treatments-xml/data/03/9E/87/039E87D5D71FFF9EC8F8FC7FFE14FED3.xml

<document id="D4E8173E1CEAAFBA174082E5A8D7BAA6" ID-CLB-Dataset="29700" ID-DOI="10.11646/zootaxa.4444.3.7" ID-GBIF-Dataset="6dc9e3d4-9de7-4ad8-8c69-105a3011740c" ID-ISSN="1175-5326" ID-Zenodo-Dep="1309723" ID-ZooBank="9C9D7C03-3A3B-4B8F-AF53-3CCD997970D9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1531314062695" checkinUser="plazi" docAuthor="Cao, Dandan, Wei, Jianrong, Liu, Shuwen, Zhao, Zhengping, Wang, Liping &amp; Ji, Baozhong" docDate="2018" docId="039E87D5D71FFF9EC8F8FC7FFE14FED3" docLanguage="en" docName="zootaxa.4444.3.7.pdf" docOrigin="Zootaxa 4444 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homalota nanjingensis Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="330" masterDocId="FFA7FFADD71EFF9AC86FFFFBFF89FFCF" masterDocTitle="A new species of Homalota Mannerheim, 1830 (Coleoptera: Staphylinidae: Aleocharinae) from Nanjing, China" masterLastPageNumber="332" masterPageNumber="327" pageNumber="328" updateTime="1698553354115" updateUser="ExternalLinkService">
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<mods:title id="4BB17BD5AEBAD7B2CD47B27DEFC96D1E">A new species of Homalota Mannerheim, 1830 (Coleoptera: Staphylinidae: Aleocharinae) from Nanjing, China</mods:title>
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<mods:namePart id="D32848A2FA89DF9B8C24AD7FE8C9E3F4">Wei, Jianrong</mods:namePart>
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<mods:namePart id="B1B7088805A66D3B3FA40AE30A82C6B5">Zhao, Zhengping</mods:namePart>
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<taxonomicName id="4C374D40D71FFF9BC8F8FC7FFDDCFC50" ID-CoL="6MB2K" authority="Cao, Ji &amp; Liu" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[151,597,900,927]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="328" phylum="Arthropoda" rank="species" species="nanjingensis" status="sp. nov.">
<emphasis id="B943EAD1D71FFF9BC8F8FC7FFE2BFC51" bold="true" box="[151,418,900,926]" italics="true" pageId="1" pageNumber="328">Homalota nanjingensis</emphasis>
Cao, Ji &amp; Liu
</taxonomicName>
, 
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<typeStatus id="548C8861D71FFF9BC8F8FC37FF5BFC2A" box="[151,210,972,997]" pageId="1" pageNumber="328">Type</typeStatus>
locality: 
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</emphasis>
, 
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rpovince, 
<collectingMunicipality id="6BECACB9D71FFF9BCA35FC37FCE3FC2A" box="[602,874,972,997]" pageId="1" pageNumber="328">Nanjing Zijin Mountain</collectingMunicipality>
.
</materialsCitation>
</paragraph>
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<materialsCitation id="3B5F3C9ED71FFF9BC8A8FC14FDABFBE3" ID-GBIF-Occurrence="1880439748" collectingDate="2013-08-15" collectorName="S. W. Liu" country="China" location="Nanjing Zijin Mountain National Forest" municipality="Type" pageId="1" pageNumber="328" specimenCount="1" specimenCount-male="1" stateProvince="Jiangsu province" typeStatus="holotype">
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materials: 
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: 
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<emphasis id="B943EAD1D71FFF9BCA3CFC14FD15FBC7" bold="true" box="[595,668,1007,1032]" pageId="1" pageNumber="328">China</emphasis>
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, 
<collectingRegion id="49F3F821D71FFF9BCAC4FC14FCE2FBC7" box="[683,875,1007,1032]" country="China" name="Jiangsu" pageId="1" pageNumber="328">Jiangsu province</collectingRegion>
, 
<location id="8EE86018D71FFF9BCB19FC14FABEFBC8" LSID="urn:lsid:plazi:treatment:039E87D5D71FFF9EC8F8FC7FFE14FED3:8EE86018D71FFF9BCB19FC14FABEFBC8" box="[886,1335,1007,1032]" country="China" municipality="Type" name="Nanjing Zijin Mountain National Forest" pageId="1" pageNumber="328" stateProvince="Jiangsu province">Nanjing Zijin Mountain National Forest</location>
Park, 
<collectingDate id="EFCDE9EBD71FFF9BCDEFFC14FEA5FBE3" pageId="1" pageNumber="328" value="2013-08-15">
<date id="FF891003D71FFF9BCDEFFC14FEA5FBE3" pageId="1" pageNumber="328" value="2013-08-15">15 August 2013</date>
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, coll. 
<collectorName id="26C25315D71FFF9BC91BFBEFFE6AFBE2" box="[372,483,1044,1069]" pageId="1" pageNumber="328">S. W. Liu</collectorName>
&amp; D
</materialsCitation>
. 
