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<document id="B2227F3D6EDA8A58C5224EF15F871D95" ID-CLB-Dataset="22339" ID-DOI="10.5852/ejt.2018.416" ID-GBIF-Dataset="2eebb037-4ab1-433c-a9f6-8fe319e9fda9" ID-Zenodo-Dep="3806109" ID-ZooBank="AD094812-5768-43E9-BCC2-9226E69F0820" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588792288471" checkinUser="valdenar" docAuthor="Stöhr, Sabine, Hugall, Andrew F., Thuy, Ben &amp; Martynov, Alexander" docDate="2018" docId="039E87E81F683023FEA3F9F87D53FD66" docLanguage="en" docName="ejt-416_ohara_stohr_hugall.pdf.imf" docOrigin="European Journal of Taxonomy 416" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ophiuridae Muller &amp; Troschel 1840" docType="treatment" docVersion="9" lastPageNumber="10" masterDocId="FFA7FF901F60302AFF92FFC27F28FFFC" masterDocTitle="Morphological diagnoses of higher taxa in Ophiuroidea (Echinodermata) in support of a new classification" masterLastPageNumber="35" masterPageNumber="1" pageNumber="9" updateTime="1698838815030" updateUser="ExternalLinkService">
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<mods:title id="F1E0BF40949F84C730222D1206AB2A10">Morphological diagnoses of higher taxa in Ophiuroidea (Echinodermata) in support of a new classification</mods:title>
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<mods:namePart id="4EF8B731981139AF313BD927E276B886">Stöhr, Sabine</mods:namePart>
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<mods:namePart id="99DABDA29FA9DE0EF9C1E1F8C5CA0D74">Thuy, Ben</mods:namePart>
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<treatment id="039E87E81F683023FEA3F9F87D53FD66" ID-DOI="http://doi.org/10.5281/zenodo.3816259" ID-GBIF-Taxon="163971273" ID-Zenodo-Dep="3816259" LSID="urn:lsid:plazi:treatment:039E87E81F683023FEA3F9F87D53FD66" httpUri="http://treatment.plazi.org/id/039E87E81F683023FEA3F9F87D53FD66" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="C32D65751F683022FEA3F9F87A2AF9A8" box="[305,1282,1593,1620]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8B8836FE1F683022FEA3F9F87A2AF9A8" blockId="8.[305,1282,1593,1620]" box="[305,1282,1593,1620]" pageId="8" pageNumber="9">
<heading id="D0C081921F683022FEA3F9F87A2AF9A8" box="[305,1282,1593,1620]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
Family
<taxonomicName id="4C374D7D1F683022FE19F9FB7C62F9A8" ID-CoL="7NK6B" authority="Muller &amp; Troschel, 1840" authorityName="Muller &amp; Troschel" authorityYear="1840" box="[395,842,1593,1620]" class="Ophiuroidea" family="Ophiuridae" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="family">
<emphasis id="B943EAEC1F683022FE19F9FB7D34F9AF" bold="true" box="[395,540,1593,1619]" pageId="8" pageNumber="9">Ophiuridae</emphasis>
<bibRefCitation id="EFA64B0F1F683022FDB6F9F87C62F9A8" author="Muller &amp; Troschel" box="[548,842,1594,1620]" firstAuthor="Muller" pageId="8" pageNumber="9" pagination="326 - 330" refId="ref18138" refString="Muller J. &amp; Troschel F. H. 1840. Uber die Gattungen der Ophiuren. Archiv Fur Naturgeschichte. Berlin 6: 326 - 330." type="journal article" year="1840">Müller &amp; Troschel, 1840</bibRefCitation>
</taxonomicName>
, restricted
<emphasis id="B943EAEC1F683022FC59F9F97B24F9A8" box="[971,1036,1595,1620]" italics="true" pageId="8" pageNumber="9">sensu</emphasis>
