/10.5281/zenodo.3652843" httpUri="https://zenodo.org/record/3652843/files/figure.png" pageId="21" pageNumber="52">Fig. 11C, D</figureCitation>
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); central tooth with rectangular basal part and weakly arcuate anterior margin and three medium long triangular broad cusps, central one equal in length but slightly narrower than lateral ones. Lateral teeth tricuspate with weakly curved basal side, attached to membrane. Outermost cusp recurved, medium long, inner cusp weakly recurved, about 2/3 of outer cusp length; inner cusp in right longitudinal row of specimen spm. no. 2 partially subdivided (
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<figureCitation id="134614DCFFDE203DFEE4FC7AFEC4C1D7" box="[267,365,951,977]" captionStart="FIG" captionStartId="20.[132,143,1406,1424]" captionTargetBox="[132,1455,215,1359]" captionTargetId="figure@20.[132,1455,214,1359]" captionTargetPageId="20" captionText="FIG. 11. — Radulae of Siphonalia spp. A, S. cassidariaeformis (Reeve, 1846) no. 1; B, S. cassidariaeformis no. 2; C, S. pfefferi G. B. Sowerby III, 1900 no. 1;" figureDoi="http://doi.org/10.5281/zenodo.3652843" httpUri="https://zenodo.org/record/3652843/files/figure.png" pageId="21" pageNumber="52">Fig. 11D</figureCitation>
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). Intermediate cusp shortest, situated closer to inner cusp; inner cusp of spm. no. 2 partially subdivided in left longitudinal row.
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</paragraph>
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<paragraph id="8BC20859FFDE203DFF6BFBFAFE54C656" blockId="21.[132,775,1078,1264]" box="[132,509,1078,1105]" pageId="21" pageNumber="52">
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<heading id="D08ABF35FFDE203DFF6BFBFAFE54C656" box="[132,509,1078,1105]" fontSize="11" level="5" pageId="21" pageNumber="52" reason="1">
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<emphasis id="B909D44BFFDE203DFF6BFBFAFE54C656" box="[132,509,1078,1105]" italics="true" pageId="21" pageNumber="52">
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Anatomy (spm. no. 1, male,
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<figureCitation id="134614DCFFDE203DFE46FBFAFE5DC656" box="[425,500,1078,1104]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC20859FFDE203DFF6BFB9BFCAAC6F6" blockId="21.[132,775,1078,1264]" pageId="21" pageNumber="52">
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Head very short and broad, tentacles short, contracted, with small eyes at lobes. Foot contracted, propodium moderately wide, operculum oval with terminal nucleus. Penis rather large (
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<figureCitation id="134614DCFFDE203DFF28FB7BFE8FC6D6" box="[199,294,1206,1232]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12C</figureCitation>
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), flattened, with long narrow seminal papilla in deepening at the top. Mantle with medium long siphon.
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</paragraph>
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<paragraph id="8BC20859FFDE203DFF6BFAD8FE99C5EB" blockId="21.[132,775,1301,2030]" pageId="21" pageNumber="52">
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<emphasis id="B909D44BFFDE203DFF6BFAD8FEFBC729" bold="true" box="[132,338,1301,1327]" pageId="21" pageNumber="52">Digestive system.</emphasis>
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Proboscis partly everted out of rhynchodaeum, with contracted walls. Multiple proboscis retractors attaching mostly along right side of anterior oesophagus (
|
||
<figureCitation id="134614DCFFDE203DFF62FAB8FEAFC796" box="[141,262,1397,1424]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12D, E</figureCitation>
|
||
, prr), connecting rhynchodaeum and lateral walls of body haemocoel. Buccal mass slightly shorter than retracted proboscis (
|
||