<materialsCitation id="3B5F3C9ED71FFF9BCA5FFBEEFEF7FB80" ID-GBIF-Occurrence="1880439749" collectingDate="2013-07-08" collectorName="D. Cao. &amp; Z. P. Zhao" country="China" location="Nanjing" pageId="1" pageNumber="328" specimenCount="3" specimenCount-female="3" stateProvince="Jiangsu province" typeStatus="paratype">
<collectorName id="26C25315D71FFF9BCA5FFBEEFD00FBE3" box="[560,649,1044,1068]" pageId="1" pageNumber="328">D. Cao.</collectorName>
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: 
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, 
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<emphasis id="B943EAD1D71FFF9BCB15FBEFFC4AFBE2" bold="true" box="[890,963,1044,1069]" pageId="1" pageNumber="328">China</emphasis>
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, 
<collectingRegion id="49F3F821D71FFF9BCBBCFBEFFB1CFBE2" box="[979,1173,1044,1069]" country="China" name="Jiangsu" pageId="1" pageNumber="328">Jiangsu province</collectingRegion>
, 
<location id="8EE86018D71FFF9BCCCCFBEFFA89FBE2" LSID="urn:lsid:plazi:treatment:039E87D5D71FFF9EC8F8FC7FFE14FED3:8EE86018D71FFF9BCCCCFBEFFA89FBE2" box="[1187,1280,1044,1069]" country="China" name="Nanjing" pageId="1" pageNumber="328" stateProvince="Jiangsu province">Nanjing</location>
, 
<collectingDate id="EFCDE9EBD71FFF9BCD62FBEFFA11FBE3" box="[1293,1432,1044,1069]" pageId="1" pageNumber="328" value="2013-07-08">
<date id="FF891003D71FFF9BCD62FBEFFA11FBE3" box="[1293,1432,1044,1069]" pageId="1" pageNumber="328" value="2013-07-08">8 July 2013</date>
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, coll. 
<collectorName id="26C25315D71FFF9BC8A0FBC3FECAFB9F" box="[207,323,1079,1104]" pageId="1" pageNumber="328">Z. P. Zhao</collectorName>
&amp; D
</materialsCitation>
. 
<materialsCitation id="3B5F3C9ED71FFF9BC9E5FBC3FF41FBBB" ID-GBIF-Occurrence="1880439754" collectingDate="2013-09-12" collectorName="D. Cao. &amp; L. P. Wang &amp; D. D. Cao" country="China" location="Nanjing" pageId="1" pageNumber="328" specimenCount="4" specimenCount-male="4" stateProvince="Jiangsu province" typeStatus="paratype">
<collectorName id="26C25315D71FFF9BC9E5FBC3FE68FB9F" box="[394,481,1080,1104]" pageId="1" pageNumber="328">D. Cao.</collectorName>
<specimenCount id="9D31FD4AD71FFF9BC987FBC3FD84FB9F" box="[488,525,1080,1104]" pageId="1" pageNumber="328" type="male">4♂</specimenCount>
, 
<collectingCountry id="F3207653D71FFF9BCA76FBCCFDD6FB9F" box="[537,607,1079,1104]" name="China" pageId="1" pageNumber="328">China</collectingCountry>
, 
<collectingRegion id="49F3F821D71FFF9BCA05FBCCFCA2FB9F" box="[618,811,1079,1104]" country="China" name="Jiangsu" pageId="1" pageNumber="328">Jiangsu province</collectingRegion>
, 
<location id="8EE86018D71FFF9BCB59FBCCFC1AFB9F" LSID="urn:lsid:plazi:treatment:039E87D5D71FFF9EC8F8FC7FFE14FED3:8EE86018D71FFF9BCB59FBCCFC1AFB9F" box="[822,915,1079,1104]" country="China" name="Nanjing" pageId="1" pageNumber="328" stateProvince="Jiangsu province">Nanjing</location>
, 
<collectingDate id="EFCDE9EBD71FFF9BCBF1FBC3FBF5FB9F" box="[926,1148,1079,1104]" pageId="1" pageNumber="328" value="2013-09-12">
<date id="FF891003D71FFF9BCBF1FBC3FBF5FB9F" box="[926,1148,1079,1104]" pageId="1" pageNumber="328" value="2013-09-12">12 September 2013</date>
</collectingDate>
, coll. 
<collectorName id="26C25315D71FFF9BCCD0FBC3FAB2FB9F" box="[1215,1339,1080,1104]" pageId="1" pageNumber="328">L. P. Wang</collectorName>
&amp; 
<collectorName id="26C25315D71FFF9BCD31FBC3FF41FBBB" pageId="1" pageNumber="328">D. D. Cao</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B5F3C9ED71FFF9BC8BDFBA7FB6BFBBA" ID-GBIF-Occurrence="1880439780" box="[210,1250,1116,1141]" country="China" location="Insect Museum of Hebei University" pageId="1" pageNumber="328" specimenCount="1" stateProvince="Hebei" typeStatus="holotype">
All 
<typeStatus id="548C8861D71FFF9BC892FBA6FEA4FBBA" box="[253,301,1117,1141]" pageId="1" pageNumber="328">type</typeStatus>
specimens are deposited in the 
<location id="8EE86018D71FFF9BCAFEFBA6FBADFBBA" LSID="urn:lsid:plazi:treatment:039E87D5D71FFF9EC8F8FC7FFE14FED3:8EE86018D71FFF9BCAFEFBA6FBADFBBA" box="[657,1060,1116,1141]" country="China" name="Insect Museum of Hebei University" pageId="1" pageNumber="328" stateProvince="Hebei">
Insect Museum of 
<collectingRegion id="49F3F821D71FFF9BCB0DFBA7FC2CFBBA" box="[866,933,1116,1141]" country="China" name="Hebei" pageId="1" pageNumber="328">Hebei</collectingRegion>
University
</location>
, Baoding, 
<collectingCountry id="F3207653D71FFF9BCCF4FBA7FB6BFBBA" box="[1179,1250,1116,1141]" name="China" pageId="1" pageNumber="328">China</collectingCountry>
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.