<bibRefCitation id="EFA64B0F1F683022FB81F9F87A2AF9A8" author="O'Hara" box="[1043,1282,1594,1620]" etAl="et al." firstAuthor="O'Hara" pageId="8" pageNumber="9" pagination="415 - 430" refId="ref18267" refString="O'Hara T. D., Hugall A. F., Thuy B., Stohr S. &amp; Martynov A. V. 2017. Restructuring higher taxonomy using broad-scale phylogenomics: The living Ophiuroidea. Molecular Phylogenetics and Evolution 107: 415 - 430. https: // doi. org / 10.1016 / j. ympev. 2016.12.006" type="journal article" year="2017">
OHara
<emphasis id="B943EAEC1F683022FBE0F9F97B85F9A8" box="[1138,1197,1594,1620]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2017)
</bibRefCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D65751F683022FF2FF9B97E17F8AB" pageId="8" pageNumber="9" type="type_taxon">
<paragraph id="8B8836FE1F683022FF2FF9B97E62F969" blockId="8.[189,330,1659,1685]" box="[189,330,1659,1685]" pageId="8" pageNumber="9">
<heading id="D0C081921F683022FF2FF9B97E62F969" bold="true" box="[189,330,1659,1685]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B943EAEC1F683022FF2FF9B97E62F969" bold="true" box="[189,330,1659,1685]" pageId="8" pageNumber="9">
<typeStatus id="548C885C1F683022FF2FF9B97FD2F969" box="[189,250,1659,1685]" pageId="8" pageNumber="9">Type</typeStatus>
genus
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836FE1F683022FF2FF9687CC2F939" blockId="8.[189,1002,1706,1733]" box="[189,1002,1706,1733]" pageId="8" pageNumber="9">
<taxonomicName id="4C374D7D1F683022FF2FF9687EF2F938" ID-CoL="7PNFY" authority="Lamarck, 1801" authorityName="Lamarck" authorityYear="1801" box="[189,474,1706,1732]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F683022FF2FF9687E08F938" box="[189,288,1706,1732]" italics="true" pageId="8" pageNumber="9">Ophiura</emphasis>
<bibRefCitation id="EFA64B0F1F683022FEBAF9687EF2F938" author="Lamarck" box="[296,474,1706,1732]" firstAuthor="Lamarck" pageId="8" pageNumber="9" refId="ref16679" refString="Lamarck J. B. 1801. Systeme des animaux sans vertebres; ou tableau general des classes, des orres et des genres de ces animaux. Deterville, Paris." type="book" year="1801">Lamarck, 1801</bibRefCitation>
</taxonomicName>
(
<typeStatus id="548C885C1F683022FE7BF9687D35F938" box="[489,541,1706,1732]" pageId="8" pageNumber="9">type</typeStatus>
species:
<taxonomicName id="4C374D7D1F683022FD1AF9687CCEF939" ID-CoL="74PFS" authority="(Linneaus, 1758))" baseAuthorityName="Linneaus" baseAuthorityYear="1758" box="[648,998,1706,1733]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="species" species="ophiura">
<emphasis id="B943EAEC1F683022FD1AF9687DB5F938" box="[648,669,1706,1732]" italics="true" pageId="8" pageNumber="9">O</emphasis>
.
<emphasis id="B943EAEC1F683022FD3EF9687C21F938" box="[684,777,1706,1732]" italics="true" pageId="8" pageNumber="9">ophiura</emphasis>
(Linneaus, 1758))
</taxonomicName>
.
</paragraph>
<paragraph id="8B8836FE1F683022FF2FF9297E4CF8FA" blockId="8.[189,356,1771,1798]" box="[189,356,1771,1798]" pageId="8" pageNumber="9">
<heading id="D0C081921F683022FF2FF9297E4CF8FA" bold="true" box="[189,356,1771,1798]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B943EAEC1F683022FF2FF9297E4CF8FA" bold="true" box="[189,356,1771,1798]" pageId="8" pageNumber="9">Other genera</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836FE1F683022FF2FF8D87E17F8AB" blockId="8.[189,1399,1818,1879]" pageId="8" pageNumber="9">
<taxonomicName id="4C374D7D1F683022FF2FF8D87D1FF8C8" ID-CoL="7PQWX" authority="H. L. Clark, 1928" authorityName="H. L. Clark" authorityYear="1928" box="[189,567,1818,1845]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiocrossota" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F683022FF2FF8D87E41F8C8" box="[189,361,1818,1844]" italics="true" pageId="8" pageNumber="9">Ophiocrossota</emphasis>
H.L.