<figureCitation id="134614DCFFDE203DFF14FA78FEFAC7C9" box="[251,339,1461,1487]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12F</figureCitation>
|
||
, bm), attaching to its walls by multiple odontophoral retractors (odr). Radula lying in middle of buccal mass and attached to proboscis walls by median retractor (
|
||
<figureCitation id="134614DCFFDE203DFF62F9D9FF49C429" box="[141,224,1556,1583]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12F</figureCitation>
|
||
, mrr). Salivary glands (
|
||
<figureCitation id="134614DCFFDE203DFE20F9D9FDE0C429" box="[463,585,1556,1583]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12D, E</figureCitation>
|
||
, sg) medium large (0.4 proboscis length), oval, with salivary ducts following on both sides of anterior oesophagus. Anterior oesophagus wide, dorso-ventrally flattened (
|
||
<figureCitation id="134614DCFFDE203DFE4CF9B9FDABC488" box="[419,514,1652,1678]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12E</figureCitation>
|
||
, aoe), valve of Leiblein rounded, medium large. Posterior oesophagus (poe) relatively narrow. Gland of Leiblein large, folded beneath nerve ring (
|
||
<figureCitation id="134614DCFFDE203DFF62F919FF4EC4E8" box="[141,231,1748,1774]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12E</figureCitation>
|
||
, gl). Stomach spanning about 0.3 whorl (
|
||
<figureCitation id="134614DCFFDE203DFD76F919FD5EC4E8" box="[665,759,1748,1774]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12G</figureCitation>
|
||
). Posterior mixing area not large (
|
||
<figureCitation id="134614DCFFDE203DFE39F939FDF1C508" box="[470,600,1780,1806]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12G, H</figureCitation>
|
||
, pma). Intestine medium wide. Opening of posterior duct of digestive gland located near oesophageal opening (
|
||
<figureCitation id="134614DCFFDE203DFE19F8FEFDE3C548" box="[502,586,1843,1870]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12I</figureCitation>
|
||
, pdg), opening of anterior duct located closer to beginning of intestine. Inner stomach wall between two openings contains longitudinal fold (
|
||
<figureCitation id="134614DCFFDE203DFF2EF85EFE8FC5AB" box="[193,294,1939,1965]" captionStart="FIG" captionStartId="22.[132,143,1964,1982]" captionTargetBox="[137,1425,241,1794]" captionTargetId="figure@22.[132,1455,215,1834]" captionTargetPageId="22" captionText="FIG. 12. — Anatomy of Siphonalia pfefferi G.B. Sowerby III,1900, specimen no.1: A, cephalopodium,ventral view;B, cephalopodium,dorsal view,mantle removed; C, penis; D, foregut, dorsal view; E, foregut, ventral view; F, proboscis opened dorsally; G, stomach, external view; H, stomach, opened dorsally. Abbreviations: see material and methods. Scale bars: A-F, 10 mm; G, H, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3652847" httpUri="https://zenodo.org/record/3652847/files/figure.png" pageId="21" pageNumber="52">Fig. 12H</figureCitation>
|
||
, lfl), lined with low oblique folds, rest part of inner and outer stomach wall lined with moderately high transverse folds.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3675BD2FFDE203FFCC2FED4FDADC375" lastPageId="23" lastPageNumber="54" pageId="21" pageNumber="52" type="discussion">
|
||
<paragraph id="8BC20859FFDE203DFCC2FED4FB54C0F7" blockId="21.[813,1456,281,2030]" pageId="21" pageNumber="52">
|
||
Results of the phylogenetic analysis suggest close affinities of
|
||
<taxonomicName id="4C7D73DAFFDE203DFCA4FEF4FC15C355" authorityName="A. Adams" authorityYear="1863" box="[843,956,313,339]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFCA4FEF4FC15C355" box="[843,956,313,339]" italics="true" pageId="21" pageNumber="52">Siphonalia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D73DAFFDE203DFC18FEF4FB3CC355" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1015,1173,313,339]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFC18FEF4FB3CC355" box="[1015,1173,313,339]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
that remained unnoticed previously.