</paragraph>
</subSubSection>
<subSubSection id="C32D6548D71FFF99C8A8FB84FA03F89B" lastPageId="3" lastPageNumber="330" pageId="1" pageNumber="328" type="description">
<paragraph id="8B8836C3D71FFF9BC8A8FB84FBF8FB73" blockId="1.[151,1436,972,1320]" pageId="1" pageNumber="328">
<emphasis id="B943EAD1D71FFF9BC8A8FB84FED2FB57" bold="true" box="[199,347,1151,1176]" pageId="1" pageNumber="328">Description:</emphasis>
Body length about 1.5–2.0 mm. Body strongly flattened dorsoventrally, parallel-sided; slightly pubescent; yellowish brown, head, elytra, and abdominal tergite VI dark brown (
<figureCitation id="130C2A46D71FFF9BCC70FB5FFBE9FB73" box="[1055,1120,1188,1213]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1954,1976]" captionTargetBox="[350,1238,1376,1933]" captionTargetId="figure@1.[340,1247,1373,1933]" captionTargetPageId="1" captionText="FIGURES 1–3. Homalota nanjingensis Cao, Ji, Liu, sp. nov.: 1 adult in dorsal view (scale bar = 0.5 mm); 2 head and pronotum, in dorsal view (scale bar = 0.1 mm); 3 antenna (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309725/files/figure.png" pageId="1" pageNumber="328">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B8836C3D71FFF9BC8A8FB3CFD2AFAE7" blockId="1.[151,1436,972,1320]" pageId="1" pageNumber="328">
<emphasis id="B943EAD1D71FFF9BC8A8FB3CFE99FB2F" bold="true" box="[199,272,1223,1248]" pageId="1" pageNumber="328">Head:</emphasis>
Subquadrate (HL = 0.29± 
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; HW = 0.27± 
<quantity id="4CCF9B26D71FFF9BCB53FB33FC2AFB10" box="[828,931,1224,1248]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="328" unit="mm" value="0.01">0.01 mm</quantity>
; HW/HL = 1.11); eyes moderate in siZe, and slightly protruding outwards (
<figureCitation id="130C2A46D71FFF9BC998FB17FDCBFACB" box="[503,578,1260,1285]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1954,1976]" captionTargetBox="[350,1238,1376,1933]" captionTargetId="figure@1.[340,1247,1373,1933]" captionTargetPageId="1" captionText="FIGURES 1–3. Homalota nanjingensis Cao, Ji, Liu, sp. nov.: 1 adult in dorsal view (scale bar = 0.5 mm); 2 head and pronotum, in dorsal view (scale bar = 0.1 mm); 3 antenna (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309725/files/figure.png" pageId="1" pageNumber="328">Fig. 2</figureCitation>
), between ommatidia with sparse setae; antenna with 11 segments, 1–3 elongate, 4–10 transverse, moniliform (
<figureCitation id="130C2A46D71FFF9BCA3EFAF4FD1FFAE7" box="[593,662,1295,1320]" captionStart="FIGURES 1–3" captionStartId="1.[151,264,1954,1976]" captionTargetBox="[350,1238,1376,1933]" captionTargetId="figure@1.[340,1247,1373,1933]" captionTargetPageId="1" captionText="FIGURES 1–3. Homalota nanjingensis Cao, Ji, Liu, sp. nov.: 1 adult in dorsal view (scale bar = 0.5 mm); 2 head and pronotum, in dorsal view (scale bar = 0.1 mm); 3 antenna (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309725/files/figure.png" pageId="1" pageNumber="328">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF48664BD71FFF9BC8F8F859FC34F818" httpUri="https://zenodo.org/record/1309725/files/figure.png" pageId="1" pageNumber="328" startId="1.[151,264,1954,1976]" targetBox="[350,1238,1376,1933]" targetPageId="1">
<paragraph id="8B8836C3D71FFF9BC8F8F859FC34F818" blockId="1.[151,1436,1954,2007]" pageId="1" pageNumber="328">
<emphasis id="B943EAD1D71FFF9BC8F8F859FEB7F877" bold="true" box="[151,318,1954,1976]" pageId="1" pageNumber="328">FIGURES 1–3.</emphasis>
<taxonomicName id="4C374D40D71FFF9BC926F859FD4CF877" authority="Cao, Ji, Liu" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[329,709,1954,1976]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="328" phylum="Arthropoda" rank="species" species="nanjingensis" status="sp. nov.">
<emphasis id="B943EAD1D71FFF9BC926F859FDB1F877" box="[329,568,1954,1976]" italics="true" pageId="1" pageNumber="328">Homalota nanjingensis</emphasis>
Cao, Ji, Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A27057AAD71FFF9BCABCF859FCA1F877" box="[723,808,1954,1976]" pageId="1" pageNumber="328" rank="species">
<emphasis id="B943EAD1D71FFF9BCABCF859FCA8F877" bold="true" box="[723,801,1954,1976]" pageId="1" pageNumber="328">sp. nov</emphasis>
.
</taxonomicNameLabel>
: 1 adult in dorsal view (scale bar = 0.5 mm); 2 head and pronotum, in dorsal view (scale bar = 0.1 mm); 3 antenna (scale bar = 0.1 mm).
