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<document id="07A969B45ED6EE19F8C40141F5EBB355" ID-DOI="10.5252/cryptogamie-bryologie2021v42a9" ID-ISSN="1776-0992" ID-Zenodo-Dep="12215190" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719004266022" checkinUser="felipe" docAuthor="Printarakul, Narin, Jampeetong, Arunothai, Pongkawong, Udon, Sukkharak, Phiangphak, Kraichak, Ekaphan, Adulkittichai, Kanonrat &amp; Chantanaorrapint, Sahut" docDate="2021" docId="039E87E2FFC1FFDDFEE11A2148A3FEF6" docLanguage="en" docName="CryptogamieBryologie.42.9.143-148.pdf" docOrigin="Cryptogamie, Bryologie 20 (9)" docSource="http://dx.doi.org/10.5252/cryptogamie-bryologie2021v42a9" docStyle="DocumentStyle:3F4A000F5B047ABA0EAB2943AA9373AE.3:CryptogamieBryologie.2019-.journal_article" docStyleId="3F4A000F5B047ABA0EAB2943AA9373AE" docStyleName="CryptogamieBryologie.2019-.journal_article" docStyleVersion="3" docTitle="Koponobryum papillosum Printarakul &amp; Chantanaorr. 2021, sp. nov." docType="treatment" docVersion="2" lastPageNumber="148" masterDocId="FFA7FF9AFFC2FFDAFFDB1C13484FFFE4" masterDocTitle="Koponobryum papillosum Printarakul &amp; Chantanaorr., sp. nov. (Pottiaceae, Bryophyta), a new moss species from northern Thailand" masterLastPageNumber="148" masterPageNumber="143" pageNumber="144" updateTime="1719018384683" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="F2F55720BC9090806FF9DCEA78EC3184">Koponobryum papillosum Printarakul &amp; Chantanaorr., sp. nov. (Pottiaceae, Bryophyta), a new moss species from northern Thailand</mods:title>
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<mods:namePart id="B2C63553EFBDA471E25329E8909C9B28">Printarakul, Narin</mods:namePart>
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<mods:namePart id="E3FF95F1586B604B877E8728B8295E73">Jampeetong, Arunothai</mods:namePart>
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<mods:namePart id="342058BBE5DBB56DAB7E00A6B303D6E2">Pongkawong, Udon</mods:namePart>
<mods:affiliation id="470F662B5B4132EBEB87922E55FE278D">Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200 (Thailand)</mods:affiliation>
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<mods:namePart id="898EC8CA752D6BE4C0DF617DCE1E86FC">Sukkharak, Phiangphak</mods:namePart>
<mods:affiliation id="7A94C0C5D8F069575EEDAD6DA2DD9E90">Department of Biology, Faculty of Science, Burapha University, Chonburi 20131 (Thailand)</mods:affiliation>
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<mods:namePart id="F6799517660A36DBFFE0D052EFD19D1B">Kraichak, Ekaphan</mods:namePart>
<mods:affiliation id="45690B692BDF84D4521FB7FED6A5D83E">Department of Botany, Faculty of Science, Kasetsart University, Chatuchak, Bangkok 10900 (Thailand)</mods:affiliation>
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<mods:namePart id="5C7C32E89ADA77DE4F21256D91A8ECEB">Adulkittichai, Kanonrat</mods:namePart>
<mods:affiliation id="6C92BF0D2FFF23EFD624C96CFC995DC9">Research and development department, Chiangmai Vanussanun Co., Ltd., Chiang Mai 50000, (Thailand)</mods:affiliation>
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<mods:namePart id="47501B6B22EF229DFF03D95BCECB4FE4">Chantanaorrapint, Sahut</mods:namePart>
<mods:affiliation id="58AD6B94BD875CB92EC80DA855F0ABB8">SU Herbarium, Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla 90110 (Thailand) sahut. c. @ psu. ac. th (corresponding author)</mods:affiliation>
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<mods:title id="FF71BE67F28D214509C5D662BE4D5867">Cryptogamie, Bryologie</mods:title>
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<mods:date id="320EBCFD9942A889EA6DD1BB7A61B5BA">2021</mods:date>
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<treatment id="039E87E2FFC1FFDDFEE11A2148A3FEF6" LSID="urn:lsid:plazi:treatment:039E87E2FFC1FFDDFEE11A2148A3FEF6" httpUri="http://treatment.plazi.org/id/039E87E2FFC1FFDDFEE11A2148A3FEF6" lastPageId="7" lastPageNumber="148" pageId="3" pageNumber="144">
<subSubSection id="C32D657FFFC1FFD9FEE11A214AC2F989" pageId="3" pageNumber="144" type="nomenclature">
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<taxonomicName id="4C374D77FFC1FFD9FEE11A214A7FF988" authority="Printarakul &amp; Chantanaorr." authorityName="Printarakul &amp; Chantanaorr." authorityYear="2021" class="Bryopsida" family="Splachnobryaceae" genus="Koponobryum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="144" phylum="Bryophyta" rank="species" species="papillosum" status="sp. nov.">
<emphasis id="B943EAE6FFC1FFD9FEE11A214A1DF9A9" bold="true" box="[314,594,1586,1613]" italics="true" pageId="3" pageNumber="144">Koponobryum papillosum</emphasis>
Printarakul &amp; Chantanaorr.