<bibRefCitation id="EFA64B0F1F683022FE39F8D87D1FF8C8" author="Clark" box="[427,567,1818,1844]" firstAuthor="Clark" pageId="8" pageNumber="9" pagination="361 - 428" refId="ref15256" refString="Clark H. L. 1928. The sea-lilies, sea-stars, brittle-stars and sea-urchins of the South Australian Museum. Records of the South Australian Museum 3: 361 - 428." type="journal article" year="1928">Clark, 1928</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F683022FDD1F8D87C4AF8C9" ID-CoL="7PNCX" authority="Lutken, 1855" authorityName="Lutken" authorityYear="1855" box="[579,866,1818,1845]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiocten" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F683022FDD1F8D87D95F8C8" box="[579,701,1818,1844]" italics="true" pageId="8" pageNumber="9">Ophiocten</emphasis>
<bibRefCitation id="EFA64B0F1F683022FD56F8D87C4AF8C9" author="Lutken" box="[708,866,1818,1845]" firstAuthor="Lutken" pageId="8" pageNumber="9" pagination="95 - 104" refId="ref17009" refString="Lutken C. F. 1855. Bidrag til Kundskab om Slangestjernerne. I. Forelobig Oversigt over Gronlandshavets Ophiurer. Videnskabelige Meddelelser fra Dansk Naturhistorisk Forening, 1854: 95 - 104." type="journal article" year="1855">Lütken, 1855</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F683022FCFFF8D87BCAF8C9" ID-CoL="7PQWY" authority="Tyler et al., 1995" authorityName="Tyler" authorityYear="1995" box="[877,1250,1818,1845]" class="Ophiuroidea" family="Ophiuridae" genus="Ophioctenella" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F683022FCFFF8D87B3BF8C8" box="[877,1043,1818,1844]" italics="true" pageId="8" pageNumber="9">Ophioctenella</emphasis>
<bibRefCitation id="EFA64B0F1F683022FB8BF8D87BCAF8C9" author="Tyler" box="[1049,1250,1818,1845]" etAl="et al." firstAuthor="Tyler" pageId="8" pageNumber="9" pagination="977 - 986" refId="ref19109" refString="Tyler P. A., Paterson G. L .., Sibuet M., Guille A., Murtons B. J. &amp; Segonzac M. 1995. A new genus of ophiuroid (Echinodermata: Ophiuroidea) from hydrothermal mounds along the Mid-Atlantic Ridge. Journal of the Marine Biological Association U. K. 75: 977 - 986." type="journal article" year="1995">
Tyler
<emphasis id="B943EAEC1F683022FBCFF8D97BBFF8C8" box="[1117,1175,1818,1844]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
, 1995
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C374D7D1F683022FB7CF8D87E13F8AB" ID-CoL="7PNDN" authority="Bell, 1902" authorityName="Bell" authorityYear="1902" class="Ophiuroidea" family="Ophiuridae" genus="Ophionotus" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F683022FB7CF8D87A5FF8C8" box="[1262,1399,1818,1844]" italics="true" pageId="8" pageNumber="9">Ophionotus</emphasis>
<bibRefCitation id="EFA64B0F1F683022FF2FF8FF7E13F8AB" author="Bell" box="[189,315,1853,1879]" firstAuthor="Bell" pageId="8" pageNumber="9" pagination="214 - 220" refId="ref14405" refString="Bell F. J. 1902. Echinodermata. Reports on the collections of Natural History made in the Antarctic regions during the voyage of the &quot; Southern Cross 8: 214 - 220, 2 pls." type="book chapter" year="1902">Bell, 1902</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D65751F683023FF2FF8BD7B0CFE93" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="diagnosis">
<paragraph id="8B8836FE1F683022FF2FF8BD7E1EF865" blockId="8.[189,310,1919,1945]" box="[189,310,1919,1945]" pageId="8" pageNumber="9">
<heading id="D0C081921F683022FF2FF8BD7E1EF865" bold="true" box="[189,310,1919,1945]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B943EAEC1F683022FF2FF8BD7E1EF865" bold="true" box="[189,310,1919,1945]" pageId="8" pageNumber="9">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836FE1F683023FF2FF86F7B0CFE93" blockId="8.[189,1398,1965,2027]" lastBlockId="9.[189,1399,267,367]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Flat disc, usually fully scaled but naked in some species. Genital papillae and arm combs present, although combs in
<taxonomicName id="4C374D7D1F683022FE0CF8137D0FF817" authorityName="Bell" authorityYear="1902" box="[414,551,2001,2027]" class="Ophiuroidea" family="Ophiuridae" genus="Ophionotus" kingdom="Animalia" order="Ophiurida" pageId="8" pageNumber="9" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F683022FE0CF8137D0FF817" box="[414,551,2001,2027]" italics="true" pageId="8" pageNumber="9">Ophionotus</emphasis>
</taxonomicName>
reduced. Dental plate entire, sockets small round holes, not perforating, with low borders; single, offset column, ventral cluster. Lateral arm plates commonly divided in two halves by central, distalwards-pointing tentacle notch. Arm spine articulation with large, irregularly round muscle opening on an elevation, nerve opening at base of elevation.
