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<mods:title id="7DAE87787C9DE307A3C419B0F3ED379D">Studies on Neotropical crickets: Aclodes paz n. sp. a new phalangopsid cricket (Orthoptera: Phalangopsidae) from the Santander caves, Colombia</mods:title>
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<treatment id="03B6878EFFB8342375C76BBB1789FA1C" ID-DOI="http://doi.org/10.5281/zenodo.6599712" ID-GBIF-Taxon="195705066" ID-Zenodo-Dep="6599712" LSID="urn:lsid:plazi:treatment:03B6878EFFB8342375C76BBB1789FA1C" httpUri="http://treatment.plazi.org/id/03B6878EFFB8342375C76BBB1789FA1C" lastPageNumber="570" pageId="2" pageNumber="570">
<subSubSection id="C3056513FFB8342375C76BBB17C5FF4B" box="[151,673,151,177]" pageId="2" pageNumber="570" type="nomenclature">
<paragraph id="8BA03698FFB8342375C76BBB17C5FF4B" blockId="2.[151,673,151,177]" box="[151,673,151,177]" pageId="2" pageNumber="570">
<heading id="D0E881F4FFB8342375C76BBB17C5FF4B" bold="true" box="[151,673,151,177]" fontSize="11" level="1" pageId="2" pageNumber="570" reason="1">
<emphasis id="B96BEA8AFFB8342375C76BBB17C5FF4B" bold="true" box="[151,673,151,177]" pageId="2" pageNumber="570">
Tribe
<taxonomicName id="4C1F4D1BFFB8342375B16BBB17C5FF4B" ID-CoL="8QHKW" authority="Desutter-Grandcolas, 1992" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[225,673,151,177]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="tribe" tribe="Aclodini">Aclodini Desutter-Grandcolas, 1992</taxonomicName>
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<paragraph id="8BA03698FFB8342375C76BF31422FE2B" blockId="2.[151,1437,223,1510]" pageId="2" pageNumber="570">
<emphasis id="B96BEA8AFFB8342375C76BF3147BFF03" bold="true" box="[151,287,223,249]" pageId="2" pageNumber="570">Comments.</emphasis>
This tribe was proposed by Desutter-Grandcolas (1992), initially as the group “Aclodae”, including
<taxonomicName id="4C1F4D1BFFB8342375C76A2F14F7FEE7" authority="Hebard, 1928" authorityName="Hebard" authorityYear="1928" box="[151,403,259,285]" class="Insecta" family="Phalangopsidae" genus="Aclodes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342375C76A2F1594FEE7" box="[151,240,259,285]" italics="true" pageId="2" pageNumber="570">Aclodes</emphasis>
<bibRefCitation id="EF8E4B69FFB8342375A76A2F14F7FEE7" author="Hebard, M." box="[247,403,259,285]" pageId="2" pageNumber="570" pagination="79 - 124" refId="ref7325" refString="Hebard, M. (1928) Studies in the Dermaptera and Orthoptera of Colombia. Fifth paper. Orthopterous family Gryllidae. Transactions of the American Entomological Society, 54 (2), 79 - 124." type="journal article" year="1928">Hebard, 1928</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1F4D1BFFB8342374CE6A2F1636FEE7" authority="Desutter-Grandcolas, 1992" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[414,850,259,285]" class="Insecta" family="Phalangopsidae" genus="Paraclodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342374CE6A2F177AFEE7" box="[414,542,259,285]" italics="true" pageId="2" pageNumber="570">Paraclodes</emphasis>
Desutter-Grandcolas, 1992
</taxonomicName>
and
<taxonomicName id="4C1F4D1BFFB8342376D96A2F11DBFEE7" authority="Chopard, 1923" authorityName="Chopard" authorityYear="1923" box="[905,1215,259,285]" class="Insecta" family="Phalangopsidae" genus="Uvaroviella" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342376D96A2F116AFEE7" box="[905,1038,259,285]" italics="true" pageId="2" pageNumber="570">Uvaroviella</emphasis>
Chopard, 1923
</taxonomicName>
. The author defines the group and suggests that two species of