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,
<taxonomicNameLabel id="A270579DFFC1FFD9FDE11A404AC2F989" box="[570,653,1619,1645]" pageId="3" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
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<heading id="D0C08198FFC1FFD9FE4A1A6149B5F968" box="[401,506,1650,1676]" centered="true" fontSize="11" level="3" pageId="3" pageNumber="144" reason="3">
(
<figureCitation id="130C2A71FFC1FFD9FE431A6149BFF968" box="[408,496,1650,1676]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="4.[132,143,1962,1980]" captionStartId-1="5.[132,143,1962,1980]" captionStartId-2="6.[132,143,1962,1980]" captionTargetBox-0="[132,1341,215,1915]" captionTargetBox-1="[136,1342,221,1915]" captionTargetBox-2="[135,1341,219,1915]" captionTargetId-0="figure-26@4.[689,1249,780,1349]" captionTargetId-1="figure-17@5.[132,1342,215,1916]" captionTargetId-2="figure-26@6.[689,1249,780,1349]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionText-0="FIG. 1. — Koponobryum papillosum Printarakul &amp; Chantanaorr.,sp. nov.: A, plants in natural habitat;B, sporophyte bearing plants;C, leaves;D, leaf apex;E, upper laminal cells and marginal cells; F, leaf bases; G, sporophyte; H, capsule; I, peristome and spores; J, calyptra. Photos by N. Printarakul. All from N. Printarakul &amp; K. Adulkittichai 19072020_1 (CMUB). Scales bars: A, G, H, 1 mm; B, 3 mm; C, J, 0.5 mm; D-F, I, 50 µm." captionText-1="FIG. 2. – Koponobryum papillosum Printarakul &amp; Chantanaorr., sp. nov.: A, KOH color reaction of plant; B, cross section of stem; C, vegetative leaf; D-G, cross sections of leaf from upper to base; H, perichaetium and perigonium; I-L, perichaetial leaves; M-O, perigonial leaves; P, axillary hair. Photos by N. Printarakul. All from N. Printarakul &amp; K. Adulkittichai 19072020_1 (CMUB). Scale bars: A, 1 mm; B, D-G, P, 25 µm; C, H-O, 0.5 mm." captionText-2="FIG. 3. – Koponobryum papillosum Printarakul &amp; Chantanaorr.,sp.nov.:A, plants with sporophytes;B, cross section of stem;C, variations of axillary hairs;D, leaves; E, cross section of midleaf; F, leaf apex; G, leaf base; H, peristome (SEM); I, spore (SEM). A-G. Drawn by N. Printarakul. All from N. Printarakul &amp; K. Adulkittichai 19072020_1 (CMUB). Scale bars: A, 1 mm; B, C, F, G, 50 µm; D, 0.5 mm; E, 25 µm; H, 10 µm; I, 1 µm." figureDoi-0="http://doi.org/10.5281/zenodo.12215192" figureDoi-1="http://doi.org/10.5281/zenodo.12215194" figureDoi-2="http://doi.org/10.5281/zenodo.12215198" httpUri-0="https://zenodo.org/record/12215192/files/figure.png" httpUri-1="https://zenodo.org/record/12215194/files/figure.png" httpUri-2="https://zenodo.org/record/12215198/files/figure.png" pageId="3" pageNumber="144">Figs 1-3</figureCitation>
)
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</paragraph>
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<subSubSection id="C32D657FFFC1FFD9FF5F1AA44A2AF8E7" pageId="3" pageNumber="144" type="diagnosis">
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. — Similar to
<taxonomicName id="4C374D77FFC1FFD9FEA81AA44AC6F92B" authority="(Gangulee) Arts" authorityName="Arts" baseAuthorityName="Gangulee" box="[371,649,1718,1743]" class="Bryopsida" family="Splachnobryaceae" genus="Koponobryum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="144" phylum="Bryophyta" rank="species" species="bengalense">
<emphasis id="B943EAE6FFC1FFD9FEA81AA449A4F92A" box="[371,491,1718,1742]" italics="true" pageId="3" pageNumber="144">K. bengalense</emphasis>
(Gangulee) Arts
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, but differing in autoicous plants, papillose spores, and axillary hairs consisting of one short basal cell and 1-4(5) longer upper cells.
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. —
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<collectingCountry id="F3207664FFC1FFD9FF381B3E4973F8A1" box="[227,316,1837,1861]" name="Thailand" pageId="3" pageNumber="144">Thailand</collectingCountry>
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.
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: Mae Rim District, Doi Suthep-Pui National Park, Mae Sa Falls,
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,
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,
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.
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elev.,
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,
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. Printarakul &amp;
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. Adulkittichai 19072020_1
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(
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,
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!;
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,
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!,
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!).
