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<document id="3769495BFEDF32E76E7206493E686739" ID-DOI="10.5281/zenodo.4650555" ID-ISSN="1638-9395" ID-Zenodo-Dep="4650555" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617194132595" checkinUser="felipe" docAuthor="Abdala, Fernando &amp; Ribeiro, Ana Maria" docDate="2000" docId="039E87A43F7044085C04FB243D2CC75D" docLanguage="en" docName="Geodiversitas.22.4.589-596.pdf" docOrigin="Geodiversitas 22 (4)" docStyle="DocumentStyle:EA833BFD04EE0E3F39962C07E351CEA5.3:Geodiversitas.2000-2008.journal_article.type1" docStyleId="EA833BFD04EE0E3F39962C07E351CEA5" docStyleName="Geodiversitas.2000-2008.journal_article.type1" docStyleVersion="3" docTitle="Charruodon tetracuspidatus Abdala &amp; Ribeiro 2000, n. sp." docType="treatment" docVersion="3" lastPageNumber="592" masterDocId="FFA7FFDC3F71440B5F1FFFE13F47C101" masterDocTitle="A new therioherpetid cynodont from the Santa Maria Formation (middle Late Triassic), southern Brazil" masterLastPageNumber="596" masterPageNumber="589" pageNumber="590" updateTime="1698897146885" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
<mods:mods id="BDC6D2CE7D2FAB2AC382BEF1A3CABA2B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6D01968CD70DEB974BDCCF256D2A202C">
<mods:title id="B87B956124B1D413450C6C16053163EB">A new therioherpetid cynodont from the Santa Maria Formation (middle Late Triassic), southern Brazil</mods:title>
</mods:titleInfo>
<mods:name id="9A696A3D570C0DE8080CB313049ED490" type="personal">
<mods:role id="B875D3738A75FEE52F3F72B845389E9B">
<mods:roleTerm id="FD025279629DE8102A1D29F14CE0A5C3">Author</mods:roleTerm>
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<mods:namePart id="EC141D61F6DF11E9AC4CAA30AF969116">Abdala, Fernando</mods:namePart>
<mods:affiliation id="122F03A2DF40589DB10454896447EECA">Ciências e Tecnologia, Pontifícia Universidade Católica de Rio Grande do Sul, Avenida Ipiranga 6681, 90619 - 900 Porto Alegre, RS (Brazil) labpaleo @ pucrs. br</mods:affiliation>
<mods:nameIdentifier id="AAAFD5D5E3CCA2C03EF37B84CDC1D167" type="email">labpaleo@pucrs.br</mods:nameIdentifier>
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<mods:role id="2AA8B84F760260062390504290D9419F">
<mods:roleTerm id="47A033A2D828EDA53D6E179B13494870">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25374D4C7D7A5B2D4617A5391BB80D32">Ribeiro, Ana Maria</mods:namePart>
<mods:affiliation id="C39EFC6512F36A6CE542039688A4F409">Museu de Ciências Naturais, Fundação Zoobotânica do Rio Grande do Sul, Rua Dr Salvador França, 1427, 90690 - 000 Porto Alegre, RS (Brazil) amr @ plug-in. com. br.</mods:affiliation>
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<mods:title id="6F59E6F582C04FD9F582BD6E656B5669">Geodiversitas</mods:title>
</mods:titleInfo>
<mods:part id="7E203CA8945293EC8F23809F5996EE02">
<mods:date id="3BC3D6ECF2D2C9255AB79112D2F79A94">2000</mods:date>
<mods:detail id="1F87E637FCE24A8AF3CEF516A935C92F" type="volume">
<mods:number id="86C324973733704718C0DB4E38EEE60C">22</mods:number>
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<mods:start id="14759A67C247B2CBDD208852D99D700D">589</mods:start>
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<mods:classification id="1D29A0DE3A0E443E91130F468B3D5BBB">journal article</mods:classification>
<mods:identifier id="0487456D91963EF847B956D401EE6149" type="DOI">10.5281/zenodo.4650555</mods:identifier>