</paragraph>
</caption>
<paragraph id="8B8836C3D71CFF98C8A8FF6CFDEBFE63" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="329">
<emphasis id="B943EAD1D71CFF98C8A8FF6CFED6FF7F" bold="true" box="[199,351,151,176]" pageId="2" pageNumber="329">Mouthparts:</emphasis>
Labrum transverse, with 8 pairs of macrosetae symmetrically arranged on the two sides of longitudinal midline.α-sensillum with a setose process, c- and γ-minute and conical, ε with a short setose process, distinctly shorter than α (
<figureCitation id="130C2A46D71CFF98C9AEFF24FD82FF37" box="[449,523,223,248]" captionStart="FIGURES 4–9" captionStartId="2.[151,264,1932,1954]" captionTargetBox="[159,1424,856,1878]" captionTargetId="figure@2.[151,1436,856,1900]" captionTargetPageId="2" captionText="FIGURES 4–9. Homalota nanjingensis Cao, Ji, Liu, sp. nov. 4 labrum, ventral aspect (scale bar = 0.02 mm); 5 right mandibles, ventral view (scale bar = 0.03 mm); 6 labium, ventral aspect (scale bar = 0.02 mm); 7 maxilla, ventral aspect (scale bar = 0.05 mm); 8 maxilla and labium (scale bar = 0.05 mm); 9 mesoventrite, ventral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309727/files/figure.png" pageId="2" pageNumber="329">Fig. 4</figureCitation>
); the molar region of the mandibles dorsal surface with small denticles, right mandible with small median tooth, prostheca well developed (
<figureCitation id="130C2A46D71CFF98CB0CFEFFFC28FED2" box="[867,929,260,285]" captionStart="FIGURES 4–9" captionStartId="2.[151,264,1932,1954]" captionTargetBox="[159,1424,856,1878]" captionTargetId="figure@2.[151,1436,856,1900]" captionTargetPageId="2" captionText="FIGURES 4–9. Homalota nanjingensis Cao, Ji, Liu, sp. nov. 4 labrum, ventral aspect (scale bar = 0.02 mm); 5 right mandibles, ventral view (scale bar = 0.03 mm); 6 labium, ventral aspect (scale bar = 0.02 mm); 7 maxilla, ventral aspect (scale bar = 0.05 mm); 8 maxilla and labium (scale bar = 0.05 mm); 9 mesoventrite, ventral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309727/files/figure.png" pageId="2" pageNumber="329">Fig.5</figureCitation>
); labial palpi with sparse setae, labium with ligula moderate in length and bifid at half, a little shorter than labial palpomere 1, labial palpomere 1 longer than 2, two medial setae present on prementum, contiguous, side by side, median field of prementum narrow and with pseudopores, mentum slightly marginated in anterior margin (
<figureCitation id="130C2A46D71CFF98CB22FE94FC18FE47" box="[845,913,367,392]" captionStart="FIGURES 4–9" captionStartId="2.[151,264,1932,1954]" captionTargetBox="[159,1424,856,1878]" captionTargetId="figure@2.[151,1436,856,1900]" captionTargetPageId="2" captionText="FIGURES 4–9. Homalota nanjingensis Cao, Ji, Liu, sp. nov. 4 labrum, ventral aspect (scale bar = 0.02 mm); 5 right mandibles, ventral view (scale bar = 0.03 mm); 6 labium, ventral aspect (scale bar = 0.02 mm); 7 maxilla, ventral aspect (scale bar = 0.05 mm); 8 maxilla and labium (scale bar = 0.05 mm); 9 mesoventrite, ventral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309727/files/figure.png" pageId="2" pageNumber="329">Fig. 6</figureCitation>
); maXillary palpomere 2 and 3 dilated distally, 4 without small spines at apeX (
<figureCitation id="130C2A46D71CFF98C997FE6FFDDCFE63" box="[504,597,404,429]" captionStart="FIGURES 4–9" captionStartId="2.[151,264,1932,1954]" captionTargetBox="[159,1424,856,1878]" captionTargetId="figure@2.[151,1436,856,1900]" captionTargetPageId="2" captionText="FIGURES 4–9. Homalota nanjingensis Cao, Ji, Liu, sp. nov. 4 labrum, ventral aspect (scale bar = 0.02 mm); 5 right mandibles, ventral view (scale bar = 0.03 mm); 6 labium, ventral aspect (scale bar = 0.02 mm); 7 maxilla, ventral aspect (scale bar = 0.05 mm); 8 maxilla and labium (scale bar = 0.05 mm); 9 mesoventrite, ventral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309727/files/figure.png" pageId="2" pageNumber="329">Fig. 7-8</figureCitation>
).