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. —
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: Mae Rim District, Doi Suthep-Pui National Park, Mae
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Sa Falls,
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elev., on soil under granitic rock,
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,
<date id="FF891034FFC1FFD9FC101CE24C65FEED" box="[971,1066,241,265]" pageId="3" pageNumber="144" value="2013-10-04">4.X.2013</date>
,
<emphasis id="B943EAE6FFC1FFD9FBEF1CE14CB5FEED" box="[1076,1274,241,265]" italics="true" pageId="3" pageNumber="144">
<collectionCode id="ED26AE31FFC1FFD9FBEF1CE14C05FEED" box="[1076,1098,242,265]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="144" type="Herbarium">N</collectionCode>
. Printarakul 6444
</emphasis>
(
<collectionCode id="ED26AE31FFC1FFD9FADC1CE24D19FEED" box="[1287,1366,241,265]" pageId="3" pageNumber="144">CMUB</collectionCode>
); Chiang Dao District, Doi Chang Dao Wildlife Sanctuary, Huay Sob On forest protection unit,
<emphasis id="B943EAE6FFC1FFD9FBDA1D344C45FEDA" box="[1025,1034,295,318]" italics="true" pageId="3" pageNumber="144">c</emphasis>
.
<quantity id="4CCF9B11FFC1FFD9FBCC1D354C1AFEDA" box="[1047,1109,294,318]" metricMagnitude="2" metricUnit="m" metricValue="7.1" pageId="3" pageNumber="144" unit="m" value="710.0">710 m</quantity>
elev., on soil under calcareous rocks,
<geoCoordinate id="EE035033FFC1FFD9FCF01D524BF7FEBD" box="[811,952,321,345]" degrees="19" direction="north" minutes="21" orientation="latitude" pageId="3" pageNumber="144" precision="1" seconds="33.7" value="19.359362">19°21’33.7”N</geoCoordinate>
,
<geoCoordinate id="EE035033FFC1FFD9FC181D524C07FEBD" box="[963,1096,321,345]" degrees="98" direction="east" minutes="43" orientation="longitude" pageId="3" pageNumber="144" precision="1" seconds="07.5" value="98.71875">98°43’07.5”E</geoCoordinate>
,
<date id="FF891034FFC1FFD9FB8C1D524C92FEBD" box="[1111,1245,321,345]" pageId="3" pageNumber="144" value="2017-08-19">19.VIII.2017</date>
,
<emphasis id="B943EAE6FFC1FFD9FB321D524DFFFEBD" box="[1257,1456,321,345]" italics="true" pageId="3" pageNumber="144">
<collectionCode id="ED26AE31FFC1FFD9FB321D524CB0FEBC" box="[1257,1279,321,344]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="144" type="Herbarium">N</collectionCode>
. Printarakul 7280
</emphasis>
(
<collectionCode id="ED26AE31FFC1FFD9FCEF1D484BCDFE97" box="[820,898,347,371]" pageId="3" pageNumber="144">CMUB</collectionCode>
,
<collectionCode id="ED26AE31FFC1FFD9FC511D484BF4FE97" box="[906,955,347,371]" country="USA" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="3" pageNumber="144" type="Museum">PSU</collectionCode>
).
<collectingRegion id="49F3F816FFC1FFD9FC111D484CCAFE97" box="[970,1157,347,371]" country="Thailand" name="Lamphun" pageId="3" pageNumber="144">Lamphun Province</collectingRegion>
:
<collectingRegion id="49F3F816FFC1FFD9FB561D484CE7FE97" box="[1165,1192,347,371]" country="United Kingdom" name="Surrey" pageId="3" pageNumber="144">Sri</collectingRegion>
Bua Ban District, the Hariphuchai Education Centre of
<collectingRegion id="49F3F816FFC1FFD9FB9D1D654CF6FE6A" box="[1094,1209,374,398]" country="Thailand" name="Chiang Mai" pageId="3" pageNumber="144">Chiang Mai</collectingRegion>
University,
<emphasis id="B943EAE6FFC1FFD9FAF31D654D7EFE69" box="[1320,1329,374,397]" italics="true" pageId="3" pageNumber="144">c</emphasis>
.