<taxonomicName id="4C1F4D1BFFB8342377D36A0B16B0FEBB" authority="Saussure, 1874" authorityName="Saussure" authorityYear="1874" box="[643,980,295,321]" class="Insecta" family="Phalangopsidae" genus="Heterogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342377D36A0B167AFEBB" box="[643,798,295,321]" italics="true" pageId="2" pageNumber="570">Heterogryllus</emphasis>
Saussure, 1874
</taxonomicName>
,
<taxonomicName id="4C1F4D1BFFB8342376B06A041027FEBB" authority="Saussure, 1878" authorityName="Saussure" authorityYear="1878" box="[992,1347,295,321]" class="Insecta" family="Phalangopsidae" genus="Heterogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="B96BEA8AFFB8342376B06A0411E9FEBB" box="[992,1165,296,321]" italics="true" pageId="2" pageNumber="570">H. crassicornis</emphasis>
Saussure, 1878
</taxonomicName>
, and
<taxonomicName id="4C1F4D1BFFB8342370D16A0414C6FE9F" authority="Chopard, 1970" authorityName="Chopard" authorityYear="1970" class="Insecta" family="Phalangopsidae" genus="Heterogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="species" species="bordoni">
<emphasis id="B96BEA8AFFB8342370D16A04158BFE9F" italics="true" pageId="2" pageNumber="570">H. bordoni</emphasis>
Chopard, 1970
</taxonomicName>
, were located in
<taxonomicName id="4C1F4D1BFFB8342377346A6717D9FE9F" authorityName="Hebard" authorityYear="1928" box="[612,701,331,357]" class="Insecta" family="Phalangopsidae" genus="Aclodes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342377346A6717D9FE9F" box="[612,701,331,357]" italics="true" pageId="2" pageNumber="570">Aclodes</emphasis>
</taxonomicName>
and
<emphasis id="B96BEA8AFFB8342377A66A671618FE9F" box="[758,892,331,357]" italics="true" pageId="2" pageNumber="570">
<taxonomicName id="4C1F4D1BFFB8342377A66A67161CFE9F" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[758,888,331,357]" class="Insecta" family="Phalangopsidae" genus="Paraclodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">Paraclodes</taxonomicName>
,
</emphasis>
respectively. Desutter-Grandcolas (1992), mentions that
<taxonomicName id="4C1F4D1BFFB83423745C6A4314C3FE73" authorityName="Saussure" authorityYear="1874" box="[268,423,367,393]" class="Insecta" family="Phalangopsidae" genus="Heterogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB83423745C6A4314C3FE73" box="[268,423,367,393]" italics="true" pageId="2" pageNumber="570">Heterogryllus</emphasis>
</taxonomicName>
only includes
<taxonomicName id="4C1F4D1BFFB8342377046A5C16F1FE73" authority="Saussure, 1874" authorityName="Saussure" authorityYear="1874" box="[596,917,367,393]" class="Insecta" family="Phalangopsidae" genus="Heterogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="species" species="ocellaris">
<emphasis id="B96BEA8AFFB8342377046A5C17BFFE73" box="[596,731,367,393]" italics="true" pageId="2" pageNumber="570">H. ocellaris</emphasis>
Saussure, 1874
</taxonomicName>
, represented by a female collected in
<collectingCountry id="F3087608FFB8342370016A4310FCFE73" box="[1361,1432,367,393]" name="Brazil" pageId="2" pageNumber="570">Brazil</collectingCountry>
, without precise locality, and that, due to its morphological characteristics, it should be included in Neoaclini (
<emphasis id="B96BEA8AFFB83423700E6AB810F8FE57" box="[1374,1436,404,429]" italics="true" pageId="2" pageNumber="570">sensu</emphasis>
<bibRefCitation id="EF8E4B69FFB8342375C76A94145DFE2B" author="Desutter, L." box="[151,313,439,466]" pageId="2" pageNumber="570" pagination="213 - 239" refId="ref6702" refString="Desutter, L. (1987) Structure et evolution du complexe phallique de Gryllidea (Orthopteres) et classification des genres neotropicaux de Grylloidea. Premiere partie. Annales de la Societe Entomologique de France, New Series, 23 (3), 213 - 239." type="journal article" year="1987">Desutter 1987</bibRefCitation>
).