|
||
<bibRefCitation id="EFEC75A8FFDE203DFC40FE94FB40C375" author="FRAUSSEN K. & HADORN R." box="[943,1257,345,371]" journalOrPublisher="Novapex" pageId="21" pageNumber="52" pagination="103 - 109" part="7" refId="ref12649" refString="FRAUSSEN K. & HADORN R. 2006. - Phaenomenella, a new genus of deep-water buccinid (Gastropoda: Buccinidae) with the description of a new species from Taiwan. Novapex 7: 103 - 109." title="Phaenomenella, a new genus of deep-water buccinid (Gastropoda: Buccinidae) with the description of a new species from Taiwan" type="journal article" year="2006">Fraussen & Hadorn (2006)</bibRefCitation>
|
||
, while describing
|
||
<taxonomicName id="4C7D73DAFFDE203DFCC2FEB5FC60C394" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[813,969,376,402]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFCC2FEB5FC60C394" box="[813,969,376,402]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
, compared it to
|
||
<taxonomicName id="4C7D73DAFFDE203DFB9AFEB4FB7BC394" authorityName="Smith" authorityYear="1906" box="[1141,1234,377,402]" class="Gastropoda" family="Buccinidae" genus="Manaria" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFB9AFEB4FB7BC394" box="[1141,1234,377,402]" italics="true" pageId="21" pageNumber="52">Manaria</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D73DAFFDE203DFAEAFEB5FAFDC394" authorityName="Thiele" authorityYear="1929" box="[1285,1364,376,402]" class="Gastropoda" family="Buccinidae" genus="Eosipho" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFAEAFEB5FAFDC394" box="[1285,1364,376,402]" italics="true" pageId="21" pageNumber="52">Eosipho</emphasis>
|
||
</taxonomicName>
|
||
, but not to
|
||
<taxonomicName id="4C7D73DAFFDE203DFCA5FE55FC10C3B4" authorityName="A. Adams" authorityYear="1863" box="[842,953,408,434]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFCA5FE55FC10C3B4" box="[842,953,408,434]" italics="true" pageId="21" pageNumber="52">Siphonalia</emphasis>
|
||
</taxonomicName>
|
||
. The shell outline of some
|
||
<taxonomicName id="4C7D73DAFFDE203DFB32FE55FAD0C3B4" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1245,1401,408,434]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFB32FE55FAD0C3B4" box="[1245,1401,408,434]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
(e.g.
|
||
<taxonomicName id="4C7D73DAFFDE203DFCC2FE75FBC6C3D4" baseAuthorityName="Okutani & Lan" baseAuthorityYear="1994" box="[813,1135,440,466]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="species" species="insulapratasensis">
|
||
<emphasis id="B909D44BFFDE203DFCC2FE75FBC6C3D4" box="[813,1135,440,466]" italics="true" pageId="21" pageNumber="52">Phaenomenella insulapratasensis</emphasis>
|
||
</taxonomicName>
|
||
) is rather similar to
|
||
<taxonomicName id="4C7D73DAFFDE203DFAD3FE75FA00C3D4" authorityName="A. Adams" authorityYear="1863" box="[1340,1449,440,466]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFAD3FE75FA00C3D4" box="[1340,1449,440,466]" italics="true" pageId="21" pageNumber="52">Siphonalia</emphasis>
|
||
</taxonomicName>
|
||
: the shell is stout, with strongly convex whorls and a recurved siphonal canal. Species of
|
||
<taxonomicName id="4C7D73DAFFDE203DFBD4FE35FB7CC014" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1083,1237,504,530]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFBD4FE35FB7CC014" box="[1083,1237,504,530]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
though have a much larger (about twice) protoconch in comparison with
|
||
<emphasis id="B909D44BFFDE203DFAD1FDD5FA19C034" box="[1342,1456,536,562]" italics="true" pageId="21" pageNumber="52">
|
||
<taxonomicName id="4C7D73DAFFDE203DFAD1FDD5FA05C034" authorityName="A. Adams" authorityYear="1863" box="[1342,1452,536,562]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">Siphonalia</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The intrageneric variability of shell shape in
|
||
<taxonomicName id="4C7D73DAFFDE203DFB16FDF5FA3DC054" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1273,1428,568,594]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFB16FDF5FA3DC054" box="[1273,1428,568,594]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
in its current definition is very high (
|
||
<bibRefCitation id="EFEC75A8FFDE203DFB4FFD95FCCCC094" author="FRAUSSEN K. & STAHLSCHMIDT P." journalOrPublisher="Novapex" pageId="21" pageNumber="52" pagination="81 - 86" part="14" refId="ref12718" refString="FRAUSSEN K. & STAHLSCHMIDT P. 2013. - The extensive Indo- Pacific radiation of Phaenomenella Fraussen & Hadorn, 2006 (Gastropoda: Buccinidae) with description of a new species. Novapex 14: 81 - 86." title="The extensive Indo- Pacific radiation of Phaenomenella Fraussen & Hadorn, 2006 (Gastropoda: Buccinidae) with description of a new species" type="journal article" year="2013">Fraussen & Stahlschmidt 2013</bibRefCitation>
|
||
) and in its extremes there is no resemblance between the two genera. It should also be born in mind that some of the most diverging species of
|
||
<taxonomicName id="4C7D73DAFFDE203DFB67FD7AFA8DC0D7" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1160,1316,695,721]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFB67FD7AFA8DC0D7" box="[1160,1316,695,721]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
were not yet sequenced and may fall into other lineages.