<quantity id="4CCF9B11FFC1FFD9FAE61D664D34FE69" box="[1341,1403,373,397]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="144" unit="m" value="500.0">500 m</quantity>
elev., on soil under calcareous-sandstones,
<geoCoordinate id="EE035033FFC1FFD9FBA61D834D5EFE4C" box="[1149,1297,400,424]" degrees="18" direction="north" minutes="32" orientation="latitude" pageId="3" pageNumber="144" precision="1" seconds="17.33" value="18.538147">18°32’17.33”N</geoCoordinate>
,
<geoCoordinate id="EE035033FFC1FFD9FAC11D834DE8FE4C" box="[1306,1447,400,424]" degrees="99" direction="east" minutes="07" orientation="longitude" pageId="3" pageNumber="144" precision="1" seconds="28.10" value="99.12447">99°07’28.10”E</geoCoordinate>
,
<date id="FF891034FFC1FFD9FCF61DB84BF7FE27" box="[813,952,427,451]" pageId="3" pageNumber="144" value="2020-08-22">22.VIII.2020</date>
,
<emphasis id="B943EAE6FFC1FFD9FC1F1DB84BF5FE39" italics="true" pageId="3" pageNumber="144">
<collectionCode id="ED26AE31FFC1FFD9FC1F1DB84B95FE26" box="[964,986,427,450]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="3" pageNumber="144" type="Herbarium">N</collectionCode>
. Printarakul,
<collectionCode id="ED26AE31FFC1FFD9FBBC1DB84C35FE26" box="[1127,1146,427,450]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="3" pageNumber="144" type="Herbarium">A</collectionCode>
. Jampeetong &amp;
<collectionCode id="ED26AE31FFC1FFD9FAC31DB84D61FE26" box="[1304,1326,427,450]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15513" name="Nationaal Herbarium Nederland" pageId="3" pageNumber="144" type="Herbarium">U</collectionCode>
. ongkawong 22082020_15
</emphasis>
(
<collectionCode id="ED26AE31FFC1FFD9FC1D1DD64C59FE39" box="[966,1046,453,477]" pageId="3" pageNumber="144">CMUB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C32D657FFFC1FFD9FCF61DE44BF4FD38" pageId="3" pageNumber="144" type="etymology">
<paragraph id="8B8836F4FFC1FFD9FCF61DE44B82FDCC" blockId="3.[813,1455,502,553]" pageId="3" pageNumber="144">
<smallCapsWord id="8D6EA028FFC1FFD9FCF61DE44BE5FDE9" baselines="520,521" box="[813,938,503,526]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="3" pageNumber="144">ETYMOLOGY</smallCapsWord>
. — The specific epithet
<taxonomicName id="4C374D77FFC1FFD9FB771DE54D50FDEA" box="[1196,1311,502,526]" class="Bryopsida" family="Splachnobryaceae" genus="Koponobryum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="144" phylum="Bryophyta" rank="species" species="papillosum">
“
<emphasis id="B943EAE6FFC1FFD9FB6D1DE54D50FDEA" box="[1206,1311,502,526]" italics="true" pageId="3" pageNumber="144">papillosum</emphasis>
</taxonomicName>
” refers to the papillose spores.
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF61E524BF4FD38" blockId="3.[811,1456,576,733]" pageId="3" pageNumber="144">
<smallCapsWord id="8D6EA028FFC1FFD9FCF61E524BCCFDB3" baselines="594,595" box="[813,899,577,600]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Habitat" pageId="3" pageNumber="144">HABITAT</smallCapsWord>
<smallCapsWord id="8D6EA028FFC1FFD9FC531E554BFEFDB3" baselines="595" box="[904,945,582,599]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="and" pageId="3" pageNumber="144">AND</smallCapsWord>
<smallCapsWord id="8D6EA028FFC1FFD9FC6D1E554C71FDB3" baselines="595" box="[950,1086,582,599]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="distribution" pageId="3" pageNumber="144">DISTRIBUTION</smallCapsWord>
. —
<taxonomicName id="4C374D77FFC1FFD9FBBD1E534BA2FD97" authority="Printarakul &amp; Chantanaorr." authorityName="Printarakul &amp; Chantanaorr." authorityYear="2021" class="Bryopsida" family="Splachnobryaceae" genus="Koponobryum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pottiales" pageId="3" pageNumber="144" phylum="Bryophyta" rank="species" species="papillosum" status="sp. nov.">
<emphasis id="B943EAE6FFC1FFD9FBBD1E534D1CFDBC" box="[1126,1363,576,600]" italics="true" pageId="3" pageNumber="144">Koponobryum papillosum</emphasis>
Printarakul &amp; Chantanaorr.