</paragraph>
<paragraph id="8BA03698FFB8342375976AF710F9FD9B" blockId="2.[151,1437,223,1510]" pageId="2" pageNumber="570">
<bibRefCitation id="EF8E4B69FFB8342375976AF714A1FE0F" author="Nischk, F. &amp; Otte, D." box="[199,453,475,501]" pageId="2" pageNumber="570" pagination="229 - 254" refId="ref7565" refString="Nischk, F. &amp; Otte, D. (2000) Bioacoustics, ecology and systematics of Ecuadorian rainforest crickets (Orthoptera: Gryllidae: Phalangopsinae), with a description of four new genera and ten new species. Journal of Orthoptera Research, 9, 229 - 254. https: // doi. org / 10.2307 / 3503651" type="journal article" year="2000">Nischk &amp; Otte (2000)</bibRefCitation>
described several taxa for
<collectingCountry id="F3087608FFB8342377AF6AF7163AFE0F" box="[767,862,475,501]" name="Ecuador" pageId="2" pageNumber="570">Ecuador</collectingCountry>
, one species for
<taxonomicName id="4C1F4D1BFFB8342371746AF711C0FE0F" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[1060,1188,475,501]" class="Insecta" family="Phalangopsidae" genus="Paraclodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342371746AF711C0FE0F" box="[1060,1188,475,501]" italics="true" pageId="2" pageNumber="570">Paraclodes</emphasis>
</taxonomicName>
and three species for
<taxonomicName id="4C1F4D1BFFB8342375C76AD31594FDE3" authorityName="Hebard" authorityYear="1928" box="[151,240,511,537]" class="Insecta" family="Phalangopsidae" genus="Aclodes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342375C76AD31594FDE3" box="[151,240,511,537]" italics="true" pageId="2" pageNumber="570">Aclodes</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF8E4B69FFB8342375AD6AD3141AFDE0" author="Otte, D." box="[253,382,511,538]" pageId="2" pageNumber="570" pagination="299 - 418" refId="ref7627" refString="Otte, D. (2006) Eighty-four new cricket species (Orthoptera: Grylloidea) from La Selva, Costa Rica. Transactions of the American Entomological Society, 132 (3 - 4), 299 - 418." type="journal article" year="2006">Otte (2006)</bibRefCitation>
describes five additional species from
<collectingCountry id="F3087608FFB83423767C6AD316CDFDE3" box="[812,937,511,537]" name="Costa Rica" pageId="2" pageNumber="570">Costa Rica</collectingCountry>
.
<bibRefCitation id="EF8E4B69FFB8342376E26AD31199FDE3" author="Otte, D. &amp; Perez-Gelabert, D. E." box="[946,1277,511,537]" pageId="2" pageNumber="570" refId="ref7668" refString="Otte, D. &amp; Perez-Gelabert, D. E. (2009) Caribbean crickets. The Orthopterists' Society, Philadelphia, Pennsylvania, iii + 792 pp. https: // doi. org / 10.3157 / 0002 - 8320 (2006) 132 [299: ENCSOG] 2.0. CO; 2" type="book" year="2009">Otte &amp; Perez-Gelabert (2009)</bibRefCitation>
added 16 species to
<taxonomicName id="4C1F4D1BFFB8342375B6690F140FFDC7" authorityName="Chopard" authorityYear="1923" box="[230,363,547,573]" class="Insecta" family="Phalangopsidae" genus="Uvaroviella" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342375B6690F140FFDC7" box="[230,363,547,573]" italics="true" pageId="2" pageNumber="570">Uvaroviella</emphasis>
</taxonomicName>
, distributed on several Caribbean islands. Subsequently, several taxa described by the above authors will be reassigned to other genera or synonymized, and not all species will retain their original combinations.
</paragraph>
<paragraph id="8BA03698FFB8342375976947103EFC5C" blockId="2.[151,1437,223,1510]" pageId="2" pageNumber="570">
<bibRefCitation id="EF8E4B69FFB834237597694714E8FD7F" author="Gorochov, A. V." box="[199,396,619,645]" pageId="2" pageNumber="570" pagination="1183 - 1195" refId="ref7116" refString="Gorochov, A. V. (2007) New and little known crickets of the subfamily Phalangopsinae (Orthoptera, Gryllidae). 4. Neotropical genus Uvaroviella. Zoologicheskii Zhurnal, 86 (10), 1183 - 1195. https: // doi. org / 10.1134 / S 0013873806040087" type="journal article" year="2007">Gorochov (2007)</bibRefCitation>
proposed that the Aclodae group comprises a single genus.
