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<mods:title id="36F61CD66E6D69FB3D2A9DAD5C1B7837">Morphological diagnoses of higher taxa in Ophiuroidea (Echinodermata) in support of a new classification</mods:title>
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<paragraph id="8B8836FE1F6D3027FD8BF89C7B33F884" blockId="13.[537,1051,1886,1912]" box="[537,1051,1886,1912]" pageId="13" pageNumber="14">
<heading id="D0C081921F6D3027FD8BF89C7C79F884" box="[537,849,1886,1912]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
Family
<taxonomicName id="4C374D7D1F6D3027FDE1F89C7C79F884" authority="Ljungman, 1867" authorityName="Ljungman" authorityYear="1867" box="[627,849,1886,1912]" class="Ophiuroidea" family="Ophiodermatidae" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="family">
<emphasis id="B943EAEC1F6D3027FDE1F89C7C79F884" bold="true" box="[627,849,1886,1912]" pageId="13" pageNumber="14">Ophiodermatidae</emphasis>
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<bibRefCitation id="EFA64B0F1F6D3027FCCAF89C7B33F884" author="Ljungman" box="[856,1051,1886,1912]" firstAuthor="Ljungman" pageId="13" pageNumber="14" pagination="303 - 336" refId="ref16865" refString="Ljungman A. V. 1867. Ophiuroidea viventia huc usque cognita. Ofversigt af Kungliga Vetenskaps- Akademiens Forhandlingar 1866 23: 303 - 336." type="journal article" year="1867">Ljungman, 1867</bibRefCitation>
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<heading id="D0C081921F6D3027FF2FF8637E62F847" bold="true" box="[189,330,1953,1979]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B943EAEC1F6D3027FF2FF8637E62F847" bold="true" box="[189,330,1953,1979]" pageId="13" pageNumber="14">
<typeStatus id="548C885C1F6D3027FF2FF8637FD2F847" box="[189,250,1953,1979]" pageId="13" pageNumber="14">Type</typeStatus>
genus
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836FE1F6D3027FF2FF8137B91F817" blockId="13.[189,1209,2000,2027]" box="[189,1209,2000,2027]" pageId="13" pageNumber="14">
<taxonomicName id="4C374D7D1F6D3027FF2FF8137D55F817" authority="Muller &amp; Troschel, 1840" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[189,637,2001,2027]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6D3027FF2FF8137E79F817" box="[189,337,2001,2027]" italics="true" pageId="13" pageNumber="14">Ophioderma</emphasis>
<bibRefCitation id="EFA64B0F1F6D3027FECBF8137D55F817" author="Muller &amp; Troschel" box="[345,637,2001,2027]" firstAuthor="Muller" pageId="13" pageNumber="14" pagination="326 - 330" refId="ref18138" refString="Muller J. &amp; Troschel F. H. 1840. Uber die Gattungen der Ophiuren. Archiv Fur Naturgeschichte. Berlin 6: 326 - 330." type="journal article" year="1840">Müller &amp; Troschel, 1840</bibRefCitation>
</taxonomicName>
(
<typeStatus id="548C885C1F6D3027FD1EF8137DE8F817" box="[652,704,2001,2027]" pageId="13" pageNumber="14">type</typeStatus>
species:
<taxonomicName id="4C374D7D1F6D3027FCB9F8137B9DF817" authority="(Bruzelius, 1805))" authorityName="" baseAuthorityName="Bruzelius" baseAuthorityYear="1805" box="[811,1205,2000,2027]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophioderma" kingdom="Animalia" order="Ophiurida" pageId="13" pageNumber="14" phylum="Echinodermata" rank="species" species="longicauda">
<emphasis id="B943EAEC1F6D3027FCB9F8137C68F817" box="[811,832,2001,2027]" italics="true" pageId="13" pageNumber="14">O</emphasis>
.
<emphasis id="B943EAEC1F6D3027FCDDF8137CFBF817" box="[847,979,2001,2027]" italics="true" pageId="13" pageNumber="14">longicauda</emphasis>
(
<bibRefCitation id="EFA64B0F1F6D3027FC71F8137B89F817" author="Bruzelius" box="[995,1185,2000,2027]" firstAuthor="Bruzelius" pageId="13" pageNumber="14" refId="ref14584" refString="Bruzelius N. 1805. Dissertatio sistens species cognitas asteriarum, quamr. sub praesidio D. M. and. J. Retzii, pro laurea modeste exhibet Nicolaus Bruzelius. Lundae." type="book" year="1805">Bruzelius, 1805</bibRefCitation>
))
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.