|
||
</paragraph>
|
||
<paragraph id="8BC20859FFDE203DFCACFD3AFA06C6B6" blockId="21.[813,1456,281,2030]" pageId="21" pageNumber="52">
|
||
Representatives of both genera are also anatomically similar, particularly in the digestive system characters. Both
|
||
<taxonomicName id="4C7D73DAFFDE203DFAB5FCDAFCD7C157" authorityName="Fraussen & Hadorn" authorityYear="2006" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFAB5FCDAFCD7C157" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D73DAFFDE203DFC57FCFAFB80C157" authorityName="A. Adams" authorityYear="1863" box="[952,1065,823,849]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFC57FCFAFB80C157" box="[952,1065,823,849]" italics="true" pageId="21" pageNumber="52">Siphonalia</emphasis>
|
||
</taxonomicName>
|
||
have a broad, medium long in the contracted state proboscis, medium large salivary glands and a large gland of Leiblein. It should be mentioned that despite these general similarities, there are no unique morphological characters uniting both genera. The radular morphology is very similar in both genera (
|
||
<taxonomicName id="4C7D73DAFFDE203DFBA1FC1BFB3BC1F6" box="[1102,1170,982,1008]" class="Jungermanniopsida" family="Radulaceae" genus="Radula" kingdom="Animalia" order="Porellales" pageId="21" pageNumber="52" phylum="Bryophyta" rank="genus">radula</taxonomicName>
|
||
of one more species,
|
||
<taxonomicName id="4C7D73DAFFDE203DFA82FC1AFC51C616" authority="Parth, 1996" authorityName="Parth" authorityYear="1996" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="species" species="marybethi">
|
||
<emphasis id="B909D44BFFDE203DFA82FC1AFCDCC616" italics="true" pageId="21" pageNumber="52">S. marybethi</emphasis>
|
||
Parth, 1996
|
||
</taxonomicName>
|
||
was illustrated in
|
||
<bibRefCitation id="EFEC75A8FFDE203DFB5BFC3BFA09C617" author="ZHANG S. & ZHANG S." box="[1204,1440,1014,1041]" journalOrPublisher="Journal of Oceanology and Limnology" pageId="21" pageNumber="52" pagination="2333 - 2336" part="36" refId="ref13572" refString="ZHANG S. & ZHANG S. 2018. - Three species of Siphonalia Adams, 1863 (Gastropoda, Buccinidae) from China seas, with descriptions of two new species. Journal of Oceanology and Limnology 36: 2333 - 2336. https: // doi. org / 10.1007 / s 00343 - 019 - 7218 - x" title="Three species of Siphonalia Adams, 1863 (Gastropoda, Buccinidae) from China seas, with descriptions of two new species" type="journal article" year="2018">Zhang & Zhang 2018</bibRefCitation>
|
||
), however, as in the case with the body anatomy, it is of rather generalized buccinid appearance; similar radular morphology can be found in many unrelated genera – eg.