</paragraph>
</subSubSection>
<subSubSection id="C32D65751F693023FF2FFE5A7D53FD66" pageId="9" pageNumber="10" type="discussion">
<paragraph id="8B8836FE1F693023FF2FFE5A7E07FE4E" blockId="9.[189,303,408,434]" box="[189,303,408,434]" pageId="9" pageNumber="10">
<heading id="D0C081921F693023FF2FFE5A7E07FE4E" bold="true" box="[189,303,408,434]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B943EAEC1F693023FF2FFE5A7E07FE4E" bold="true" box="[189,303,408,434]" pageId="9" pageNumber="10">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836FE1F693023FF2FFE0A7D53FD66" blockId="9.[189,1399,456,667]" pageId="9" pageNumber="10">
<taxonomicName id="4C374D7D1F693023FF2FFE0A7E08FE1E" authorityName="Lamarck" authorityYear="1801" box="[189,288,456,482]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F693023FF2FFE0A7E08FE1E" box="[189,288,456,482]" italics="true" pageId="9" pageNumber="10">Ophiura</emphasis>
</taxonomicName>
may be polyphyletic and needs to be revised (e.g.,
<taxonomicName id="4C374D7D1F693023FCE5FE0B7BB2FE1F" authority="(Ayres, 1852)" baseAuthorityName="Ayres" baseAuthorityYear="1852" box="[887,1178,456,483]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiura" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="species" species="robusta">
<emphasis id="B943EAEC1F693023FCE5FE0B7CA4FE1F" box="[887,908,457,483]" italics="true" pageId="9" pageNumber="10">O</emphasis>
.
<emphasis id="B943EAEC1F693023FC09FE0A7CDCFE1E" box="[923,1012,456,482]" italics="true" pageId="9" pageNumber="10">robusta</emphasis>
(Ayres, 1852)
</taxonomicName>
has hooked spines on the distal arm, a character of
<taxonomicName id="4C374D7D1F693023FDDDFE2F7C81FDFB" authority="Perrier, 1893" authorityName="Perrier" authorityYear="1893" box="[591,937,493,519]" class="Ophiuroidea" family="Ophiopyrgidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="family">
Ophiopyrgidae
<bibRefCitation id="EFA64B0F1F693023FC98FE2F7C81FDFB" author="Perrier" box="[778,937,493,519]" firstAuthor="Perrier" pageId="9" pageNumber="10" pagination="781 - 864" refId="ref18476" refString="Perrier E. 1893. In: Traite de zoologie. 3 ser. Echinodermata: 781 - 864. Masson, Paris." type="book chapter" year="1893">Perrier, 1893</bibRefCitation>
</taxonomicName>
). The subgenus
<taxonomicName id="4C374D7D1F693023FBE3FE2F7FD0FDD0" authority="Hertz, 1927" authorityName="Hertz" authorityYear="1927" class="Ophiuroidea" family="Ophiopyrgidae" genus="Ophiuroglypha" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F693023FBE3FE2F7A0BFDFB" box="[1137,1315,493,519]" italics="true" pageId="9" pageNumber="10">Ophiuroglypha</emphasis>
<bibRefCitation id="EFA64B0F1F693023FABCFE2F7FD0FDD0" author="Hertz" firstAuthor="Hertz" pageId="9" pageNumber="10" pagination="59 - 122" refId="ref16081" refString="Hertz M. 1927. Die Ophiuroiden der deutschen Tiefsee-Expedition. 1. Chilophiurida Mats. (Ophiolepididae: Ophioleucidae: Ophiodermatidae: Ophiocomidae). Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899 22: 59 - 122, pls 6 - 9." type="journal article" year="1927">Hertz, 1927</bibRefCitation>
</taxonomicName>
is herein raised to genus-rank and referred to the