<emphasis id="B96BEA8AFFB834237116694711B5FD7F" box="[1094,1233,619,645]" italics="true" pageId="2" pageNumber="570">
<taxonomicName id="4C1F4D1BFFB834237116694711A9FD7F" authorityName="Chopard" authorityYear="1923" box="[1094,1229,619,645]" class="Insecta" family="Phalangopsidae" genus="Uvaroviella" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">Uvaroviella</taxonomicName>
,
</emphasis>
the oldest genus, retained generic status, the others as subgenera: four originally described as genera (
<taxonomicName id="4C1F4D1BFFB83423710869A31054FD53" authority="Hebard, 1928" authorityName="Hebard" authorityYear="1928" box="[1112,1328,655,681]" class="Insecta" family="Phalangopsidae" genus="Acla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB83423710869A311EFFD53" box="[1112,1163,655,681]" italics="true" pageId="2" pageNumber="570">Acla</emphasis>
<bibRefCitation id="EF8E4B69FFB8342371C469A31054FD53" author="Hebard, M." box="[1172,1328,655,681]" pageId="2" pageNumber="570" pagination="79 - 124" refId="ref7325" refString="Hebard, M. (1928) Studies in the Dermaptera and Orthoptera of Colombia. Fifth paper. Orthopterous family Gryllidae. Transactions of the American Entomological Society, 54 (2), 79 - 124." type="journal article" year="1928">Hebard, 1928</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C1F4D1BFFB83423706D69A310F2FD53" authorityName="Hebard" authorityYear="1928" box="[1341,1430,655,681]" class="Insecta" family="Phalangopsidae" genus="Aclodes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB83423706D69A310F2FD53" box="[1341,1430,655,681]" italics="true" pageId="2" pageNumber="570">Aclodes</emphasis>
</taxonomicName>
,
<emphasis id="B96BEA8AFFB8342375C7699F147AFD37" box="[151,286,691,717]" italics="true" pageId="2" pageNumber="570">
<taxonomicName id="4C1F4D1BFFB8342375C7699F147DFD37" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[151,281,691,717]" class="Insecta" family="Phalangopsidae" genus="Paraclodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">Paraclodes</taxonomicName>
,
</emphasis>
and
<emphasis id="B96BEA8AFFB834237404699F1767FD37" box="[340,515,691,717]" italics="true" pageId="2" pageNumber="570">
<taxonomicName id="4C1F4D1BFFB834237404699F14BDFD37" authorityName="Chopard" authorityYear="1923" box="[340,473,691,717]" class="Insecta" family="Phalangopsidae" genus="Uvaroviella" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus" sensu="stricto">Uvaroviella</taxonomicName>
<taxonomicNameLabel id="A25857F1FFB83423748F69981767FD37" box="[479,515,692,717]" pageId="2" pageNumber="570" sensu="stricto">s.s.</taxonomicNameLabel>
</emphasis>
) and five as new subgenera (
<taxonomicName id="4C1F4D1BFFB834237619699F16C7FD37" authorityName="Gorochov" authorityYear="2007" box="[841,931,691,717]" class="Insecta" family="Phalangopsidae" genus="Subacla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB834237619699F16C7FD37" box="[841,931,691,717]" italics="true" pageId="2" pageNumber="570">Subacla</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1F4D1BFFB8342376E1699F1164FD37" baseAuthorityName="sensu Gorochov" baseAuthorityYear="2007" box="[945,1024,691,717]" class="Insecta" family="Phalangopsidae" genus="Euacla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342376E1699F1164FD37" box="[945,1024,691,717]" italics="true" pageId="2" pageNumber="570">Euacla</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1F4D1BFFB83423715E699F1138FD37" authorityName="Gorochov" authorityYear="2007" box="[1038,1116,691,717]" class="Insecta" family="Phalangopsidae" genus="Reacla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB83423715E699F1138FD37" box="[1038,1116,691,717]" italics="true" pageId="2" pageNumber="570">Reacla</emphasis>