<mods:identifier id="308538819559F5EDA58C7121ABA4BE2F" type="ISSN">1638-9395</mods:identifier>
<mods:identifier id="7B19DB88EEBD3F8DC136A90C00A99C80" type="Zenodo-Dep">4650555</mods:identifier>
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<treatment id="039E87A43F7044085C04FB243D2CC75D" LSID="urn:lsid:plazi:treatment:039E87A43F7044085C04FB243D2CC75D" httpUri="http://treatment.plazi.org/id/039E87A43F7044085C04FB243D2CC75D" lastPageId="3" lastPageNumber="592" pageId="1" pageNumber="590">
<subSubSection id="C32D65393F70440A5C04FB243B09C5FC" pageId="1" pageNumber="590" type="nomenclature">
<paragraph id="8B8836B23F70440A5C04FB243B09C5FC" blockId="1.[795,1166,1220,1277]" pageId="1" pageNumber="590">
<heading id="D0C081DE3F70440A5C04FB243B09C5FC" centered="true" fontSize="11" level="2" pageId="1" pageNumber="590" reason="2">
<taxonomicName id="4C374D313F70440A5C04FB243B0CC5DE" authority="Abdala &amp; Ribeiro, 2000" authorityName="Abdala &amp; Ribeiro" authorityYear="2000" box="[795,1099,1221,1247]" class="Reptilia" family="Cynodontia" genus="Charruodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="1" pageNumber="590" phylum="Chordata" rank="species" species="tetracuspidatus" status="sp. nov.">
<emphasis id="B943EAA03F70440A5C04FB243B0CC5DE" bold="true" box="[795,1099,1221,1247]" italics="true" pageId="1" pageNumber="590">Charruodon tetracuspidatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27057DB3F70440A5B4CFB253BC9C5DF" box="[1107,1166,1220,1246]" pageId="1" pageNumber="590" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="130C2A373F70440A5C7EFB023CE1C5FC" box="[865,934,1251,1277]" captionStart="FIG" captionStartId="3.[151,162,1069,1086]" captionTargetBox="[224,1201,168,1030]" captionTargetId="figure-112@3.[224,1201,168,1030]" captionTargetPageId="3" captionText="FIG. 2. — Stereo pair and line drawing of the mandibular ramus of Charruodon tetracuspidatus n. sp. (MCP-3934 PV); A, labial view; B, lingual view. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4650557" httpUri="https://zenodo.org/record/4650557/files/figure.png" pageId="1" pageNumber="590">Figs 2</figureCitation>
;
<figureCitation id="130C2A373F70440A5CADFB023C86C5FC" box="[946,961,1251,1277]" captionStart="FIG" captionStartId="4.[151,162,771,788]" captionTargetBox="[163,1232,170,728]" captionTargetId="figure-376@4.[163,1232,170,728]" captionTargetPageId="4" captionText="FIG. 3. — Fifth tooth of Charruodon tetracuspidatus n. sp. (MCP-3934 PV); A, labial view; B, lingual view; C, crown view. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4650559" httpUri="https://zenodo.org/record/4650559/files/figure.png" pageId="1" pageNumber="590">3</figureCitation>
;
<tableCitation id="C6B503093F70440A5CD1FB023B6DC5FC" box="[974,1066,1251,1277]" captionStart="TABLE" captionStartId="3.[151,162,1163,1180]" captionText="TABLE 1. — Charruodon tetracuspidatus n. sp., holotype. Measurements of the specimen MCP-3934 PV (mm)." pageId="1" pageNumber="590">Tables 1</tableCitation>
;
<tableCitation id="C6B503093F70440A5B27FB023B00C5FC" box="[1080,1095,1251,1277]" captionStart="TABLE" captionStartId="3.[151,162,1472,1489]" captionText="TABLE 2. — Charruodon tetracuspidatus n. sp., holotype. Measurements of the fifth postcanine (mm)." pageId="1" pageNumber="590">2</tableCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D65393F70440A5DD5FAFD3BF0C480" pageId="1" pageNumber="590" type="materials_examined">