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<paragraph id="8B8836FE1F6E3024FF2FFEC97E4CFED9" blockId="14.[189,356,267,293]" box="[189,356,267,293]" pageId="14" pageNumber="15">
<heading id="D0C081921F6E3024FF2FFEC97E4CFED9" bold="true" box="[189,356,267,293]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B943EAEC1F6E3024FF2FFEC97E4CFED9" bold="true" box="[189,356,267,293]" pageId="14" pageNumber="15">Other genera</emphasis>
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<taxonomicName id="4C374D7D1F6E3024FF2FFEF87D49FEA8" authority="H. L. Clark, 1909 b" authorityName="H. L. Clark" authorityYear="1909" box="[189,609,314,341]" class="Ophiuroidea" family="Ophiodermatidae" genus="Bathypectinura" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FF2FFEF87E5AFEA8" box="[189,370,314,340]" italics="true" pageId="14" pageNumber="15">Bathypectinura</emphasis>
H.L.
<bibRefCitation id="EFA64B0F1F6E3024FE52FEF87D49FEA8" author="Clark" box="[448,609,314,340]" firstAuthor="Clark" pageId="14" pageNumber="15" pagination="109 - 135" refId="ref14933" refString="Clark H. L. 1909 b. Notes on some Australian and Indo-Pacific echinoderms. Bulletin of the Museum of Comparative Zoology at Harvard University 52: 109 - 135." type="journal article" year="1909" yearSuffix="b">Clark, 1909b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FDE0FEF87CC5FEA8" authority="H. L. Clark, 1909 b" authorityName="H. L. Clark" authorityYear="1909" box="[626,1005,314,341]" class="Ophiuroidea" family="Ophiodermatidae" genus="Cryptopelta" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FDE0FEF87DD6FEA8" box="[626,766,314,340]" italics="true" pageId="14" pageNumber="15">Cryptopelta</emphasis>
H.L.
<bibRefCitation id="EFA64B0F1F6E3024FCDFFEF87CC5FEA8" author="Clark" box="[845,1005,314,340]" firstAuthor="Clark" pageId="14" pageNumber="15" pagination="109 - 135" refId="ref14933" refString="Clark H. L. 1909 b. Notes on some Australian and Indo-Pacific echinoderms. Bulletin of the Museum of Comparative Zoology at Harvard University 52: 109 - 135." type="journal article" year="1909" yearSuffix="b">Clark, 1909b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FC6DFEF87A59FEA8" authority="H. L. Clark, 1913" authorityName="H. L. Clark" authorityYear="1913" box="[1023,1393,314,341]" class="Ophiuroidea" family="Ophiodermatidae" genus="Diopederma" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FC6DFEF87BBAFEA8" box="[1023,1170,314,340]" italics="true" pageId="14" pageNumber="15">Diopederma</emphasis>
H.L.
<bibRefCitation id="EFA64B0F1F6E3024FB72FEF87A59FEA8" author="Clark" box="[1248,1393,314,340]" firstAuthor="Clark" pageId="14" pageNumber="15" pagination="185 - 236" refId="ref14998" refString="Clark H. L. 1913. Scientific Results of the Expedition to the Gulf of California in charge of C. H. Townsend, by the U. S. Fisheries Steamship ' Albatross' in 1911. Commander C. H. Burrage, U. S. N., Commanding. V. Echinoderms from Lower California, with descriptions of new species. Bulletin of the American Museum of Natural History 32: 185 - 236." type="journal article" year="1913">Clark, 1913</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FF2FFE9C7D7EFE85" authority="Ljungman, 1872" authorityName="Ljungman" authorityYear="1872" box="[189,598,350,377]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiarachnella" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FF2FFE9C7E5FFE84" box="[189,375,350,376]" italics="true" pageId="14" pageNumber="15">Ophiarachnella</emphasis>
<bibRefCitation id="EFA64B0F1F6E3024FE1AFE9C7D7EFE85" author="Ljungman" box="[392,598,350,377]" firstAuthor="Ljungman" pageId="14" pageNumber="15" pagination="615 - 658" refId="ref16891" refString="Ljungman A. V. 1872. Forteckning ofver uti Vestindien af Dr A. Goes samt under korvetten Josefinas expedition i Atlantiska Oceanen samlade Ophiurider. Ofversigt af Kungliga Vetenskapsakademiens Forhandlingar 1871 6: 615 - 658." type="journal article" year="1872">Ljungman, 1872</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FDFEFE9C7C97FE84" authority="Grube, 1868" authorityName="Grube" authorityYear="1868" box="[620,959,350,376]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiochasma" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FDFEFE9C7C27FE84" box="[620,783,350,376]" italics="true" pageId="14" pageNumber="15">Ophiochasma</emphasis>
<bibRefCitation id="EFA64B0F1F6E3024FCB3FE9C7C97FE84" author="Grube" box="[801,959,350,376]" firstAuthor="Grube" pageId="14" pageNumber="15" pagination="44 - 45" refId="ref15958" refString="Grube A. E. 1868. Uber einige seltenere oder neue Ophiuriden. Jahresberichte der schlesischen Gesellschaft fur vaterlandische Kultur 45: 44 - 45." type="journal article" year="1868">Grube, 1868</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FC44FE9C7A5AFE85" authority="H. L. Clark, 1915 a" authorityName="H. L. Clark" authorityYear="1915" box="[982,1394,350,377]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiocormus" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FC44FE9C7B5EFE84" box="[982,1142,350,376]" italics="true" pageId="14" pageNumber="15">Ophiocormus</emphasis>
H.L.