</taxonomicName>
,
<emphasis id="B96BEA8AFFB83423713A699F11AFFD37" box="[1130,1227,691,717]" italics="true" pageId="2" pageNumber="570">Holacla,</emphasis>
and
<taxonomicName id="4C1F4D1BFFB834237051699F103EFD37" authorityName="Gorochov" authorityYear="2007" box="[1281,1370,691,717]" class="Insecta" family="Phalangopsidae" genus="Topacla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB834237051699F103EFD37" box="[1281,1370,691,717]" italics="true" pageId="2" pageNumber="570">Topacla</emphasis>
</taxonomicName>
). The same author adds six species in
<taxonomicName id="4C1F4D1BFFB8342374AE69FB17E7FD0B" authorityName="Chopard" authorityYear="1923" box="[510,643,727,753]" class="Insecta" family="Phalangopsidae" genus="Uvaroviella" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B96BEA8AFFB8342374AE69FB17E7FD0B" box="[510,643,727,753]" italics="true" pageId="2" pageNumber="570">Uvaroviella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25857F1FFB8342377DA69FB17CDFD08" box="[650,681,727,754]" pageId="2" pageNumber="570" sensu="lato">
<emphasis id="B96BEA8AFFB8342377DA69FB17C0FD0B" box="[650,676,727,753]" italics="true" pageId="2" pageNumber="570">s.l</emphasis>
.
</taxonomicNameLabel>
, and keeps this taxon in the tribe
<taxonomicName id="4C1F4D1BFFB83423717569FB11A3FD0B" authorityName="Blanchard" authorityYear="1845" box="[1061,1223,727,753]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="tribe" tribe="Phalangopsini">Phalangopsini</taxonomicName>
; for
<taxonomicName id="4C1F4D1BFFB8342371AA69FB10E5FD0B" authorityName="Desutter-Grandcolas" authorityYear="1987" box="[1274,1409,727,753]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="tribe" tribe="Paragryllini">Paragryllini</taxonomicName>
, it gives a new status, including subtribes described as tribes by Desutter (Paragryllina
<emphasis id="B96BEA8AFFB83423711369D01103FCEF" box="[1091,1127,764,789]" italics="true" pageId="2" pageNumber="570">s.s.</emphasis>
, Neoaclina and Strogulomorphina). This contribution initiated some synonymization of taxa described by Otte and other authors (
<bibRefCitation id="EF8E4B69FFB83423707A683315B7FCA7" author="Gorochov, A. V." pageId="2" pageNumber="570" pagination="1055 - 1069" refId="ref7228" refString="Gorochov, A. V. (2011) New and little known Phalangopsinae (Orthoptera, Gryllidae) 7. Neotropic taxa of the tribes Paragryllini and Luzarini. Zoologicheskii Zhurnal, 90 (9), 1055 - 1069." type="journal article" year="2011">Gorochov 2011</bibRefCitation>
).
<bibRefCitation id="EF8E4B69FFB8342375B7686F14CFFCA7" author="Gorochov, A. V." box="[231,427,835,861]" pageId="2" pageNumber="570" pagination="7 - 88" refId="ref7270" refString="Gorochov, A. V. (2014) Classification of the Phalangopsinae subfamily group, and new taxa from the subfamilies Phalangopsinae and Phaloriinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 23 (1), 7 - 88. https: // doi. org / 10.31610 / zsr / 2014.23.1.7" type="journal article" year="2014">Gorochov (2014)</bibRefCitation>
proposed a classification for the
<taxonomicName id="4C1F4D1BFFB834237679686F16BDFCA7" box="[809,985,835,861]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="subFamily" subFamily="Phalangopsinae">Phalangopsinae</taxonomicName>
subfamily group.