</paragraph>
<paragraph id="8B8836C3D71CFF98C8A8FE4CFE54FD4B" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="329">
<emphasis id="B943EAD1D71CFF98C8A8FE4CFEA0FE1F" bold="true" box="[199,297,439,464]" pageId="2" pageNumber="329">Thorax:</emphasis>
Pronotum transverse, about 1.29 times wider than long (PW = 0.35± 
<quantity id="4CCF9B26D71CFF98CC54FE4CFB2BFE00" box="[1083,1186,439,464]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="329" unit="mm" value="0.01">0.01 mm</quantity>
; PL = 0.27± 
<quantity id="4CCF9B26D71CFF98CD5FFE43FA11FE00" box="[1328,1432,440,464]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="329" unit="mm" value="0.01">0.01 mm</quantity>
; PW/PL = 1.29), posterior margin straight. Elytra slightly wider than pronotum, postero-laterally slightly sinuate. Elytra (
<figureCitation id="130C2A46D71CFF98C884FE04FEB5FDD7" box="[235,316,511,536]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="2" pageNumber="329">Fig. 10</figureCitation>
) longer than wide (EL = 0.34± 
<quantity id="4CCF9B26D71CFF98CAF7FDFBFD76FDD8" box="[664,767,512,536]" metricMagnitude="-5" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="329" unit="mm" value="0.02">0.02 mm</quantity>
; EW = 0.18± 
<quantity id="4CCF9B26D71CFF98CBFBFDFBFC72FDD8" box="[916,1019,512,536]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="329" unit="mm" value="0.03">0.03 mm</quantity>
; EL/EW = 1.91), slightly longer than pronotum (EL/PL = 1.35), hind wings developed, 1-1.5 times than body length. Scutellum postero-medially round. MesocoXae narrowly separated (
<figureCitation id="130C2A46D71CFF98CA6CFDBCFDCEFDAF" box="[515,583,583,608]" captionStart="FIGURES 4–9" captionStartId="2.[151,264,1932,1954]" captionTargetBox="[159,1424,856,1878]" captionTargetId="figure@2.[151,1436,856,1900]" captionTargetPageId="2" captionText="FIGURES 4–9. Homalota nanjingensis Cao, Ji, Liu, sp. nov. 4 labrum, ventral aspect (scale bar = 0.02 mm); 5 right mandibles, ventral view (scale bar = 0.03 mm); 6 labium, ventral aspect (scale bar = 0.02 mm); 7 maxilla, ventral aspect (scale bar = 0.05 mm); 8 maxilla and labium (scale bar = 0.05 mm); 9 mesoventrite, ventral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309727/files/figure.png" pageId="2" pageNumber="329">Fig. 9</figureCitation>
). Tarsomere 1 as long as 
<quantity id="4CCF9B26D71CFF98CB0BFDB3FC06FDAF" box="[868,911,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="329" unit="in" value="2.0">2 in</quantity>
front and middle leg, but longer than 
<quantity id="4CCF9B26D71CFF98CD56FDB3FAEAFDAF" box="[1337,1379,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="329" unit="in" value="2.0">2 in</quantity>
hind legs. Empodial setae present.
</paragraph>
<paragraph id="8B8836C3D71CFF98C8A8FD74FE3CFD03" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="329">
<emphasis id="B943EAD1D71CFF98C8A8FD74FECCFD67" bold="true" box="[199,325,655,680]" pageId="2" pageNumber="329">Abdomen:</emphasis>
Tergites III-?transversely impressed, with clear basal transverse lines; behind them uniformly covered with dense setae.
</paragraph>
<paragraph id="8B8836C3D71CFF98C8A8FD2CFAA3FCDB" blockId="2.[151,1437,151,789]" pageId="2" pageNumber="329">
<emphasis id="B943EAD1D71CFF98C8A8FD2CFEAEFD3F" bold="true" box="[199,295,727,752]" pageId="2" pageNumber="329">Female:</emphasis>
Posterior margin of female tergite VIII slightly sunk in middle part (
<figureCitation id="130C2A46D71CFF98CC3BFD2CFB20FD3F" box="[1108,1193,727,752]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="2" pageNumber="329">Fig. 11</figureCitation>
); sternite? (
<figureCitation id="130C2A46D71CFF98CD50FD2CFA1DFD3F" box="[1343,1428,727,752]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="2" pageNumber="329">Fig. 12</figureCitation>
) broadly rounded apically. Spermatheca simple and elongate, spermathecal tube slightly curved (
<figureCitation id="130C2A46D71CFF98CCA6FD07FA95FCDB" box="[1225,1308,764,789]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="2" pageNumber="329">Fig. 13</figureCitation>
).
</paragraph>
<caption id="DF48664BD71CFF98C8F8F877FA90F810" httpUri="https://zenodo.org/record/1309727/files/figure.png" pageId="2" pageNumber="329" startId="2.[151,264,1932,1954]" targetBox="[159,1424,856,1878]" targetPageId="2">
<paragraph id="8B8836C3D71CFF98C8F8F877FA90F810" blockId="2.[151,1436,1932,2015]" pageId="2" pageNumber="329">
<emphasis id="B943EAD1D71CFF98C8F8F877FEC9F86D" bold="true" box="[151,320,1932,1954]" pageId="2" pageNumber="329">FIGURES 4–9.</emphasis>
<taxonomicName id="4C374D40D71CFF98C924F877FD42F86D" authority="Cao, Ji, Liu" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[331,715,1932,1954]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="329" phylum="Arthropoda" rank="species" species="nanjingensis" status="sp. nov.">
<emphasis id="B943EAD1D71CFF98C924F877FDB5F86D" box="[331,572,1932,1954]" italics="true" pageId="2" pageNumber="329">Homalota nanjingensis</emphasis>
Cao, Ji, Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A27057AAD71CFF98CAB4F877FCB8F86D" box="[731,817,1932,1954]" pageId="2" pageNumber="329" rank="species">
<emphasis id="B943EAD1D71CFF98CAB4F877FCA2F86D" bold="true" box="[731,811,1932,1954]" pageId="2" pageNumber="329">sp. nov</emphasis>
.
</taxonomicNameLabel>
4 labrum, ventral aspect (scale bar = 0.02 mm); 5 right mandibles, ventral view (scale bar = 0.03 mm); 6 labium, ventral aspect (scale bar = 0.02 mm); 7 maxilla, ventral aspect (scale bar = 0.05 mm); 8 maxilla and labium (scale bar = 0.05 mm); 9 mesoventrite, ventral aspect (scale bar = 0.1 mm).