|
||
<taxonomicName id="4C7D73DAFFDE203DFAFCFB9BFBA5C697" authority="Dall, 1916" authorityName="Dall" authorityYear="1916" class="Gastropoda" family="Buccinidae" genus="Latisipho" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFAFCFB9BFADAC676" box="[1299,1395,1110,1136]" italics="true" pageId="21" pageNumber="52">Latisipho</emphasis>
|
||
Dall, 1916 (
|
||
<bibRefCitation id="EFEC75A8FFDE203DFC9BFBBAFBABC696" author="KOSYAN A. R." box="[884,1026,1142,1169]" journalOrPublisher="Ruthenica, Russian Malacological Journal" pageId="21" pageNumber="52" pagination="17 - 42" part="16" refId="ref13116" refString="KOSYAN A. R. 2006. - Anatomy and taxonomic composition of the genus Latisipho Dall (Gastropoda: Buccinidae) from the Russian waters. Ruthenica, Russian Malacological Journal 16: 17 - 42." title="Anatomy and taxonomic composition of the genus Latisipho Dall (Gastropoda: Buccinidae) from the Russian waters" type="journal article" year="2006">Kosyan 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7D73DAFFDE203DFBF6FBBBFB5FC696" authority="Dall, 1902" authorityName="Dall" authorityYear="1902" box="[1049,1270,1142,1168]" class="Gastropoda" family="Buccinidae" genus="Plicifusus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFBF6FBBBFBD3C696" box="[1049,1146,1142,1168]" italics="true" pageId="21" pageNumber="52">Plicifusus</emphasis>
|
||
Dall, 1902
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFEC75A8FFDE203DFAEAFBBAFC23C6B6" author="KOSYAN A. R. & KANTOR Y. I." journalOrPublisher="Ruthenica, Russian Malacological Journal" pageId="21" pageNumber="52" pagination="55 - 92" part="22" refId="ref13154" refString="KOSYAN A. R. & KANTOR Y. I. 2012. - Revision of the genus Plicifusus Dall, 1902 (Gastropoda: Buccinidae). Ruthenica, Russian Malacological Journal 22: 55 - 92." title="Revision of the genus Plicifusus Dall, 1902 (Gastropoda: Buccinidae)" type="journal article" year="2012">Kosyan & Kantor 2012</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C7D73DAFFDE203DFC4FFB5BFB3AC6B6" authority="Dall, 1918" authorityName="Dall" authorityYear="1918" box="[928,1171,1174,1200]" class="Gastropoda" family="Buccinidae" genus="Aulacofusus" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFC4FFB5BFBB1C6B6" box="[928,1048,1174,1200]" italics="true" pageId="21" pageNumber="52">Aulacofusus</emphasis>
|
||
Dall, 1918
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFEC75A8FFDE203DFB4DFB5BFA36C6B6" author="KOSYAN A. R. & KANTOR Y. I." box="[1186,1439,1174,1200]" journalOrPublisher="Ruthenica, Russian Malacological Journal" pageId="21" pageNumber="52" pagination="1 - 33" part="23" refId="ref13193" refString="KOSYAN A. R. & KANTOR Y. I. 2013. - Revision of the genus Aulacofusus Dall, 1918 (Gastropoda: Buccinidae). Ruthenica, Russian Malacological Journal 23: 1 - 33." title="Revision of the genus Aulacofusus Dall, 1918 (Gastropoda: Buccinidae)" type="journal article" year="2013">Kosyan & Kantor 2013</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC20859FFDE203DFCACFB7BFB33C528" blockId="21.[813,1456,281,2030]" pageId="21" pageNumber="52">
|
||
Our molecular analysis did not recover
|
||
<taxonomicName id="4C7D73DAFFDE203DFB19FB7BFA3BC6D6" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1270,1426,1206,1232]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFB19FB7BFA3BC6D6" box="[1270,1426,1206,1232]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
as monophyletic. In both COI and combined COI+28S analyses the internal relationships within
|
||
<emphasis id="B909D44BFFDE203DFB65FB38FA06C709" box="[1162,1455,1269,1296]" italics="true" pageId="21" pageNumber="52">
|
||
<taxonomicName id="4C7D73DAFFDE203DFB65FB38FA8FC709" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1162,1318,1269,1295]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">Phaenomenella</taxonomicName>
|
||
–
|
||
<taxonomicName id="4C7D73DAFFDE203DFAAFFB38FA06C709" authorityName="A. Adams" authorityYear="1863" box="[1344,1455,1269,1295]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">Siphonalia</taxonomicName>
|
||
</emphasis>
|
||
clade are not resolved.