<bibRefCitation id="EFA64B0F1F6E3024FB5CFE9C7A5AFE85" author="Clark" box="[1230,1394,350,377]" firstAuthor="Clark" pageId="14" pageNumber="15" pagination="163 - 376" refId="ref15107" refString="Clark H. L. 1915 a. Catalogue of recent Ophiurans. Memoirs of the Museum of Comparative Zoology 25: 163 - 376." type="journal article" year="1915" yearSuffix="a">Clark, 1915a</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FF2FFE417D1AFE61" authority="H. L. Clark, 1915 b" authorityName="H. L. Clark" authorityYear="1915" box="[189,562,386,413]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiocryptus" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FF2FFE417E75FE61" box="[189,349,387,413]" italics="true" pageId="14" pageNumber="15">Ophiocryptus</emphasis>
H.L.
<bibRefCitation id="EFA64B0F1F6E3024FE08FE417D1AFE61" author="Clark" box="[410,562,386,413]" firstAuthor="Clark" pageId="14" pageNumber="15" pagination="64 - 66" refId="ref15133" refString="Clark H. L. 1915 b. A remarkable new brittle star. Journal of Entomology and Zoology 7: 64 - 66." type="journal article" year="1915" yearSuffix="b">Clark, 1915b</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FDA9FE417C5DFE61" authority="Koehler, 1930" authorityName="Koehler" authorityYear="1930" box="[571,885,387,413]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiocypris" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FDA9FE417DE4FE61" box="[571,716,387,413]" italics="true" pageId="14" pageNumber="15">Ophiocypris</emphasis>
<bibRefCitation id="EFA64B0F1F6E3024FD42FE417C5DFE61" author="Koehler" box="[720,885,387,413]" firstAuthor="Koehler" pageId="14" pageNumber="15" pagination="1 - 295" refId="ref16630" refString="Koehler R. 1930. Ophiures recueillies par le Docteur Th. Mortensen dans les Mers d'Australie et dans l'Archipel Malais. Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 16. LIV. Videnskabelige Meddelelser fra Dansk naturhistorisk Forening 89: 1 - 295." type="journal article" year="1930">Koehler, 1930</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FCECFE417B9FFE61" authority="Koehler, 1904" authorityName="Koehler" authorityYear="1904" box="[894,1207,387,413]" class="Ophiuroidea" family="Ophiolepididae" genus="Ophiomidas" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FCECFE417B25FE61" box="[894,1037,387,413]" italics="true" pageId="14" pageNumber="15">Ophiomidas</emphasis>
<bibRefCitation id="EFA64B0F1F6E3024FB80FE417B9FFE61" author="Koehler" box="[1042,1207,387,413]" firstAuthor="Koehler" pageId="14" pageNumber="15" pagination="1 - 36" refId="ref16336" refString="Koehler R. 1904. Ophiures de mer profonde. Siboga Expeditie Monographs 45 a: 1 - 176, pls 1 - 36." type="book chapter" year="1904">Koehler, 1904</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F6E3024FB52FE417E75FE3D" authority="Lutken, 1869" authorityName="Lutken" authorityYear="1869" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiopsammus" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FB52FE417A5FFE61" box="[1216,1399,387,413]" italics="true" pageId="14" pageNumber="15">Ophiopsammus</emphasis>
<bibRefCitation id="EFA64B0F1F6E3024FF2FFE657E75FE3D" author="Lutken" box="[189,349,423,449]" firstAuthor="Lutken" pageId="14" pageNumber="15" pagination="24 - 109" refId="ref17121" refString="Lutken C. F. 1869. Additamenta ad historiam Ophiuridarum. 3. Beskrivende og kritiske Bidrag til Kundskab om Slangestjernerne. Det kongelige danske Videnskabernes Selskabs Skrifter 5: 24 - 109." type="journal article" year="1869">Lütken, 1869</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D65751F6E3024FF2FFE2B7C9EFD63" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8B8836FE1F6E3024FF2FFE2B7E1EFDFF" blockId="14.[189,310,489,515]" box="[189,310,489,515]" pageId="14" pageNumber="15">