<paragraph id="8B8836B23F70440A5DD5FAFD3BF0C480" blockId="1.[714,1247,1308,1669]" pageId="1" pageNumber="590">
<typeStatus id="548C88103F70440A5DD5FAFD3C73C433" box="[714,820,1308,1331]" pageId="1" pageNumber="590" type="holotype">HOLOTYPE</typeStatus>
. — Fragment of the left dentary horizontal ramus, with alveoli for three incisors, roots of the canine and three postcanines, and a fifth tooth with a well-preserved crown (
<figureCitation id="130C2A373F70440A5CBAFA883CA4C480" box="[933,995,1385,1409]" captionStart="FIG" captionStartId="3.[151,162,1069,1086]" captionTargetBox="[224,1201,168,1030]" captionTargetId="figure-112@3.[224,1201,168,1030]" captionTargetPageId="3" captionText="FIG. 2. — Stereo pair and line drawing of the mandibular ramus of Charruodon tetracuspidatus n. sp. (MCP-3934 PV); A, labial view; B, lingual view. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4650557" httpUri="https://zenodo.org/record/4650557/files/figure.png" pageId="1" pageNumber="590">Figs 2</figureCitation>
;
<figureCitation id="130C2A373F70440A5CF1FA883CBCC480" box="[1006,1019,1385,1409]" captionStart="FIG" captionStartId="4.[151,162,771,788]" captionTargetBox="[163,1232,170,728]" captionTargetId="figure-376@4.[163,1232,170,728]" captionTargetPageId="4" captionText="FIG. 3. — Fifth tooth of Charruodon tetracuspidatus n. sp. (MCP-3934 PV); A, labial view; B, lingual view; C, crown view. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4650559" httpUri="https://zenodo.org/record/4650559/files/figure.png" pageId="1" pageNumber="590">3</figureCitation>
) (MCP-3934 PV).
</paragraph>
</subSubSection>
<subSubSection id="C32D65393F70440A5DD5FA6D3BE6C715" pageId="1" pageNumber="590" type="etymology">
<paragraph id="8B8836B23F70440A5DD5FA6D3B40C4BE" blockId="1.[714,1247,1308,1669]" pageId="1" pageNumber="590">
ETYMOLOGY. —
<taxonomicName id="4C374D313F70440A5C65FA6C3B49C4A4" authorityName="Abdala &amp; Ribeiro" authorityYear="2000" box="[890,1038,1421,1445]" class="Reptilia" family="Cynodontia" genus="Charruodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="1" pageNumber="590" phylum="Chordata" rank="species" species="tetracuspidatus">
<emphasis id="B943EAA03F70440A5C65FA6C3B49C4A4" box="[890,1038,1421,1445]" italics="true" pageId="1" pageNumber="590">tetracuspidatus</emphasis>
</taxonomicName>
, in reference to the four-cusped crown morphology.
</paragraph>
<paragraph id="8B8836B23F70440A5DD5FA293BE6C715" blockId="1.[714,1247,1308,1669]" pageId="1" pageNumber="590">
LOCALITY. —
<geoCoordinate id="EE0350753F70440A5C76FA283B52C4E0" box="[873,1045,1481,1505]" degrees="29" direction="south" minutes="42" orientation="latitude" pageId="1" pageNumber="590" precision="1" seconds="07.44" value="-29.702068">29°42’07.44’’S</geoCoordinate>
,
<geoCoordinate id="EE0350753F70440A5B3CFA283B9DC4E0" box="[1059,1242,1481,1505]" degrees="52" direction="west" minutes="51" orientation="longitude" pageId="1" pageNumber="590" precision="1" seconds="43.95" value="-52.862206">52°51’43.95’’W</geoCoordinate>
, approximately
<quantity id="4CCF9B573F70440A5C42FA023CD6C4FA" box="[861,913,1507,1531]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="590" unit="km" value="8.0">8 km</quantity>
West of the city of Candelaria in the State of
<collectingRegion id="49F3F8503F70440A5C5FFA1D3CBCC715" box="[832,1019,1532,1556]" country="Brazil" name="Rio Grande do Sul" pageId="1" pageNumber="590">Rio Grande do Sul</collectingRegion>