|
||
<taxonomicName id="4C7D73DAFFDE203DFBF7FAD8FB49C729" baseAuthorityName="Reeve" baseAuthorityYear="1847" box="[1048,1248,1301,1327]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="species" species="spadicea">
|
||
<emphasis id="B909D44BFFDE203DFBF7FAD8FB49C729" box="[1048,1248,1301,1327]" italics="true" pageId="21" pageNumber="52">Siphonalia spadicea</emphasis>
|
||
</taxonomicName>
|
||
cluster without significant support with
|
||
<taxonomicName id="4C7D73DAFFDE203DFBFCFAFBFB23C749" authority="Kantor & Kosyan & Sorokin, 2020" authorityName="Kantor & Kosyan & Sorokin" authorityYear="2020" box="[1043,1162,1333,1359]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="species" species="samadiae" status="sp. nov.">
|
||
<emphasis id="B909D44BFFDE203DFBFCFAFBFB23C749" box="[1043,1162,1333,1359]" italics="true" pageId="21" pageNumber="52">P. samadiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A6930FFDE203DFB60FAFBFB61C756" box="[1167,1224,1334,1360]" pageId="21" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
|
||
We have only a single species of
|
||
<taxonomicName id="4C7D73DAFFDE203DFC7AFA98FBADC769" authorityName="A. Adams" authorityYear="1863" box="[917,1028,1365,1391]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFC7AFA98FBADC769" box="[917,1028,1365,1391]" italics="true" pageId="21" pageNumber="52">Siphonalia</emphasis>
|
||
</taxonomicName>
|
||
in our analyses so it is too preliminary to change the classification on the basis of the incomplete dataset. Therefore we presently retain the validity of
|
||
<taxonomicName id="4C7D73DAFFDE203DFAE3FA58FA01C7A9" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1292,1448,1429,1455]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFAE3FA58FA01C7A9" box="[1292,1448,1429,1455]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
, although it is possible that
|
||
<taxonomicName id="4C7D73DAFFDE203DFBA8FA78FB4AC7C9" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1095,1251,1461,1487]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFBA8FA78FB4AC7C9" box="[1095,1251,1461,1487]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7D73DAFFDE203DFAFAFA78FA2DC7C9" authorityName="A. Adams" authorityYear="1863" box="[1301,1412,1461,1487]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFAFAFA78FA2DC7C9" box="[1301,1412,1461,1487]" italics="true" pageId="21" pageNumber="52">Siphonalia</emphasis>
|
||
</taxonomicName>
|
||
can belong to a single genus. One of the distinctions between the genera is the depth range of known species. Generally, species of
|
||
<taxonomicName id="4C7D73DAFFDE203DFC76F9D9FBA1C428" authorityName="A. Adams" authorityYear="1863" box="[921,1032,1556,1582]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFC76F9D9FBA1C428" box="[921,1032,1556,1582]" italics="true" pageId="21" pageNumber="52">Siphonalia</emphasis>
|
||
</taxonomicName>
|
||
dwell at shallower depths – from
|
||
<quantity id="4C85A5BCFFDE203DFA99F9D8FCDDC449" metricMagnitude="2" metricUnit="m" metricValue="1.55" metricValueMax="3.0" metricValueMin="0.1" pageId="21" pageNumber="52" unit="m" value="155.0" valueMax="300.0" valueMin="10.0">10 to 300 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFEC75A8FFDE203DFC6AF9F9FB88C449" author="OKUTANI T." bookContentInfo="1221 p." box="[901,1057,1588,1615]" journalOrPublisher="Tokyo, Tokai University Press" pageId="21" pageNumber="52" refId="ref13277" refString="OKUTANI T. 2000. - Marine mollusks in Japan. Tokyo, Tokai University Press, 1221 p." title="Marine mollusks in Japan" type="book" year="2000">Okutani 2000</bibRefCitation>
|
||
), while
|
||
<taxonomicName id="4C7D73DAFFDE203DFB95F9F9FABFC448" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1146,1302,1588,1614]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFB95F9F9FABFC448" box="[1146,1302,1588,1614]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
is recorded at
|
||
<quantity id="4C85A5BCFFDE203DFCC2F999FC1FC469" box="[813,950,1620,1647]" metricMagnitude="2" metricUnit="m" metricValue="7.895" metricValueMax="13.89" metricValueMin="1.9" pageId="21" pageNumber="52" unit="m" value="789.5" valueMax="1389.0" valueMin="190.0">190-1389 m</quantity>