<taxonomicName id="4C374D7D1F693023FCE1FDD07B0EFDD0" authorityName="Perrier" authorityYear="1893" box="[883,1062,530,556]" class="Ophiuroidea" family="Ophiopyrgidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="family">Ophiopyrgidae</taxonomicName>
. The available genus name
<taxonomicName id="4C374D7D1F693023FF2FFDF47EC3FDAC" authority="Fell, 1961 b" authorityName="Fell" authorityYear="1961" box="[189,491,566,592]" class="Ophiuroidea" family="Ophiopyrgidae" genus="Glaciacantha" higherTaxonomySource="CoL" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F693023FF2FFDF47E75FDAC" box="[189,349,566,592]" italics="true" pageId="9" pageNumber="10">Glaciacantha</emphasis>
<bibRefCitation id="EFA64B0F1F693023FEF7FDF47EC3FDAC" author="Fell" box="[357,491,566,592]" firstAuthor="Fell" pageId="9" pageNumber="10" pagination="9 - 79" refId="ref15767" refString="Fell H. B. 1961 b. The Fauna of the Ross Sea. Part 1, Ophiuroidea. Memoirs of the New Zealand Oceanographic Institute 18: 9 - 79, 19 pls." type="book chapter" year="1961" yearSuffix="b">Fell, 1961b</bibRefCitation>
</taxonomicName>
is used herein to contain two Southern Ocean species of
<taxonomicName id="4C374D7D1F693023FB00FDF47A24FDAC" authorityName="Lutken" authorityYear="1855" box="[1170,1292,566,592]" class="Ophiuroidea" family="Ophiuridae" genus="Ophiocten" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="genus">
<emphasis id="B943EAEC1F693023FB00FDF47A24FDAC" box="[1170,1292,566,592]" italics="true" pageId="9" pageNumber="10">Ophiocten</emphasis>
</taxonomicName>
(
<emphasis id="B943EAEC1F693023FA8CFDF47A5EFDAC" box="[1310,1398,566,592]" italics="true" pageId="9" pageNumber="10">dubium</emphasis>
<bibRefCitation id="EFA64B0F1F693023FF2FFD997E4EFD89" author="Koehler" box="[189,358,603,629]" firstAuthor="Koehler" pageId="9" pageNumber="10" pagination="222 - 231" refId="ref16304" refString="Koehler R. 1901. Note preliminaire sur quelques ophiures nouvelles provenant des campagnes de la Princesse Alice. Bulletin de la Societe zoologique de France 26: 222 - 231." type="journal article" year="1901">Koehler, 1901</bibRefCitation>
and
<emphasis id="B943EAEC1F693023FE30FD997D00FD89" box="[418,552,603,629]" italics="true" pageId="9" pageNumber="10">doederleini</emphasis>
<bibRefCitation id="EFA64B0F1F693023FDA2FD9E7D96FD89" author="Hertz" box="[560,702,603,630]" firstAuthor="Hertz" pageId="9" pageNumber="10" pagination="59 - 122" refId="ref16081" refString="Hertz M. 1927. Die Ophiuroiden der deutschen Tiefsee-Expedition. 1. Chilophiurida Mats. (Ophiolepididae: Ophioleucidae: Ophiodermatidae: Ophiocomidae). Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot; 1898 - 1899 22: 59 - 122, pls 6 - 9." type="journal article" year="1927">Hertz, 1927</bibRefCitation>
) with raised disc plates and hooked distal arm spines, and also transferred to the
<taxonomicName id="4C374D7D1F693023FE51FD427D5EFD66" authorityName="Perrier" authorityYear="1893" box="[451,630,640,666]" class="Ophiuroidea" family="Ophiopyrgidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ophiurida" pageId="9" pageNumber="10" phylum="Echinodermata" rank="family">Ophiopyrgidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>