</paragraph>
</caption>
<paragraph id="8B8836C3D71DFF99C8A8FF6CFB8AFE8F" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="330">
<emphasis id="B943EAD1D71DFF99C8A8FF6CFE84FF7F" bold="true" box="[199,269,151,176]" pageId="3" pageNumber="330">Male:</emphasis>
Posterior margin of male tergite VIII with twao long and acute lateral processes, median area with 4 short and broad processes (
<figureCitation id="130C2A46D71DFF99C9B5FF47FDBAFF1B" box="[474,563,188,213]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="3" pageNumber="330">Fig. 14</figureCitation>
); sternite VIII narrowly protruding medially (
<figureCitation id="130C2A46D71DFF99CC3FFF47FB21FF1A" box="[1104,1192,188,213]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="3" pageNumber="330">Fig. 15</figureCitation>
); aedeagus (
<figureCitation id="130C2A46D71DFF99CD51FF47FA1DFF1A" box="[1342,1428,188,213]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="3" pageNumber="330">Fig. 16</figureCitation>
) median lobe elongate, bulbous at base, apical process slightly longer than basal bulb (
<figureCitation id="130C2A46D71DFF99CCE8FF24FB54FF37" box="[1159,1245,223,248]" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="3" pageNumber="330">Fig. 17</figureCitation>
), flagellum well sclerotiZed and short (
<figureCitation id="130C2A46D71DFF99C9E0FEFFFE6AFED3" box="[399,483,260,285]" captionStart="FIGURES 19–20" captionStartId="4.[151,264,1277,1299]" captionTargetBox="[149,1438,355,1244]" captionTargetId="figure@4.[149,1438,337,1256]" captionTargetPageId="4" captionText="FIGURES 19–20. Homalota nanjingensis Cao, Ji, Liu, sp. nov. 19 median lobe, ventral view (scale bar = 0.08 mm); 20 paramere, lateral aspect (scale bar = 0.08 mm)." httpUri="https://zenodo.org/record/1309731/files/figure.png" pageId="3" pageNumber="330">Fig. 19</figureCitation>
); the apical lobe of parameres long and nearly fusiform, the middle part dilated (
<figureCitation id="130C2A46D71DFF99CD00FEFFFF3CFE8F" captionStart="FIGURES 10–18" captionStartId="3.[151,264,1368,1390]" captionTargetBox="[180,1386,397,1329]" captionTargetId="figure@3.[151,1436,373,1347]" captionTargetPageId="3" captionText="FIGURES 10–18. Homalota nanjingensis Cao, Ji &amp; Liu, sp. nov. 10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm)." httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="3" pageNumber="330">Fig. 18</figureCitation>
), with two setae per paramere (
<figureCitation id="130C2A46D71DFF99CA7AFEDCFDE1FE8F" box="[533,616,295,320]" captionStart="FIGURES 19–20" captionStartId="4.[151,264,1277,1299]" captionTargetBox="[149,1438,355,1244]" captionTargetId="figure@4.[149,1438,337,1256]" captionTargetPageId="4" captionText="FIGURES 19–20. Homalota nanjingensis Cao, Ji, Liu, sp. nov. 19 median lobe, ventral view (scale bar = 0.08 mm); 20 paramere, lateral aspect (scale bar = 0.08 mm)." httpUri="https://zenodo.org/record/1309731/files/figure.png" pageId="3" pageNumber="330">Fig. 20</figureCitation>
), and approXimately equal in length.
</paragraph>
<caption id="DF48664BD71DFF99C8F8FAA3FBF0FA05" httpUri="https://zenodo.org/record/1309729/files/figure.png" pageId="3" pageNumber="330" startId="3.[151,264,1368,1390]" targetBox="[180,1386,397,1329]" targetPageId="3">
<paragraph id="8B8836C3D71DFF99C8F8FAA3FBF0FA05" blockId="3.[151,1436,1368,1482]" pageId="3" pageNumber="330">
<emphasis id="B943EAD1D71DFF99C8F8FAA3FEDDFAA1" bold="true" box="[151,340,1368,1390]" pageId="3" pageNumber="330">FIGURES 10–18.</emphasis>
<taxonomicName id="4C374D40D71DFF99C933FAA3FD57FAA1" authority="Cao, Ji &amp; Liu" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[348,734,1368,1390]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="nanjingensis" status="sp. nov.">
<emphasis id="B943EAD1D71DFF99C933FAA3FE48FAA1" box="[348,449,1368,1390]" italics="true" pageId="3" pageNumber="330">Homalota</emphasis>
<emphasis id="B943EAD1D71DFF99C9A7FAA2FDC1FAA1" box="[456,584,1369,1390]" italics="true" pageId="3" pageNumber="330">nanjingensis</emphasis>
Cao, Ji &amp; Liu
</taxonomicName>
, 
<taxonomicNameLabel id="A27057AAD71DFF99CA85FAA3FCB2FAA1" box="[746,827,1368,1390]" pageId="3" pageNumber="330" rank="species">
<emphasis id="B943EAD1D71DFF99CA85FAA3FCBCFAA1" bold="true" box="[746,821,1368,1390]" pageId="3" pageNumber="330">sp. nov</emphasis>
.
</taxonomicNameLabel>
10 elytrum in dorsal view (scale bar = 0.1 mm); 11 female tergite VIII (scale bar=0.1 mm); 12 female sternite VIII (scale bar=0.1 mm); 13 spermatheca (scale bar = 0.1 mm); 14 male tergite VIII (scale bar=0.1 mm); 15 male sternite VIII (scale bar=0.1 mm); 16 aedeagus, dorsal view (scale bar = 0.1 mm); 17 median lobe, lateral aspect (scale bar = 0.1 mm); 18 paramere, lateral aspect (scale bar = 0.1 mm).