<heading id="D0C081921F6E3024FF2FFE2B7E1EFDFF" bold="true" box="[189,310,489,515]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B943EAEC1F6E3024FF2FFE2B7E1EFDFF" bold="true" box="[189,310,489,515]" pageId="14" pageNumber="15">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836FE1F6E3024FF2FFDDB7C9EFD63" blockId="14.[189,1399,537,671]" pageId="14" pageNumber="15">Dorsal disc covered with granules. Dental plate fragmented, with finely meshed stereom, tooth sockets as broadly oval, not penetrating depressions with swollen borders, small tooth cluster on ventral end. Lateral arm plate with two proximal spurs, visible on external and internal surface.Arm spine articulation with weak sigmoidal fold, sunken in notches of distal plate edge.</paragraph>
</subSubSection>
<subSubSection id="C32D65751F6E3024FF2FFD0A7CF8FCC9" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8B8836FE1F6E3024FF2FFD0A7E07FD1E" blockId="14.[189,303,712,738]" box="[189,303,712,738]" pageId="14" pageNumber="15">
<heading id="D0C081921F6E3024FF2FFD0A7E07FD1E" bold="true" box="[189,303,712,738]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
<emphasis id="B943EAEC1F6E3024FF2FFD0A7E07FD1E" bold="true" box="[189,303,712,738]" pageId="14" pageNumber="15">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836FE1F6E3024FF2FFD357CF8FCC9" blockId="14.[189,1398,759,822]" pageId="14" pageNumber="15">
<taxonomicName id="4C374D7D1F6E3024FF2FFD357E5FFCED" authorityName="Ljungman" authorityYear="1872" box="[189,375,759,785]" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiarachnella" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F6E3024FF2FFD357E5FFCED" box="[189,375,759,785]" italics="true" pageId="14" pageNumber="15">Ophiarachnella</emphasis>
</taxonomicName>
is polyphyletic and requires revision. The
<typeStatus id="548C885C1F6E3024FCF8FD357CB5FCED" box="[874,925,759,785]" pageId="14" pageNumber="15">type</typeStatus>
species
<taxonomicName id="4C374D7D1F6E3024FB92FD357FD2FCC9" authority="Muller &amp; Troschel, 1842" authorityName="Muller &amp; Troschel" authorityYear="1842" class="Ophiuroidea" family="Ophiodermatidae" genus="Ophiarachnella" kingdom="Animalia" order="Ophiurida" pageId="14" pageNumber="15" phylum="Echinodermata" rank="species" species="gorgonia">
<emphasis id="B943EAEC1F6E3024FB92FD357BA6FCED" box="[1024,1166,759,785]" italics="true" pageId="14" pageNumber="15">O. gorgonia</emphasis>
<bibRefCitation id="EFA64B0F1F6E3024FB07FD357FD2FCC9" author="Muller &amp; Troschel" firstAuthor="Muller" pageId="14" pageNumber="15" refId="ref18166" refString="Muller J. H. &amp; Troschel F. H. 1842. System der Asteriden. Vieweg &amp; Sohn, Braunschweig." type="book" year="1842">Müller &amp; Troschel, 1842</bibRefCitation>
</taxonomicName>
, however, is a confirmed ophiodermatid (
<bibRefCitation id="EFA64B0F1F6E3024FD73FCD97CECFCC9" author="O'Hara" box="[737,964,795,821]" etAl="et al." firstAuthor="O'Hara" pageId="14" pageNumber="15" pagination="415 - 430" refId="ref18267" refString="O'Hara T. D., Hugall A. F., Thuy B., Stohr S. &amp; Martynov A. V. 2017. Restructuring higher taxonomy using broad-scale phylogenomics: The living Ophiuroidea. Molecular Phylogenetics and Evolution 107: 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
OHara
<emphasis id="B943EAEC1F6E3024FCD0FCDE7C55FCC9" box="[834,893,795,821]" italics="true" pageId="14" pageNumber="15">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>