</taxonomicName>
,
<taxonomicNameLabel id="A270579DFFC1FFD9FC2F1E484C72FD96" box="[1012,1085,603,626]" pageId="3" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
is currently known only from northern
<collectingCountry id="F3207664FFC1FFD9FCF01E664BCAFD69" box="[811,901,629,653]" name="Thailand" pageId="3" pageNumber="144">Thailand</collectingCountry>
. It grows on disturbed soils on a degraded calcareous substrate, sandstones and granitic rocks in deciduous, dipterocarp-oak, and also in mixed evergreen-deciduous seasonal forests at elevation of
<quantity id="4CCF9B11FFC1FFD9FC9D1ED64BF8FD38" box="[838,951,709,733]" metricMagnitude="2" metricUnit="m" metricValue="5.65" metricValueMax="7.1" metricValueMin="4.2" pageId="3" pageNumber="144" unit="m" value="565.0" valueMax="710.0" valueMin="420.0">
420-
<specimenCount id="9D31FD7DFFC1FFD9FCA81ED64BF8FD38" box="[883,951,709,733]" count="710" pageId="3" pageNumber="144" type="generic">710 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D657FFFC1FFDDFCF61EEE48A3FEF6" lastPageId="7" lastPageNumber="148" pageId="3" pageNumber="144" type="description">
<paragraph id="8B8836F4FFC1FFD9FCF61EEE4C52FC81" blockId="3.[813,1456,765,869]" pageId="3" pageNumber="144">
<smallCapsWord id="8D6EA028FFC1FFD9FCF61EEE4B89FCF0" baselines="783,784" box="[813,966,765,789]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Conservation" pageId="3" pageNumber="144">CONSERVATION</smallCapsWord>
<smallCapsWord id="8D6EA028FFC1FFD9FC161F104C5CFCF0" baselines="784" box="[973,1043,771,788]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="status" pageId="3" pageNumber="144">STATUS</smallCapsWord>
. —
<bibRefCitation id="EFA64B05FFC1FFD9FB9B1EEE4C85FCF1" author="IUCN" box="[1088,1226,765,789]" pageId="3" pageNumber="144" refId="ref2593" refString="IUCN 2012. - IUCN Red List categories and criteria: version 3.1, second edition. IUCN, Gland Switzerland and Cambridge, United Kingdom." type="book" year="2012">IUCN (2012)</bibRefCitation>
category Least concern (LC). Although only three locations were found in
<collectingCountry id="F3207664FFC1FFD9FAF91F0B4D34FCD4" box="[1314,1403,792,816]" name="Thailand" pageId="3" pageNumber="144">Thailand</collectingCountry>
, they are within the well-protected areas. Therefore, this species is not under immediate threat.
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF61F864BF2FC49" blockId="3.[813,957,917,943]" box="[813,957,917,943]" pageId="3" pageNumber="144">
<heading id="D0C08198FFC1FFD9FCF61F864BF2FC49" box="[813,957,917,943]" fontSize="9" level="3" pageId="3" pageNumber="144" reason="5">
<smallCapsWord id="8D6EA028FFC1FFD9FCF61F864BF2FC49" baselines="936,936" box="[813,957,917,943]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="3" pageNumber="144">DESCRIPTION</smallCapsWord>
</heading>
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF61FA64B22FC2B" blockId="3.[813,1456,949,1039]" box="[813,877,949,975]" pageId="3" pageNumber="144">
<heading id="D0C08198FFC1FFD9FCF61FA64B22FC2B" box="[813,877,949,975]" fontSize="11" level="3" pageId="3" pageNumber="144" reason="9">
<emphasis id="B943EAE6FFC1FFD9FCF61FA64B22FC2B" box="[813,877,949,975]" italics="true" pageId="3" pageNumber="144">Plants</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF61FC64C03FBEA" blockId="3.[813,1456,949,1039]" pageId="3" pageNumber="144">
Up to
<quantity id="4CCF9B11FFC1FFD9FCA31FC64B84FC0B" box="[888,971,981,1007]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="3" pageNumber="144" unit="mm" value="17.0">17 mm</quantity>
high (including sporophytes), pale to light green, forming loose turfs.
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF618274BCAFBAA" blockId="3.[813,901,1076,1102]" box="[813,901,1076,1102]" pageId="3" pageNumber="144">
<heading id="D0C08198FFC1FFD9FCF618274BCAFBAA" box="[813,901,1076,1102]" fontSize="11" level="3" pageId="3" pageNumber="144" reason="9">
<emphasis id="B943EAE6FFC1FFD9FCF618274BCAFBAA" box="[813,901,1076,1102]" italics="true" pageId="3" pageNumber="144">Rhizoids</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF618474D50FB8A" blockId="3.[813,1311,1108,1135]" box="[813,1311,1108,1135]" pageId="3" pageNumber="144">Reddish to pale brown, clustered at stem base.</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF618874B28FB4A" blockId="3.[813,871,1172,1198]" box="[813,871,1172,1198]" pageId="3" pageNumber="144">
<heading id="D0C08198FFC1FFD9FCF618874B28FB4A" box="[813,871,1172,1198]" fontSize="11" level="3" pageId="3" pageNumber="144" reason="9">
<emphasis id="B943EAE6FFC1FFD9FCF618874B28FB4A" box="[813,871,1172,1198]" italics="true" pageId="3" pageNumber="144">Stems</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF618A74D77FAA9" blockId="3.[813,1457,1204,1357]" pageId="3" pageNumber="144">Erect, unbranched, dark green to reddish brown, in cross section 8-10 layers of cells across, 120-150 µm in diameter; outer cortical cells somewhat smaller with moderately thick-walled, inner cortical cells larger with thicker walls; central stand consisting of smaller and thin-walled cells.</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF619604BF7FA69" blockId="3.[813,952,1395,1421]" box="[813,952,1395,1421]" pageId="3" pageNumber="144">