<taxonomicName id="4C1F4D1BFFB8342371FC686F1055FCA7" authorityName="Chopard" authorityYear="1923" box="[1196,1329,835,861]" class="Insecta" family="Phalangopsidae" genus="Uvaroviella" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342371FC686F1055FCA7" box="[1196,1329,835,861]" italics="true" pageId="2" pageNumber="570">Uvaroviella</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1F4D1BFFB83423703A6868146DFC7B" authorityName="Saussure" authorityYear="1874" class="Insecta" family="Phalangopsidae" genus="Heterogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB83423703A6868146DFC7B" italics="true" pageId="2" pageNumber="570">Heterogryllus</emphasis>
</taxonomicName>
are included in the subtribe
<taxonomicName id="4C1F4D1BFFB834237718684B16C2FC78" authority="(Phalangopsini)" baseAuthorityName="Phalangopsini" box="[584,934,871,898]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="subTribe" subTribe="Heterogryllina">Heterogryllina (Phalangopsini)</taxonomicName>
, sugesting the probable relationship between both taxa, and indicating that Aclodae is probably junior synonym of
<taxonomicName id="4C1F4D1BFFB8342376F168A71120FC5F" box="[929,1092,907,933]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="subTribe" subTribe="Heterogryllina">Heterogryllina</taxonomicName>
(
<bibRefCitation id="EF8E4B69FFB83423710368A71062FC5F" author="Gorochov, A. V." box="[1107,1286,907,933]" pageId="2" pageNumber="570" pagination="7 - 88" refId="ref7270" refString="Gorochov, A. V. (2014) Classification of the Phalangopsinae subfamily group, and new taxa from the subfamilies Phalangopsinae and Phaloriinae (Orthoptera: Gryllidae). Zoosystematica Rossica, 23 (1), 7 - 88. https: // doi. org / 10.31610 / zsr / 2014.23.1.7" type="journal article" year="2014">Gorochov 2014</bibRefCitation>
,
<bibRefCitation id="EF8E4B69FFB83423704168A7102AFC5C" author="Gorochov, A. V." box="[1297,1358,907,934]" pageId="2" pageNumber="570" pagination="600 - 611" refId="ref7169" refString="Gorochov, A. V. (2015) New and little-known crickets of the subfamily Phalangopsinae (Orthoptera, Gryllidae). 8. The genus Paragryllodes and notes on the subfamily classification. Entomological Review, 95 (5), 600 - 611. https: // doi. org / 10.1134 / S 001387381505005 X" type="journal article" year="2015">2015</bibRefCitation>
).
</paragraph>
<paragraph id="8BA03698FFB8342375976883162CFACB" blockId="2.[151,1437,223,1510]" pageId="2" pageNumber="570">
<bibRefCitation id="EF8E4B69FFB83423759768831764FC33" author="Desutter-Grandcolas, L." box="[199,512,943,969]" pageId="2" pageNumber="570" pagination="398 - 420" refId="ref6909" refString="Desutter-Grandcolas, L. (2014) New taxa and data for Neotropical Phalangopsidae (Orthoptera, Grylloidea). Zootaxa, 3866 (3), 398 - 420. https: // doi. org / 10.11646 / zootaxa. 3866.3.5" type="journal article" year="2014">Desutter-Grandcolas (2014)</bibRefCitation>
officially proposed
<taxonomicName id="4C1F4D1BFFB8342377B268831671FC33" authorityName="Hebard" authorityYear="1928" box="[738,789,943,969]" class="Insecta" family="Phalangopsidae" genus="Acla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342377B268831671FC33" box="[738,789,943,969]" italics="true" pageId="2" pageNumber="570">Acla</emphasis>
</taxonomicName>
, as a synonym of
<taxonomicName id="4C1F4D1BFFB8342376B06883115DFC33" authorityName="Hebard" authorityYear="1928" box="[992,1081,943,969]" class="Insecta" family="Phalangopsidae" genus="Aclodes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342376B06883115DFC33" box="[992,1081,943,969]" italics="true" pageId="2" pageNumber="570">Aclodes</emphasis>
</taxonomicName>
, something that apparently was missing in the authors contribution of 1992, where
<taxonomicName id="4C1F4D1BFFB83423778768FF115BFC17" authority="(Saussure, 1878)" baseAuthorityName="Saussure" baseAuthorityYear="1878" box="[727,1087,979,1005]" class="Insecta" family="Phalangopsidae" genus="Heterogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="B96BEA8AFFB83423778768FF161AFC17" box="[727,894,979,1005]" italics="true" pageId="2" pageNumber="570">A. crassicornis</emphasis>