, southern
<collectingCountry id="F32076223F70440A5B7EFA1D3BDAC715" box="[1121,1181,1532,1556]" name="Brazil" pageId="1" pageNumber="590">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D65393F70440A5DD5F9C13C9AC784" pageId="1" pageNumber="590" type="description">
<paragraph id="8B8836B23F70440A5DD5F9C13C9AC784" blockId="1.[714,1247,1308,1669]" pageId="1" pageNumber="590">
HORIZON. — Santa Maria Formation. The presence of
<taxonomicName id="4C374D313F70440A5DF8F9DB3C26C753" authorityName="Cabrera" authorityYear="1943" box="[743,865,1594,1618]" class="Reptilia" family="Traversodontidae" genus="Exaeretodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="1" pageNumber="590" phylum="Chordata" rank="genus">
<emphasis id="B943EAA03F70440A5DF8F9DB3C26C753" box="[743,865,1594,1618]" italics="true" pageId="1" pageNumber="590">Exaeretodon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C374D313F70440A5C86F9DB3B72C753" authorityName="Reig" authorityYear="1959" box="[921,1077,1594,1618]" class="Reptilia" family="Proterochampsidae" genus="Proterochampsa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="1" pageNumber="590" phylum="Chordata" rank="genus">
<emphasis id="B943EAA03F70440A5C86F9DB3B72C753" box="[921,1077,1594,1618]" italics="true" pageId="1" pageNumber="590">Proterochampsa</emphasis>
</taxonomicName>
here is strongly suggestive of a Carnian age for this stratum of the formation (Rogers
<emphasis id="B943EAA03F70440A5C78F98D3CD1C784" box="[871,918,1644,1669]" italics="true" pageId="1" pageNumber="590">et al.</emphasis>
1993).
</paragraph>
</subSubSection>
<paragraph id="8B8836B23F7344095C57FEE93BDCC244" pageId="2" pageNumber="591">
<table id="F937C4123F73BBF45FA3FF3B3BE3C258" box="[188,1188,218,857]" gridcols="4" gridrows="8" pageId="2" pageNumber="591">
<tr id="350734F03F73BBF45FA3FF3B3BE3C01E" box="[188,1188,218,287]" gridrow="0" pageId="2" pageNumber="591" rowspan-0="1" rowspan-1="1" rowspan-3="1">
<th id="76D65D8C3F73BBF45DF5FF3B3CC1C01E" box="[746,902,218,287]" gridcol="2" gridrow="0" pageId="2" pageNumber="591">
<emphasis id="B943EAA03F7344095C57FEE93C2EC002" box="[840,873,218,287]" italics="true" pageId="2" pageNumber="591">Rio Pardo</emphasis>
</th>
</tr>
<tr id="350734F03F73BBF45FA3FEDF3BE3C0B1" box="[188,1188,318,432]" gridrow="1" pageId="2" pageNumber="591">
<th id="76D65D8C3F73BBF45FA3FEDF3E0EC0B1" box="[188,329,318,432]" gridcol="0" gridrow="1" pageId="2" pageNumber="591">SANTA MARIA</th>
<td id="76D65D8C3F73BBF45EA8FEDF3D1BC0B1" box="[439,604,318,432]" gridcol="1" gridrow="1" pageId="2" pageNumber="591">
<emphasis id="B943EAA03F7344095D1AFE893D61C065" box="[517,550,318,384]" italics="true" pageId="2" pageNumber="591">Rio Jacui</emphasis>
</td>
<td id="76D65D8C3F73BBF45DF5FEDF3CC1C0B1" box="[746,902,318,432]" gridcol="2" gridrow="1" pageId="2" pageNumber="591">CANDELÁRIA</td>
<td id="76D65D8C3F73BBF45CC4FEDF3BE3C0B1" box="[987,1188,318,432]" gridcol="3" gridrow="1" pageId="2" pageNumber="591">VENÂNCIO AIRES</td>
</tr>
<tr id="350734F03F73BBF45FA3FE283BE3C0DC" box="[188,1188,457,477]" gridrow="2" pageId="2" pageNumber="591">
<th id="76D65D8C3F73BBF45FA3FE283BE3C0DC" box="[188,1188,457,477]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="2" pageNumber="591">BR 287</th>
</tr>
<tr id="350734F03F73BBF45FA3FE1C3BE3C345" box="[188,1188,509,580]" gridrow="3" pageId="2" pageNumber="591" rowspan-2="1">