<heading id="D0C08198FFC1FFD9FCF619604BF7FA69" box="[813,952,1395,1421]" fontSize="11" level="3" pageId="3" pageNumber="144" reason="9">
<emphasis id="B943EAE6FFC1FFD9FCF619604BF7FA69" box="[813,952,1395,1421]" italics="true" pageId="3" pageNumber="144">Axillary hairs</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF619804BF2FA09" blockId="3.[811,1455,1427,1517]" pageId="3" pageNumber="144">Filiform, uniseriate, 2-5(-6) cells long, the basal cell shortest and yellowish to pale brown, the upper cells distinctly longer and colorless.</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF61A004B20F9C8" blockId="3.[813,879,1555,1580]" box="[813,879,1555,1580]" pageId="3" pageNumber="144">
<heading id="D0C08198FFC1FFD9FCF61A004B20F9C8" box="[813,879,1555,1580]" fontSize="11" level="3" pageId="3" pageNumber="144" reason="9">
<emphasis id="B943EAE6FFC1FFD9FCF61A004B20F9C8" box="[813,879,1555,1580]" italics="true" pageId="3" pageNumber="144">Leaves</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836F4FFC1FFD9FCF61A214DE9F80F" blockId="3.[810,1457,1586,2028]" pageId="3" pageNumber="144">
Contorted when dry, erecto-patent to spreading when moist; lanceolate to oblong-lanceolate, 1.1-1.5 ×
<quantity id="4CCF9B11FFC1FFD9FB2B1A414D3CF989" box="[1264,1395,1618,1645]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="3" pageNumber="144" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3-0.4 mm</quantity>
; apex acute to acuminate; base slightly decurrent; margins plane above, slightly revolute below, bordered throughout by 1-3 rows of elongate cells, remotely serrulate to denticulate above the midleaf; costa single, stout, short excurrent, ventral and dorsal surfaces smooth, not prominently bulging; costa cross section at midleaf semicircular to elliptic, with one stereid band; lamina unistratose, orange to red with KOH; upper laminal cells subquadrangular, polygonal to rounded-hexagonal, (20-) 25-30 µm in diameter, with a central papilla on both sides, firm to slightly thick-walled; basal marginal cells elongate rectangular, (35-) 40-80 × 12.5-15 µm wide, thin-walled; basal juxtacostal cells broader, rectangular, (37.5-)50-90 × 25-35 µm, thin-walled.
</paragraph>
<caption id="DF48667CFFC6FFDEFF5F1BB94BAEF80D" ID-DOI="http://doi.org/10.5281/zenodo.12215192" ID-Zenodo-Dep="12215192" httpUri="https://zenodo.org/record/12215192/files/figure.png" pageId="4" pageNumber="145" startId="4.[132,143,1962,1980]" targetBox="[132,1341,215,1915]" targetPageId="4" targetType="figure">
<paragraph id="8B8836F4FFC6FFDEFF5F1BB94BAEF80D" blockId="4.[132,1455,1962,2025]" pageId="4" pageNumber="145">
<smallCapsWord id="8D6EA028FFC6FFDEFF5F1BB948D2F85F" baselines="1975,1976" box="[132,157,1962,1980]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="4" pageNumber="145">FIG</smallCapsWord>
. 1. —
<taxonomicName id="4C374D77FFC6FFDEFF091BB94AC9F858" authority="Printarakul &amp; Chantanaorr." authorityName="Printarakul &amp; Chantanaorr." authorityYear="2021" box="[210,646,1962,1980]" class="Bryopsida" family="Splachnobryaceae" genus="Koponobryum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pottiales" pageId="4" pageNumber="145" phylum="Bryophyta" rank="species" species="papillosum" status="sp. nov.">
<emphasis id="B943EAE6FFC6FFDEFF091BB949E9F858" box="[210,422,1962,1980]" italics="true" pageId="4" pageNumber="145">Koponobryum papillosum</emphasis>
Printarakul &amp; Chantanaorr.
</taxonomicName>
,
<taxonomicNameLabel id="A270579DFFC6FFDEFD511BB94A81F858" box="[650,718,1962,1980]" pageId="4" pageNumber="145" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="B943EAE6FFC6FFDEFD0F1BB94AAEF858" bold="true" box="[724,737,1962,1980]" pageId="4" pageNumber="145">A</emphasis>
, plants in natural habitat;
<emphasis id="B943EAE6FFC6FFDEFC681BB94B8FF858" bold="true" box="[947,960,1962,1980]" pageId="4" pageNumber="145">B</emphasis>
, sporophyte bearing plants;
<emphasis id="B943EAE6FFC6FFDEFB711BB94CF7F858" bold="true" box="[1194,1208,1962,1980]" pageId="4" pageNumber="145">C</emphasis>
, leaves;
<emphasis id="B943EAE6FFC6FFDEFB241BB94D42F858" bold="true" box="[1279,1293,1962,1980]" pageId="4" pageNumber="145">D</emphasis>
, leaf apex;
<emphasis id="B943EAE6FFC6FFDEFAB21BB94D3AF858" bold="true" box="[1385,1397,1962,1980]" pageId="4" pageNumber="145">E</emphasis>
, upper laminal cells and marginal cells;
<emphasis id="B943EAE6FFC6FFDEFE541BD249D1F837" bold="true" box="[399,414,1985,2003]" pageId="4" pageNumber="145">F,</emphasis>
leaf bases;
<emphasis id="B943EAE6FFC6FFDEFDDB1BD34A41F836" bold="true" box="[512,526,1984,2002]" pageId="4" pageNumber="145">G</emphasis>
, sporophyte;
<emphasis id="B943EAE6FFC6FFDEFD5A1BD24AC0F837" bold="true" box="[641,655,1985,2003]" pageId="4" pageNumber="145">H</emphasis>
, capsule;
<emphasis id="B943EAE6FFC6FFDEFD3E1BD24AA4F837" bold="true" box="[741,747,1985,2003]" pageId="4" pageNumber="145">I</emphasis>
, peristome and spores;
<emphasis id="B943EAE6FFC6FFDEFC6F1BD24BF1F837" bold="true" box="[948,958,1985,2003]" pageId="4" pageNumber="145">J</emphasis>
, calyptra. Photos by N. Printarakul. All from
<emphasis id="B943EAE6FFC6FFDEFAF41BD24923F80D" italics="true" pageId="4" pageNumber="145">N. Printarakul &amp; K. Adulkittichai 19072020_1</emphasis>
(CMUB). Scales bars: A, G, H, 1 mm; B, 3 mm; C, J, 0.5 mm; D-F, I, 50 µm.