(Saussure, 1878)
</taxonomicName>
is included in
<taxonomicName id="4C1F4D1BFFB8342371B168FF105EFC17" authorityName="Hebard" authorityYear="1928" box="[1249,1338,979,1005]" class="Insecta" family="Phalangopsidae" genus="Aclodes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342371B168FF105EFC17" box="[1249,1338,979,1005]" italics="true" pageId="2" pageNumber="570">Aclodes</emphasis>
</taxonomicName>
, without any formal nomenclatural act. Recently, Desutter-Grandcolas &amp; Faberon (2020) revalidated and elevated
<taxonomicName id="4C1F4D1BFFB83423706A68DB10F8FBEB" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[1338,1436,1015,1041]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="tribe" tribe="Aclodini">Aclodini</taxonomicName>
(=Aclodae) to tribal status and provided the initial opinion on the position of
<taxonomicName id="4C1F4D1BFFB83423715B6F3711C2FBCF" authorityName="Saussure" authorityYear="1874" box="[1035,1190,1051,1077]" class="Insecta" family="Phalangopsidae" genus="Heterogryllus" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB83423715B6F3711C2FBCF" box="[1035,1190,1051,1077]" italics="true" pageId="2" pageNumber="570">Heterogryllus</emphasis>
</taxonomicName>
and the non-relationship with the other
<taxonomicName id="4C1F4D1BFFB8342374236F1314B1FBA3" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[371,469,1087,1113]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="tribe" tribe="Aclodini">Aclodini</taxonomicName>
(Desutter-Grandcolas 1992). Likewise, they contrast with the
<bibRefCitation id="EF8E4B69FFB8342371CB6F131463FB84" author="Chintauan-Marquier, I. &amp; Legendre, F. &amp; Robillard, T. &amp; Hugel, S. &amp; Nel, A. &amp; Grandcolas, P. &amp; Zuccon, D. &amp; Desutter-Grandcolas, L." pageId="2" pageNumber="570" pagination="54 - 81" refId="ref6563" refString="Chintauan-Marquier, I., Legendre, F., Robillard, T., Hugel, S., Nel, A., Grandcolas, P., Zuccon, D. &amp; Desutter-Grandcolas, L. (2016) Laying the foundation of a new, phylogenetic classification of crickets (Insecta, Orthoptera): a multilocus phylogenetic approach. Cladistis, 32 (1), 54 - 81. https: // doi. org / 10.1111 / cla. 12114" type="journal article" year="2016">
Chintauan-Marquier
<emphasis id="B96BEA8AFFB8342370D86F6C15C9FB87" italics="true" pageId="2" pageNumber="570">et al</emphasis>
. (2016)
</bibRefCitation>
molecular study and other unpublished studies, highlighting that the
<taxonomicName id="4C1F4D1BFFB8342371456F4F1113FB87" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[1045,1143,1123,1149]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="tribe" tribe="Aclodini">Aclodini</taxonomicName>
are a well-defined monophyletic clade within the
<taxonomicName id="4C1F4D1BFFB8342374E46FAB1728FB5B" authorityName="Desutter-Grandcolas" authorityYear="1987" box="[436,588,1159,1185]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="subFamily" subFamily="Paragryllinae">Paragryllinae</taxonomicName>
, thus supporting the new tribal status (
<bibRefCitation id="EF8E4B69FFB8342376AE6FAB104DFB5B" author="Desutter-Grandcolas, L." box="[1022,1321,1159,1185]" pageId="2" pageNumber="570" pagination="398 - 420" refId="ref6909" refString="Desutter-Grandcolas, L. (2014) New taxa and data for Neotropical Phalangopsidae (Orthoptera, Grylloidea). Zootaxa, 3866 (3), 398 - 420. https: // doi. org / 10.11646 / zootaxa. 3866.3.5" type="journal article" year="2014">Desutter-Grandcolas 2014</bibRefCitation>
, Desutter-Grandcolas &amp; Faberon 2020). Also, they morphologically redefine
<taxonomicName id="4C1F4D1BFFB8342376C56F87169EFB3F" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[917,1018,1195,1221]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="tribe" tribe="Aclodini">Aclodini</taxonomicName>