<th id="76D65D8C3F73BBF45FA3FE1C3E0EC345" box="[188,329,509,580]" gridcol="0" gridrow="3" pageId="2" pageNumber="591">ARGENTINA</th>
<td id="76D65D8C3F73BBF45EA8FE1C3D1BC345" box="[439,604,509,580]" gridcol="1" gridrow="3" pageId="2" pageNumber="591">N</td>
<td id="76D65D8C3F73BBF45CC4FE1C3BE3C345" box="[987,1188,509,580]" gridcol="3" gridrow="3" pageId="2" pageNumber="591">SANTA CRUZ DO SUL</td>
</tr>
<tr id="350734F03F73BBF45FA3FDBB3BE3C36F" box="[188,1188,602,622]" gridrow="4" pageId="2" pageNumber="591" rowspan-2="1" rowspan-3="1">
<th id="76D65D8C3F73BBF45FA3FDBB3D1BC36F" box="[188,604,602,622]" colspan="2" colspanRight="1" gridcol="0" gridrow="4" pageId="2" pageNumber="591">RIO GRANDE DO SUL</th>
</tr>
<tr id="350734F03F73BBF45FA3FD243BE3C3F8" box="[188,1188,709,761]" gridrow="5" pageId="2" pageNumber="591" rowspan-0="1">
<td id="76D65D8C3F73BBF45EA8FD243D1BC3F8" box="[439,604,709,761]" gridcol="1" gridrow="5" pageId="2" pageNumber="591">BRAZIL</td>
<td id="76D65D8C3F73BBF45DF5FD243CC1C3F8" box="[746,902,709,761]" gridcol="2" gridrow="5" pageId="2" pageNumber="591">CACHOEIRA DO SUL</td>
<td id="76D65D8C3F73BBF45CC4FD243BE3C3F8" box="[987,1188,709,761]" gridcol="3" gridrow="5" pageId="2" pageNumber="591">BR 471</td>
</tr>
<tr id="350734F03F73BBF45FA3FCE63BE3C224" box="[188,1188,775,805]" gridrow="6" pageId="2" pageNumber="591" rowspan-1="1" rowspan-2="1">
<th id="76D65D8C3F73BBF45FA3FCE63E0EC224" box="[188,329,775,805]" gridcol="0" gridrow="6" pageId="2" pageNumber="591">URUGUAY</th>
<td id="76D65D8C3F73BBF45CC4FCE63BE3C224" box="[987,1188,775,805]" gridcol="3" gridrow="6" pageId="2" pageNumber="591">BR 290</td>
</tr>
<tr id="350734F03F73BBF45FA3FCD03BE3C258" box="[188,1188,817,857]" gridrow="7" pageId="2" pageNumber="591" rowspan-0="1">
<td id="76D65D8C3F73BBF45EA8FCD03D1BC258" box="[439,604,817,857]" gridcol="1" gridrow="7" pageId="2" pageNumber="591">120 km</td>
<td id="76D65D8C3F73BBF45DF5FCD03CC1C258" box="[746,902,817,857]" gridcol="2" gridrow="7" pageId="2" pageNumber="591">25 km</td>
<td id="76D65D8C3F73BBF45CC4FCD03BE3C258" box="[987,1188,817,857]" gridcol="3" gridrow="7" pageId="2" pageNumber="591">PANTANO GRANDE</td>
</tr>
</table>
</paragraph>
<caption id="DF48663A3F7344095ED0FC7F3CEFC2AE" box="[463,936,926,943]" pageId="2" pageNumber="591">
<paragraph id="8B8836B23F7344095ED0FC7F3CEFC2AE" blockId="2.[463,936,926,943]" box="[463,936,926,943]" pageId="2" pageNumber="591">FIG. 1. — Map showing the location of the fossil outcrop.</paragraph>
</caption>
<subSubSection id="C32D65393F7344095F88FBE93DDDC5ED" pageId="2" pageNumber="591" type="diagnosis">
<paragraph id="8B8836B23F7344095F88FBE93DDDC5ED" blockId="2.[151,684,1032,1260]" pageId="2" pageNumber="591">DIAGNOSIS. — Lower jaw with a high horizontal ramus when compared with the postcanine size, approximately 0.31 with respect to the anteroposterior length of the crown of the fifth postcanine; four welldeveloped cusps on the crown of the postcanine; second cusp “a” is the largest, followed closely in size by the third (“c”); the three posterior cusps (“a”, “c” and “d”) are directed posteriorly; a sharp constriction distinguishes the crown from the root; roots undivided.</paragraph>
</subSubSection>
<subSubSection id="C32D65393F7344085F88FACB3D2CC75D" lastPageId="3" lastPageNumber="592" pageId="2" pageNumber="591" type="description">
<paragraph id="8B8836B23F7344095F88FACB3E6FC443" blockId="2.[151,685,1322,1669]" box="[151,296,1322,1348]" pageId="2" pageNumber="591">DESCRIPTION</paragraph>