</paragraph>
</caption>
<paragraph id="8B8836C3D71DFF99C8A8FA0CFAEFF96F" blockId="3.[151,1436,1527,2020]" pageId="3" pageNumber="330">
<emphasis id="B943EAD1D71DFF99C8A8FA0CFE75F9DF" bold="true" box="[199,508,1527,1552]" pageId="3" pageNumber="330">Habitat and distribution:</emphasis>
The specimens were collected in Nanjing Zijin Mountain National Forest Park (Nanjing, 
<collectingCountry id="F3207653D71DFF99C967F9E0FED9F9FB" box="[264,336,1563,1588]" name="China" pageId="3" pageNumber="330">China</collectingCountry>
). We found this new species under bark of dead pine trees (
<taxonomicName id="4C374D40D71DFF99CB84F9E7FB02F9FC" box="[1003,1163,1564,1587]" class="Gastropoda" family="Fasciolariidae" genus="Pinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="3" pageNumber="330" phylum="Mollusca" rank="species" species="massoniana">
<emphasis id="B943EAD1D71DFF99CB84F9E7FB02F9FC" box="[1003,1163,1564,1587]" italics="true" pageId="3" pageNumber="330">P. massoniana</emphasis>
</taxonomicName>
) that had been killed by nematode 
<taxonomicName id="4C374D40D71DFF99C964F9BBFE17F998" authorityName="Steiner &amp; Buhrer" authorityYear="1934" box="[267,414,1599,1623]" class="Secernentea" family="Aphelenchoididae" genus="Bursaphelenchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Aphelenchida" pageId="3" pageNumber="330" phylum="Nematoda" rank="species" species="xylophilus">
<emphasis id="B943EAD1D71DFF99C964F9BBFE17F998" box="[267,414,1599,1623]" italics="true" pageId="3" pageNumber="330">B. xylophilus</emphasis>
</taxonomicName>
, but we did not find this new species beneath the bark of completely rotten timber. Larvae of 
<taxonomicName id="4C374D40D71DFF99C8DAF99FFEEDF9B4" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[181,356,1636,1659]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="nanjingensis">
<emphasis id="B943EAD1D71DFF99C8DAF99FFEEDF9B4" box="[181,356,1636,1659]" italics="true" pageId="3" pageNumber="330">H. nanjingensis</emphasis>
</taxonomicName>
usually occurred between May and August. In contrast, based on three years of the investigation (
<date id="FF891003D71DFF99C8F0F973FE94F96F" box="[159,285,1672,1696]" pageId="3" pageNumber="330" value="2011" valueMax="2013">2011-2013</date>
), adults were found throughout the year, indicating that 
<taxonomicName id="4C374D40D71DFF99CBE0F973FBB6F950" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[911,1087,1672,1695]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="nanjingensis">
<emphasis id="B943EAD1D71DFF99CBE0F973FBB6F950" box="[911,1087,1672,1695]" italics="true" pageId="3" pageNumber="330">H. nanjingensis</emphasis>
</taxonomicName>
may overwinter as adults.
</paragraph>
<paragraph id="8B8836C3D71DFF99C8A8F950FA03F89B" blockId="3.[151,1436,1527,2020]" pageId="3" pageNumber="330">
With the secondary invasion of other insects (e.g. 
<taxonomicName id="4C374D40D71DFF99CB79F950FC05F90B" box="[790,908,1707,1732]" class="Insecta" family="Scolytidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="family">Scolytidae</taxonomicName>
and 
<taxonomicName id="4C374D40D71DFF99CBA4F950FBE6F90B" authorityName="Latreille" authorityYear="1802" box="[971,1135,1707,1732]" class="Insecta" family="Cerambycidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="family">Cerambycidae</taxonomicName>
) and microorganisms, the number of staphylinid species increased correspondingly. There was a spatial-temporal niche overlap between 
<taxonomicName id="4C374D40D71DFF99C8F8F908FEB8F8C3" box="[151,305,1779,1804]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
(
<taxonomicName id="4C374D40D71DFF99C92EF90FFE79F8C4" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[321,496,1780,1803]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="nanjingensis">
<emphasis id="B943EAD1D71DFF99C92EF90FFE79F8C4" box="[321,496,1780,1803]" italics="true" pageId="3" pageNumber="330">H. nanjingensis</emphasis>
</taxonomicName>
was the dominant species) and wood borer pests (
<bibRefCitation id="EFA64B32D71DFF99CC4CF90FFB6BF8C3" box="[1059,1250,1779,1804]" pageId="3" pageNumber="330" refId="ref2279" refString="Cao, D. D., Ji, B. Z., Liu, S. W., Chen, X. M., Zhang, X. W., Yang, J. J. &amp; Liu, J. J. (2013 b) Niches and following performance evaluation of arthropod populations under the barks of dead pine trees caused by Bursaphelenchus xylophilus. Chinese Journal of Ecology, 32, 3015 - 3021. [in Chinese]" type="journal article">
Cao 
<emphasis id="B943EAD1D71DFF99CC37F90FFB03F8C4" box="[1112,1162,1779,1803]" italics="true" pageId="3" pageNumber="330">et al</emphasis>
. 2013b
</bibRefCitation>
), but whether 
<taxonomicName id="4C374D40D71DFF99CDEEF90FFEACF8E0" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="nanjingensis">