, returning to the 1992 classification, assigning generic status to
<taxonomicName id="4C1F4D1BFFB83423749E6FE31743FB13" authorityName="Hebard" authorityYear="1928" box="[462,551,1231,1257]" class="Insecta" family="Phalangopsidae" genus="Aclodes" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB83423749E6FE31743FB13" box="[462,551,1231,1257]" italics="true" pageId="2" pageNumber="570">Aclodes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1F4D1BFFB8342377326FE31786FB13" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[610,738,1231,1257]" class="Insecta" family="Phalangopsidae" genus="Paraclodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B96BEA8AFFB8342377326FE31786FB13" box="[610,738,1231,1257]" italics="true" pageId="2" pageNumber="570">Paraclodes</emphasis>
</taxonomicName>
, and synonymizing all the subgenera proposed by
<bibRefCitation id="EF8E4B69FFB83423707C6FE31581FAF7" author="Gorochov, A. V." pageId="2" pageNumber="570" pagination="1183 - 1195" refId="ref7116" refString="Gorochov, A. V. (2007) New and little known crickets of the subfamily Phalangopsinae (Orthoptera, Gryllidae). 4. Neotropical genus Uvaroviella. Zoologicheskii Zhurnal, 86 (10), 1183 - 1195. https: // doi. org / 10.1134 / S 0013873806040087" type="journal article" year="2007">Gorochov (2007)</bibRefCitation>
. They emphasize the incomplete study of morphological characters and that monophyly was not reflected in Gorochovs proposal (Desutter-Grandcolas &amp; Faberon 2020).
</paragraph>
<paragraph id="8BA03698FFB8342375976E171789FA1C" blockId="2.[151,1437,223,1510]" pageId="2" pageNumber="570">
As noted in the history of the classification of the Aclodae /
<taxonomicName id="4C1F4D1BFFB8342376076E1716DDFAAF" authorityName="Desutter-Grandcolas" authorityYear="1992" box="[855,953,1339,1365]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="tribe" tribe="Aclodini">Aclodini</taxonomicName>
/
<taxonomicName id="4C1F4D1BFFB83423769C6E171116FAAF" box="[972,1138,1339,1365]" class="Insecta" family="Phalangopsidae" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="570" phylum="Arthropoda" rank="subTribe" subTribe="Heterogryllina">Heterogryllina</taxonomicName>
, cricket taxonomy remains in constant debate and turmoil. At the Neotropical level, it is based on the classification proposed either by Desutter-Grandcolas or Gorochov. Based on the author a tribe can be a genus or subtribe, and according to the opinion of some other authors the taxa can be in different subfamilies (
<bibRefCitation id="EF8E4B69FFB83423767C6E8B11C2FA3B" author="Cadena-Castaneda, O. J. &amp; Tijaro, M. H." box="[812,1190,1447,1473]" pageId="2" pageNumber="570" pagination="571 - 581" refId="ref5971" refString="Cadena-Castaneda, O. J. &amp; Tijaro, M. H. (2020) Studies in Colombian Ensifera and adjacent countries: New taxa of smallest field crickets (Orthoptera: Gryllidae: Gryllinae). Zootaxa, 4809 (3), 571 - 581. https: // doi. org / 10.11646 / zootaxa. 4809.3.10" type="journal article" year="2020">Cadena-Castañeda &amp; Tíjaro 2020</bibRefCitation>
,
<bibRefCitation id="EF8E4B69FFB8342371E06E8B1411FA1F" author="Cadena-Castaneda, O. J. &amp; Garcia Garcia, A." pageId="2" pageNumber="570" pagination="273 - 290" refId="ref5902" refString="Cadena-Castaneda, O. J. &amp; Garcia Garcia, A. (2020) Studies in Colombian Ensifera and adjacent countries: Gigagryllus, a new genus of giant field crickets (Orthoptera: Gryllidae) with comments on current Neotropical field crickets classification. Zootaxa, 4830 (2), 273 - 290. https: // doi. org / 10.11646 / zootaxa. 4830.2.3" type="journal article" year="2020">Cadena-Castañeda &amp; García García 2020</bibRefCitation>
, Cadena-Castañeda
<emphasis id="B96BEA8AFFB8342377086EE017F4FA1F" box="[600,656,1483,1509]" italics="true" pageId="2" pageNumber="570">et al.</emphasis>
2021b).
</paragraph>
</subSubSection>
</treatment>
</document>