<paragraph id="8B8836B23F7344095F88FAAA3BEFC784" blockId="2.[151,685,1322,1669]" lastBlockId="2.[714,1247,1034,1669]" pageId="2" pageNumber="591">
The dentary fragment extends from an open and high symphysis to the level of the third postcanine (
<figureCitation id="130C2A373F7344095F80FA6A3E51C4A5" box="[159,278,1418,1445]" captionStart="FIG" captionStartId="3.[151,162,1069,1086]" captionTargetBox="[224,1201,168,1030]" captionTargetId="figure-112@3.[224,1201,168,1030]" captionTargetPageId="3" captionText="FIG. 2. — Stereo pair and line drawing of the mandibular ramus of Charruodon tetracuspidatus n. sp. (MCP-3934 PV); A, labial view; B, lingual view. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4650557" httpUri="https://zenodo.org/record/4650557/files/figure.png" pageId="2" pageNumber="591">Fig. 2A, B</figureCitation>
). There are three alveoli for the incisors. The root of the canine is larger than those of the incisors and postcanines, but has undergone some transverse compression during fossilization. The mental foramen is located at the level of the second postcanine, and closer to the symphysis are some nutritive foramina (
<figureCitation id="130C2A373F7344095EB2F9AA3D44C764" box="[429,515,1611,1637]" captionStart="FIG" captionStartId="3.[151,162,1069,1086]" captionTargetBox="[224,1201,168,1030]" captionTargetId="figure-112@3.[224,1201,168,1030]" captionTargetPageId="3" captionText="FIG. 2. — Stereo pair and line drawing of the mandibular ramus of Charruodon tetracuspidatus n. sp. (MCP-3934 PV); A, labial view; B, lingual view. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4650557" httpUri="https://zenodo.org/record/4650557/files/figure.png" pageId="2" pageNumber="591">Fig. 2B</figureCitation>
). Three postcanines are represented only by their roots. All ap- pear relatively small compared with the anterior height of the jaw. Originally, the region preserved extended as far posteriorly as the fifth alveolus, which contained a replacing tooth that preserved a complete crown. This tooth was isolated and prepared in order to examine its crown morphology (
<figureCitation id="130C2A373F7344095DCDFB2A3C53C5E4" box="[722,788,1227,1253]" captionStart="FIG" captionStartId="4.[151,162,771,788]" captionTargetBox="[163,1232,170,728]" captionTargetId="figure-376@4.[163,1232,170,728]" captionTargetPageId="4" captionText="FIG. 3. — Fifth tooth of Charruodon tetracuspidatus n. sp. (MCP-3934 PV); A, labial view; B, lingual view; C, crown view. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4650559" httpUri="https://zenodo.org/record/4650559/files/figure.png" pageId="2" pageNumber="591">Fig. 3</figureCitation>
). Four aligned cusps are present and there is no indication of a cingulum. The main cusp “a” is slightly larger than the first posterior accessory cusp “c” from which it is separated by a notch. Both of these cusps are slightly recurved posteriorly. The second posterior accessory cusp “d” is the lowest. It is posteriorly directed and wellseparated from cusp “c” by a notch. The anterior accessory cusp “b” is relatively small and closely united with the main cusp by a high crest. In occlusal view, the crown is bulbous, with convex labial and lingual sides, the latter being particulary rounded. The root is undivided and distinguished from the crown by a prominent constriction.