|
||
(
|
||
<bibRefCitation id="EFEC75A8FFDE203DFC27F998FAB3C468" author="FRAUSSEN K. & STAHLSCHMIDT P." box="[968,1306,1620,1647]" journalOrPublisher="Novapex" pageId="21" pageNumber="52" pagination="81 - 86" part="14" refId="ref12718" refString="FRAUSSEN K. & STAHLSCHMIDT P. 2013. - The extensive Indo- Pacific radiation of Phaenomenella Fraussen & Hadorn, 2006 (Gastropoda: Buccinidae) with description of a new species. Novapex 14: 81 - 86." title="The extensive Indo- Pacific radiation of Phaenomenella Fraussen & Hadorn, 2006 (Gastropoda: Buccinidae) with description of a new species" type="journal article" year="2013">Fraussen & Stahlschmidt 2013</bibRefCitation>
|
||
; herein). The new species are attributed to
|
||
<taxonomicName id="4C7D73DAFFDE203DFBB7F9B9FB5BC488" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1112,1266,1652,1678]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFBB7F9B9FB5BC488" box="[1112,1266,1652,1678]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
based on stronger conchological similarity to other species of the genus rather than to species of
|
||
<taxonomicName id="4C7D73DAFFDE203DFC02F979FBF5C4C8" authorityName="A. Adams" authorityYear="1863" box="[1005,1116,1716,1742]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFC02F979FBF5C4C8" box="[1005,1116,1716,1742]" italics="true" pageId="21" pageNumber="52">Siphonalia</emphasis>
|
||
</taxonomicName>
|
||
. Unfortunately the protoconch of
|
||
<taxonomicName id="4C7D73DAFFDE203DFCA7F918FC17C4E8" authority="Kantor & Kosyan & Sorokin, 2020" authorityName="Kantor & Kosyan & Sorokin" authorityYear="2020" box="[840,958,1748,1774]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="species" species="samadiae" status="sp. nov.">
|
||
<emphasis id="B909D44BFFDE203DFCA7F918FC17C4E8" box="[840,958,1748,1774]" italics="true" pageId="21" pageNumber="52">P. samadiae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A6930FFDE203DFC2CF919FC55C4E8" box="[963,1020,1748,1774]" pageId="21" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
|
||
was decollated in all available specimens, but the protoconch of
|
||
<taxonomicName id="4C7D73DAFFDE203DFBCDF938FBC4C508" authority="Kantor & Kosyan & Sorokin, 2020" authorityName="Kantor & Kosyan & Sorokin" authorityYear="2020" box="[1058,1133,1781,1806]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="species" species="nicoi" status="sp. nov.">
|
||
<emphasis id="B909D44BFFDE203DFBCDF938FBC4C508" box="[1058,1133,1781,1806]" italics="true" pageId="21" pageNumber="52">P. nicoi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23A6930FFDE203DFB9BF939FB06C508" box="[1140,1199,1780,1806]" pageId="21" pageNumber="52" rank="species">n. sp.</taxonomicNameLabel>
|
||
is large globose, similar to other species of
|
||
<taxonomicName id="4C7D73DAFFDE203DFC19F8D9FB3BC528" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[1014,1170,1812,1838]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFC19F8D9FB3BC528" box="[1014,1170,1812,1838]" italics="true" pageId="21" pageNumber="52">Phaenomenella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC20859FFDE203FFCACF8FEFDADC375" blockId="21.[813,1456,281,2030]" lastBlockId="23.[132,776,217,371]" lastPageId="23" lastPageNumber="54" pageId="21" pageNumber="52">
|
||
The analysis of a broader dataset of Buccinoidea rejected the monophyly of
|
||
<taxonomicName id="4C7D73DAFFDE203DFC18F89EFB23C56B" authorityName="Finlay" authorityYear="1928" box="[1015,1162,1875,1901]" class="Gastropoda" family="Buccinidae" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="subFamily" subFamily="Siphonaliinae">Siphonaliinae</taxonomicName>
|
||
in its original scope. None of the Recent genera, originally included by
|
||
<bibRefCitation id="EFEC75A8FFDE203DFAEDF8BEFA3DC588" author="FINLAY H. J." box="[1282,1428,1907,1934]" journalOrPublisher="Transactions and Proceedings of the Royal Society of New Zealand" pageId="21" pageNumber="52" pagination="232 - 286" part="59" refId="ref12507" refString="FINLAY H. J. 1928. - The recent mollusca of the Chatham Islands. Transactions and Proceedings of the Royal Society of New Zealand 59: 232 - 286, pls 38 - 43." title="The recent mollusca of the Chatham Islands" type="journal article" year="1928">Finlay (1928)</bibRefCitation>