</paragraph>
</caption>
<caption id="DF48667CFFC7FFDFFF5F1BB94C13F80D" ID-DOI="http://doi.org/10.5281/zenodo.12215194" ID-Zenodo-Dep="12215194" httpUri="https://zenodo.org/record/12215194/files/figure.png" pageId="5" pageNumber="146" startId="5.[132,143,1962,1980]" targetBox="[136,1342,221,1915]" targetPageId="5" targetType="figure">
<paragraph id="8B8836F4FFC7FFDFFF5F1BB94C13F80D" blockId="5.[132,1457,1962,2025]" pageId="5" pageNumber="146">
<smallCapsWord id="8D6EA028FFC7FFDFFF5F1BB948D2F85F" baselines="1975,1976" box="[132,157,1962,1980]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="5" pageNumber="146">FIG</smallCapsWord>
. 2. –
<taxonomicName id="4C374D77FFC7FFDFFF171BB94AC7F858" authority="Printarakul &amp; Chantanaorr." authorityName="Printarakul &amp; Chantanaorr." authorityYear="2021" box="[204,648,1962,1980]" class="Bryopsida" family="Splachnobryaceae" genus="Koponobryum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pottiales" pageId="5" pageNumber="146" phylum="Bryophyta" rank="species" species="papillosum" status="sp. nov.">
<emphasis id="B943EAE6FFC7FFDFFF171BB949EDF858" box="[204,418,1962,1980]" italics="true" pageId="5" pageNumber="146">Koponobryum papillosum</emphasis>
Printarakul &amp; Chantanaorr.
</taxonomicName>
,
<taxonomicNameLabel id="A270579DFFC7FFDFFD551BB94A9BF858" box="[654,724,1962,1980]" pageId="5" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
:
<emphasis id="B943EAE6FFC7FFDFFD011BB94AA8F858" bold="true" box="[730,743,1962,1980]" pageId="5" pageNumber="146">A</emphasis>
, KOH color reaction of plant;
<emphasis id="B943EAE6FFC7FFDFFC3A1BB94BA1F858" bold="true" box="[993,1006,1962,1980]" pageId="5" pageNumber="146">B</emphasis>
, cross section of stem;
<emphasis id="B943EAE6FFC7FFDFFB631BB94C89F858" bold="true" box="[1208,1222,1962,1980]" pageId="5" pageNumber="146">C</emphasis>
, vegetative leaf;
<emphasis id="B943EAE6FFC7FFDFFA8F1BB94D38F858" bold="true" box="[1364,1399,1962,1980]" pageId="5" pageNumber="146">D -G</emphasis>
, cross sections of leaf from upper to base;
<emphasis id="B943EAE6FFC7FFDFFE6E1BD2498CF837" bold="true" box="[437,451,1985,2003]" pageId="5" pageNumber="146">H</emphasis>
, perichaetium and perigonium;
<emphasis id="B943EAE6FFC7FFDFFD151BD24AAAF837" bold="true" box="[718,741,1985,2003]" pageId="5" pageNumber="146">I -L</emphasis>
, perichaetial leaves;
<emphasis id="B943EAE6FFC7FFDFFC4D1BD24BF2F836" bold="true" box="[918,957,1984,2003]" pageId="5" pageNumber="146">M -O</emphasis>
, perigonial leaves;
<emphasis id="B943EAE6FFC7FFDFFB841BD24C24F837" bold="true" box="[1119,1131,1985,2003]" pageId="5" pageNumber="146">P</emphasis>
, axillary hair. Photos by N. Printarakul. All from
<emphasis id="B943EAE6FFC7FFDFFF121BC44A78F80D" box="[201,567,2007,2025]" italics="true" pageId="5" pageNumber="146">N. Printarakul &amp; K. Adulkittichai 19072020_1</emphasis>
(CMUB). Scale bars: A, 1 mm; B, D-G, P, 25 µm; C, H-O, 0.5 mm.
</paragraph>
</caption>
<caption id="DF48667CFFC4FFDCFF5F1BB94B9CF80D" ID-DOI="http://doi.org/10.5281/zenodo.12215198" ID-Zenodo-Dep="12215198" httpUri="https://zenodo.org/record/12215198/files/figure.png" pageId="6" pageNumber="147" startId="6.[132,143,1962,1980]" targetBox="[135,1341,219,1915]" targetPageId="6" targetType="figure">
<paragraph id="8B8836F4FFC4FFDCFF5F1BB94B9CF80D" blockId="6.[132,1457,1962,2025]" pageId="6" pageNumber="147">
<smallCapsWord id="8D6EA028FFC4FFDCFF5F1BB948D2F85F" baselines="1975,1976" box="[132,157,1962,1980]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="6" pageNumber="147">FIG</smallCapsWord>
. 3. –
<taxonomicName id="4C374D77FFC4FFDCFF131BB94A35F858" authority="Printarakul &amp; Chantanaorr." authorityName="Printarakul &amp; Chantanaorr." authorityYear="2021" box="[200,634,1962,1980]" class="Bryopsida" family="Splachnobryaceae" genus="Koponobryum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pottiales" pageId="6" pageNumber="147" phylum="Bryophyta" rank="species" species="papillosum" status="sp. nov.">
<emphasis id="B943EAE6FFC4FFDCFF131BB949D5F858" box="[200,410,1962,1980]" italics="true" pageId="6" pageNumber="147">Koponobryum papillosum</emphasis>
Printarakul &amp; Chantanaorr.
</taxonomicName>
,
<taxonomicNameLabel id="A270579DFFC4FFDCFDA51BB94A8EF858" box="[638,705,1962,1980]" pageId="6" pageNumber="147" rank="species">sp.nov.</taxonomicNameLabel>
:
<emphasis id="B943EAE6FFC4FFDCFD1D1BB94A9CF858" bold="true" box="[710,723,1962,1980]" pageId="6" pageNumber="147">A</emphasis>
, plants with sporophytes;
<emphasis id="B943EAE6FFC4FFDCFC7C1BB94BFBF858" bold="true" box="[935,948,1962,1980]" pageId="6" pageNumber="147">B</emphasis>
, cross section of stem;
<emphasis id="B943EAE6FFC4FFDCFBA81BB94CCEF858" bold="true" box="[1139,1153,1962,1980]" pageId="6" pageNumber="147">C</emphasis>
, variations of axillary hairs;
<emphasis id="B943EAE6FFC4FFDCFA841BB94D22F858" bold="true" box="[1375,1389,1962,1980]" pageId="6" pageNumber="147">D</emphasis>
, leaves;
<emphasis id="B943EAE6FFC4FFDCFF5F1BD248DFF837" bold="true" box="[132,144,1985,2003]" pageId="6" pageNumber="147">E</emphasis>
, cross section of midleaf;
<emphasis id="B943EAE6FFC4FFDCFEB01BD24939F837" bold="true" box="[363,374,1985,2003]" pageId="6" pageNumber="147">F</emphasis>
, leaf apex;
<emphasis id="B943EAE6FFC4FFDCFE0D1BD349ABF836" bold="true" box="[470,484,1984,2002]" pageId="6" pageNumber="147">G</emphasis>
, leaf base;
<emphasis id="B943EAE6FFC4FFDCFD981BD24A1EF837" bold="true" box="[579,593,1985,2003]" pageId="6" pageNumber="147">H</emphasis>
, peristome (SEM);
<emphasis id="B943EAE6FFC4FFDCFD341BD24ABAF837" bold="true" box="[751,757,1985,2003]" pageId="6" pageNumber="147">I</emphasis>
, spore (SEM). A-G. Drawn by N. Printarakul. All from
<emphasis id="B943EAE6FFC4FFDCFB751BD248A4F80D" italics="true" pageId="6" pageNumber="147">N. Printarakul &amp; K. Adulkittichai 19072020_1</emphasis>
(CMUB). Scale bars: A, 1 mm; B, C, F, G, 50 µm; D, 0.5 mm; E, 25 µm; H, 10 µm; I, 1 µm.
</paragraph>
</caption>
<paragraph id="8B8836F4FFC5FFDDFF5F1CC44913FF15" blockId="7.[132,348,215,241]" box="[132,348,215,241]" pageId="7" pageNumber="148">
<heading id="D0C08198FFC5FFDDFF5F1CC44913FF15" box="[132,348,215,241]" fontSize="11" level="3" pageId="7" pageNumber="148" reason="9">
<emphasis id="B943EAE6FFC5FFDDFF5F1CC44913FF15" box="[132,348,215,241]" italics="true" pageId="7" pageNumber="148">Asexual reproduction</emphasis>
</heading>
</paragraph>
<paragraph id="8B8836F4FFC5FFDDFF5F1CEB48A3FEF6" blockId="7.[132,236,248,274]" box="[132,236,248,274]" pageId="7" pageNumber="148">Not seen.</paragraph>
</subSubSection>
</treatment>
</document>
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