</paragraph>
<caption id="DF48663A3F7244085F88FBCC3EF4C555" ID-DOI="http://doi.org/10.5281/zenodo.4650557" ID-Zenodo-Dep="4650557" httpUri="https://zenodo.org/record/4650557/files/figure.png" pageId="3" pageNumber="592" startId="3.[151,162,1069,1086]" targetBox="[224,1201,168,1030]" targetPageId="3">
<paragraph id="8B8836B23F7244085F88FBCC3EF4C555" blockId="3.[151,1248,1069,1108]" pageId="3" pageNumber="592">
FIG. 2. — Stereo pair and line drawing of the mandibular ramus of
<taxonomicName id="4C374D313F7244085DA1FBCC3CE0C53F" authority="Abdala &amp; Ribeiro, 2000" authorityName="Abdala &amp; Ribeiro" authorityYear="2000" box="[702,935,1069,1086]" class="Reptilia" family="Cynodontia" genus="Charruodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="3" pageNumber="592" phylum="Chordata" rank="species" species="tetracuspidatus" status="sp. nov.">
<emphasis id="B943EAA03F7244085DA1FBCC3CE0C53F" box="[702,935,1069,1086]" italics="true" pageId="3" pageNumber="592">Charruodon tetracuspidatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27057DB3F7244085CB3FBCC3C9BC53F" box="[940,988,1069,1086]" pageId="3" pageNumber="592" rank="species">n. sp.</taxonomicNameLabel>
(MCP-3934 PV);
<emphasis id="B943EAA03F7244085B73FBCC3B3EC53F" bold="true" box="[1132,1145,1069,1086]" pageId="3" pageNumber="592">A</emphasis>
, labial view;
<emphasis id="B943EAA03F7244085F88FBA23FE3C555" bold="true" box="[151,164,1091,1108]" pageId="3" pageNumber="592">B</emphasis>
, lingual view. Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="DF48663A3F7244085F88FB6A3D08C5B0" pageId="3" pageNumber="592">
<paragraph id="8B8836B23F7244085F88FB6A3D08C5B0" blockId="3.[151,685,1163,1201]" pageId="3" pageNumber="592">
TABLE 1. —
<taxonomicName id="4C374D313F7244085E1BFB6A3D4DC59D" authority="Abdala &amp; Ribeiro, 2000" authorityName="Abdala &amp; Ribeiro" authorityYear="2000" box="[260,522,1163,1180]" class="Reptilia" family="Cynodontia" genus="Charruodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Therapsida" pageId="3" pageNumber="592" phylum="Chordata" rank="species" species="tetracuspidatus" status="sp. nov.">
<emphasis id="B943EAA03F7244085E1BFB6A3D4DC59D" box="[260,522,1163,1180]" italics="true" pageId="3" pageNumber="592">Charruodon tetracuspidatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27057DB3F7244085D0DFB6A3D0CC59D" box="[530,587,1163,1180]" pageId="3" pageNumber="592" rank="species">n. sp.</taxonomicNameLabel>
, holotype. Measurements of the specimen MCP-3934 PV (mm).
</paragraph>
</caption>
<paragraph id="8B8836B23F7244085F88FB013D2CC75D" blockId="3.[151,684,1248,1396]" lastBlockId="3.[151,619,1557,1628]" pageId="3" pageNumber="592">Total length of specimen 23.5 Height at the level of the first postcanine 15.2 Anteroposterior length of the root of the canine 5.0 Anteroposterior length of the root of postcanine 1 2.8 Anteroposterior length of the root of postcanine 2 3.0 Anteroposterior length of the root of postcanine 3 3.0 Crown length 4.8 Crown height 3.0 Crown width 1.8</paragraph>
</subSubSection>
</treatment>
</document>
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