<emphasis id="B943EAD1D71DFF99CDEEF90FFEACF8E0" italics="true" pageId="3" pageNumber="330">H. nanjingensis</emphasis>
</taxonomicName>
interacts with these wood borers remains unknown. In addition, feeding habits of 
<taxonomicName id="4C374D40D71DFF99CD68F8ECFA15F8FF" box="[1287,1436,1815,1840]" class="Insecta" family="Aleocharinae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="family">Aleocharinae</taxonomicName>
beetles are compleXed (
<bibRefCitation id="EFA64B32D71DFF99C9F1F8C7FDD2F89B" box="[414,603,1851,1876]" pageId="3" pageNumber="330" refId="ref2194" refString="Cao, D. D., Ji, B. Z., Liu, S. W., Chen, X. M., Wang, L. P., Zhao, Z. P. &amp; Ji, S. L. (2013 a) The feeding, defensive and reproductive behavior of adult staphlinids (Insecta: Coleoptera: Staphylinidae). Journal of Environmental Entomology, 35, 815 - 826. [in Chinese]" type="journal article">
Cao 
<emphasis id="B943EAD1D71DFF99C9BBF8C7FD8CF89C" box="[468,517,1851,1875]" italics="true" pageId="3" pageNumber="330">et al</emphasis>
. 2013a
</bibRefCitation>
), research on biology and behavior of 
<taxonomicName id="4C374D40D71DFF99CC66F8C7FB30F89C" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[1033,1209,1852,1875]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="nanjingensis">
<emphasis id="B943EAD1D71DFF99CC66F8C7FB30F89C" box="[1033,1209,1852,1875]" italics="true" pageId="3" pageNumber="330">H. nanjingensis</emphasis>
</taxonomicName>
will be continued.
</paragraph>
</subSubSection>
<subSubSection id="C32D6548D71DFF9EC8A8F8A4FE14FED3" lastPageId="4" lastPageNumber="331" pageId="3" pageNumber="330" type="etymology">
<paragraph id="8B8836C3D71DFF99C8A8F8A4FAC8F853" blockId="3.[151,1436,1527,2020]" pageId="3" pageNumber="330">
<emphasis id="B943EAD1D71DFF99C8A8F8A4FED8F8B7" bold="true" box="[199,337,1887,1912]" pageId="3" pageNumber="330">Etymology:</emphasis>
In our study, all samples of this species were collected from the Zijin Mountain National Forest Park, in the east of Nanjing City. The specific name is derived from the name of the 
<typeStatus id="548C8861D71DFF99CC27F87FFBF1F853" box="[1096,1144,1924,1948]" pageId="3" pageNumber="330">type</typeStatus>
locality, Nanjing.
</paragraph>
<paragraph id="8B8836C3D71DFF9EC8A8F85CFE14FED3" blockId="3.[151,1436,1527,2020]" lastBlockId="4.[151,1437,151,285]" lastPageId="4" lastPageNumber="331" pageId="3" pageNumber="330">
<emphasis id="B943EAD1D71DFF99C8A8F85CFE50F80F" bold="true" box="[199,473,1959,1984]" pageId="3" pageNumber="330">Comparative remarks:</emphasis>
<bibRefCitation id="EFA64B32D71DFF99C98EF85CFD44F80F" box="[481,717,1959,1984]" pageId="3" pageNumber="330" refId="ref2539" refString="Kim, Y. H. &amp; Ahn, K. J. (2014) Taxonomy of the genus Homalota Mannerheim in Korea (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys, 447, 109 - 123. https: // doi. org / 10.3897 / zookeys. 447.7728" type="journal article">Kim and Ahn (2014)</bibRefCitation>
revised the Korean species of 
<taxonomicName id="4C374D40D71DFF99CC46F85CFB10F870" authorityName="Mannerheim" authorityYear="1830" box="[1065,1177,1959,1983]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="genus">
<emphasis id="B943EAD1D71DFF99CC46F85CFB10F870" box="[1065,1177,1959,1983]" italics="true" pageId="3" pageNumber="330">Homalota</emphasis>
</taxonomicName>
, and provided detailed descriptions of five species and a key. 
<taxonomicName id="4C374D40D71DFF99CA2FF830FCCCF82C" authorityName="Cao &amp; Wei &amp; Liu &amp; Zhao &amp; Wang &amp; Ji" authorityYear="2018" box="[576,837,1995,2019]" class="Insecta" family="Staphylinidae" genus="Homalota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="330" phylum="Arthropoda" rank="species" species="nanjingensis">
<emphasis id="B943EAD1D71DFF99CA2FF830FCCCF82C" box="[576,837,1995,2019]" italics="true" pageId="3" pageNumber="330">Homalota nanjingensis</emphasis>
</taxonomicName>
can be distinguished from all species known in 
<collectingCountry id="F3207653D71DFF99CD36F830FA15F82B" box="[1369,1436,1995,2020]" name="China" pageId="3" pageNumber="330">China</collectingCountry>
by the following combination of characters: body strongly flattened dorsoventrally, parallel-sided, slightly pubescent; head slightly narrower than pronotum, posterior margin of pronotum relatively straight; male tergite? with two elongate lateral teeth, and with two setae per paramere. The key of species recorded in 
<collectingCountry id="F3207653D71AFF9ECD4DFF24FAECFF37" box="[1314,1381,223,248]" name="China" pageId="4" pageNumber="331">China</collectingCountry>
was enclosed (AppendiX 1).
</paragraph>
</subSubSection>
</treatment>
</document>