|
||
in the subfamily, that are
|
||
<taxonomicName id="4C7D73DAFFDE203DFBCBF859FA2CC5AB" authority=", Aeneator and Glaphyrina" authorityName="Aeneator and Glaphyrina" box="[1060,1413,1939,1966]" class="Gastropoda" family="Buccinidae" genus="Penion" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B909D44BFFDE203DFBCBF859FB7DC5A8" box="[1060,1236,1940,1966]" italics="true" pageId="21" pageNumber="52">Penion, Aeneator</emphasis>
|
||
and
|
||
<emphasis id="B909D44BFFDE203DFAE4F85EFA2CC5AB" box="[1291,1413,1939,1965]" italics="true" pageId="21" pageNumber="52">Glaphyrina</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B909D44BFFDE203DFA6AF85EFA20C5AB" box="[1413,1417,1939,1965]" italics="true" pageId="21" pageNumber="52">,</emphasis>
|
||
are closely related neither to each other, nor to
|
||
<emphasis id="B909D44BFFDE203DFAE7F87EFAD6C5CB" box="[1288,1407,1971,1997]" italics="true" pageId="21" pageNumber="52">
|
||
<taxonomicName id="4C7D73DAFFDE203DFAE7F87EFAD2C5CB" authorityName="A. Adams" authorityYear="1863" box="[1288,1403,1971,1997]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="genus">Siphonalia</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The system of
|
||
<taxonomicName id="4C7D73DAFFDE203DFC74F81EFBBBC5EB" authorityName="Rafinesque" authorityYear="1815" box="[923,1042,2003,2029]" class="Gastropoda" family="Buccinidae" kingdom="Animalia" order="Neogastropoda" pageId="21" pageNumber="52" phylum="Mollusca" rank="family">Buccinidae</taxonomicName>
|
||
and Buccinoidea in general is still far from being resolved, with many problematic buccinoidean lineages (see e.g.
|
||
<bibRefCitation id="EFEC75A8FFDC203FFEDEFF34FE4FC315" author="COUTO D. R. & BOUCHET P. & KANTOR Y. I. & SIMONE L. R. L. & GIRIBET G." box="[305,486,249,275]" journalOrPublisher="Molecular Phylogenics and Evolution" pageId="23" pageNumber="54" pagination="309 - 322" part="99" refId="ref12398" refString="COUTO D. R., BOUCHET P., KANTOR Y. I., SIMONE L. R. L. & GIRIBET G. 2016. - A multilocus molecular phylogeny of Fasciolariidae (Neogastropoda: Buccinoidea). Molecular Phylogenics and Evolution 99: 309 - 322. https: // doi. org / 10.1016 / j. ympev. 2016.03.025" title="A multilocus molecular phylogeny of Fasciolariidae (Neogastropoda: Buccinoidea)" type="journal article" year="2016">
|
||
Couto
|
||
<emphasis id="B909D44BFFDC203FFE95FF37FE02C315" box="[378,427,249,275]" italics="true" pageId="23" pageNumber="54">et al.</emphasis>
|
||
2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEC75A8FFDC203FFE1DFF34FD07C315" author="HARASEWYCH M. G." box="[498,686,249,275]" journalOrPublisher="The Nautilus" pageId="23" pageNumber="54" pagination="35 - 44" part="132" refId="ref12910" refString="HARASEWYCH M. G. 2018. - The anatomy of Tudicla spirillus (Linnaeus, 1767) and the relationships of the Tudiclidae (Gastropoda: Neogastropoda). The Nautilus 132: 35 - 44." title="The anatomy of Tudicla spirillus (Linnaeus, 1767) and the relationships of the Tudiclidae (Gastropoda: Neogastropoda)" type="journal article" year="2018">Harasewych 2018</bibRefCitation>
|
||
). Therefore the rank of the inferred clade
|
||
<emphasis id="B909D44BFFDC203FFE0AFED4FCA1C335" box="[485,776,281,307]" italics="true" pageId="23" pageNumber="54">
|
||
<taxonomicName id="4C7D73DAFFDC203FFE0AFED4FDFAC335" authorityName="A. Adams" authorityYear="1863" box="[485,595,281,307]" class="Gastropoda" family="Buccinidae" genus="Siphonalia" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="54" phylum="Mollusca" rank="genus">Siphonalia</taxonomicName>
|
||
+
|
||
<taxonomicName id="4C7D73DAFFDC203FFD82FED4FCA1C335" authorityName="Fraussen & Hadorn" authorityYear="2006" box="[621,776,281,307]" class="Gastropoda" family="Buccinidae" genus="Phaenomenella" kingdom="Animalia" order="Neogastropoda" pageId="23" pageNumber="54" phylum="Mollusca" rank="genus">Phaenomenella</taxonomicName>
|
||
</emphasis>
|
||
can be resolved only after obtaining the robust phylogeny of the entire superfamily Buccinoidea.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |