treatments-xml/data/03/9E/87/039E8789496B2F1FFF59F837FC9CDD76.xml

<document id="2FECC3B6B79F75C6F06F7FFE6B335E75" ID-CLB-Dataset="277847" ID-DOI="10.11646/zootaxa.5361.2.8" ID-GBIF-Dataset="5bf93237-8248-486c-855f-c0f87553a671" ID-ISSN="1175-5326" ID-Zenodo-Dep="10146585" ID-ZooBank="9FB81B0C-412B-4978-B2CF-4151DA505F93" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700189545860" checkinUser="plazi" docAuthor="Yuan, Zi-Ming, Jiang, Wei &amp; Sha, Zhong-Li" docDate="2023" docId="039E8789496B2F1FFF59F837FC9CDD76" docLanguage="en" docName="zootaxa.5361.2.8.pdf" docOrigin="Zootaxa 5361 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.2.8/52183" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cymo mazu Yuan &amp; Jiang &amp; Sha 2023, sp. nov." docType="treatment" docVersion="5" lastPageNumber="281" masterDocId="FFA7FFF1496A2F19FFCEFFBAFFA3DA15" masterDocTitle="A new species of the coral-symbiont crab genus Cymo de Haan, 1833 (Decapoda, Brachyura, Xanthidae) from Nansha Islands, the South China Sea" masterLastPageNumber="286" masterPageNumber="275" pageNumber="276" updateTime="1700241860690" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="8D10D5C127C39E309CA6437CE9F02D81" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="14B8CFB58E05A1E702EDB5751ECF53D6">
<mods:title id="9849F762E968A20CDE04B4DB9F9C65E4">A new species of the coral-symbiont crab genus Cymo de Haan, 1833 (Decapoda, Brachyura, Xanthidae) from Nansha Islands, the South China Sea</mods:title>
</mods:titleInfo>
<mods:name id="B203BEB855E9C5E5EEE6E97AF00EAA55" type="personal">
<mods:role id="0B424D45E40E4F07080C4897FFD81C7B">
<mods:roleTerm id="5D627A4B417F250A01231F4FC95BD770">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="52B402177127E2ABFC8CFA08DEBE0E83">Yuan, Zi-Ming</mods:namePart>
<mods:affiliation id="ED6E7194BD9E4BDDC2B97781AB5146ED">Laboratory of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China. yuanziming @ qdio. ac. cn; jiangwei @ qdio. ac. cn; shazl @ qdio. ac. cn &amp; University of Chinese Academy of Sciences, Beijing, China</mods:affiliation>
<mods:nameIdentifier id="18E50415D8F5D76099CAAB183B803E2A" type="email">yuanziming@qdio.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="8F9B58C5AD41E1D4AA0BAF63B1B0476C" type="personal">
<mods:role id="4ECC7651B962336CB2416F472AC9F20B">
<mods:roleTerm id="66930CA6F5E9BCAC88D378FBB95C79DF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="13E045D4262AD1E1A943E3CB4A93D9E9">Jiang, Wei</mods:namePart>
<mods:affiliation id="B7B6C08429C5B833D794EE82BC5D58C0">Laboratory of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China. yuanziming @ qdio. ac. cn; jiangwei @ qdio. ac. cn; shazl @ qdio. ac. cn &amp; Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China &amp; Laoshan Laboratory, Qingdao, China</mods:affiliation>
<mods:nameIdentifier id="6D8C7F885B440A86B57701B8288E0D98" type="email">yuanziming@qdio.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="B8BBB3684898578AFF6F2E846CDF2BEE" type="personal">
<mods:role id="FF4644041A895DAC0A4117454403DB18">
<mods:roleTerm id="4E7EE9A7D06C74D3035D6D85D83EFE4D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="344576200DAC09006913D520943D98ED">Sha, Zhong-Li</mods:namePart>
<mods:affiliation id="4DE7A7C39703845F29CC0A431B0E6A0E">Laboratory of Marine Organism Taxonomy and Phylogeny, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China. yuanziming @ qdio. ac. cn; jiangwei @ qdio. ac. cn; shazl @ qdio. ac. cn &amp; Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao, China &amp; Laoshan Laboratory, Qingdao, China</mods:affiliation>
<mods:nameIdentifier id="8BE18162C569AA49DB939CD9BC7AF1A1" type="email">yuanziming@qdio.ac.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="0FFDABA4250F3045879E718A6F2C50AF">text</mods:typeOfResource>
<mods:relatedItem id="F998D5432EFF551D97B47D5F56585D80" type="host">
<mods:titleInfo id="6A1661228C2851748D91DE9E1CD2D3B1">
<mods:title id="89B0F23E98EEC08E4EAD8A91FBCEBF86">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5B378EE3F0918B31F906453117AE2300">
<mods:date id="BB8125BD9FF771C367D07CE9F9A35A82">2023</mods:date>
<mods:detail id="FF8638913FB60E16C92EF1F950FCF60F" type="pubDate">
<mods:number id="426851D49D5F088CE84FC62CD36E936B">2023-11-01</mods:number>
</mods:detail>
<mods:detail id="9AF3B35BCC734777199EAED107ABC4A8" type="volume">
<mods:number id="1E525DD36F7912E708D98843021735EC">5361</mods:number>
</mods:detail>
<mods:detail id="CBF529D5CA228246033F77709F781471" type="issue">
<mods:number id="3CF82CAAB5730C78BA261B1595DA301E">2</mods:number>
</mods:detail>
<mods:extent id="37790C25DF64BE60E4AC0BB6C81D7024" unit="page">
<mods:start id="14B63621D7646EF7995D0E7EA4D5FA83">275</mods:start>
<mods:end id="3EE5298549B8435DB2EBEC8CFD7BDD7A">286</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="EA0B1FE673A5F8941568D55DAF122123">
<mods:url id="9154701C4AD2B3C433939057A9C5E6CE">https://www.mapress.com/zt/article/download/zootaxa.5361.2.8/52183</mods:url>
</mods:location>
<mods:classification id="B0B7F36E0A648D45C62786AC6119A2EA">journal article</mods:classification>
<mods:identifier id="A8DD9026D8B0A5BD414BDC0272B98D71" type="CLB-Dataset">277847</mods:identifier>
<mods:identifier id="B988CDAA00875819368061B65E061C3A" type="DOI">10.11646/zootaxa.5361.2.8</mods:identifier>
<mods:identifier id="8516FF76C97F06357BB72D967AD7B9AD" type="GBIF-Dataset">5bf93237-8248-486c-855f-c0f87553a671</mods:identifier>
<mods:identifier id="9DBD258D52C4116EA265025DFA26F1A6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="33A91A44F43ACB7503C6856C9F98FD8B" type="Zenodo-Dep">10146585</mods:identifier>
<mods:identifier id="1918066E92118F2469A8726F0F0EEFEF" type="ZooBank">9FB81B0C-412B-4978-B2CF-4151DA505F93</mods:identifier>
</mods:mods>
<treatment id="039E8789496B2F1FFF59F837FC9CDD76" LSID="urn:lsid:plazi:treatment:039E8789496B2F1FFF59F837FC9CDD76" httpUri="http://treatment.plazi.org/id/039E8789496B2F1FFF59F837FC9CDD76" lastPageId="6" lastPageNumber="281" pageId="1" pageNumber="276">
<subSubSection id="C32D6514496B2F18FF59F837FE28DDB2" box="[151,395,1933,1959]" pageId="1" pageNumber="276" type="nomenclature">
<paragraph id="8B88369F496B2F18FF59F837FE28DDB2" blockId="1.[151,395,1933,1959]" box="[151,395,1933,1959]" pageId="1" pageNumber="276">
<heading id="D0C081F3496B2F18FF59F837FE28DDB2" bold="true" box="[151,395,1933,1959]" fontSize="11" level="1" pageId="1" pageNumber="276" reason="1">
<emphasis id="B943EA8D496B2F18FF59F837FE28DDB2" bold="true" box="[151,395,1933,1959]" pageId="1" pageNumber="276">
<taxonomicName id="4C374D1C496B2F18FF59F837FE85DDB2" authority="Yuan &amp; Jiang &amp; Sha, 2023" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[151,294,1933,1959]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="mazu" status="sp. nov.">
<emphasis id="B943EA8D496B2F18FF59F837FE85DDB2" bold="true" box="[151,294,1933,1959]" italics="true" pageId="1" pageNumber="276">Cymo mazu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27057F6496B2F18FEE3F837FE28DDB2" box="[301,395,1933,1959]" pageId="1" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32D6514496B2F1BFF59F86FFE77DAC0" lastPageId="2" lastPageNumber="277" pageId="1" pageNumber="276" type="materials_examined">
<paragraph id="8B88369F496B2F1BFF59F86FFE77DAC0" blockId="1.[151,1436,2004,2031]" lastBlockId="2.[151,1437,150,753]" lastPageId="2" lastPageNumber="277" pageId="1" pageNumber="276">
<emphasis id="B943EA8D496B2F18FF59F86FFEE6DDFB" bold="true" box="[151,325,2004,2031]" pageId="1" pageNumber="276">Type material.</emphasis>
<materialsCitation id="3B5F3CC2496B2F18FE85F86FFA3FDDFA" box="[331,1436,2004,2031]" collectionCode="MBM" country="China" location="Fiery Cross Reef" municipality="Nansha Islands" pageId="1" pageNumber="276" specimenCode="MBM287035" specimenCount="1" typeStatus="holotype">
<typeStatus id="548C883D496B2F18FE85F86FFE1BDDFA" box="[331,440,2005,2031]" pageId="1" pageNumber="276" type="holotype">Holotype</typeStatus>
: 
<specimenCode id="DB919EE4496B2F18FE0CF86EFDFDDDFA" box="[450,606,2004,2031]" collectionCode="MBM" country="0" httpUri="http://grbio.org/cool/fc7c-p25f" name="San Jose State University, Museum of Birds and Mammals" pageId="1" pageNumber="276">MBM287035</specimenCode>
, male, CW 
<quantity id="4CCF9B7A496B2F18FD2EF86FFC9BDDFA" box="[736,824,2005,2031]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="276" unit="mm" value="4.1">4.1 mm</quantity>
, CL 
<quantity id="4CCF9B7A496B2F18FCA2F86FFC67DDFA" box="[876,964,2005,2031]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="276" unit="mm" value="3.8">3.8 mm</quantity>
, 
<location id="8EE86044496B2F18FC01F86FFB33DDFA" LSID="urn:lsid:plazi:treatment:039E8789496B2F1FFF59F837FC9CDD76:8EE86044496B2F18FC01F86FFB33DDFA" box="[975,1168,2005,2031]" country="China" municipality="Nansha Islands" name="Fiery Cross Reef" pageId="1" pageNumber="276">Fiery Cross Reef</location>
, 
<collectingMunicipality id="6BECACE5496B2F18FB55F86FFAE4DDFA" box="[1179,1351,2005,2031]" pageId="1" pageNumber="276">Nansha Islands</collectingMunicipality>
, 
<collectingCountry id="F320760F496B2F18FA9CF86FFA3BDDFA" box="[1362,1432,2005,2031]" name="China" pageId="1" pageNumber="276">China</collectingCountry>
,
</materialsCitation>
<geoCoordinate id="EE03505849682F1BFF59FF2CFEA7DAA4" box="[151,260,150,177]" degrees="112" direction="east" minutes="59" orientation="longitude" pageId="2" pageNumber="277" precision="925" value="112.98333">112°59’E</geoCoordinate>
, 
<geoCoordinate id="EE03505849682F1BFEDDFF2DFECBDAA4" box="[275,360,151,177]" degrees="9" direction="north" minutes="39" orientation="latitude" pageId="2" pageNumber="277" precision="925" value="9.65">9°39’N</geoCoordinate>
, 
<quantity id="4CCF9B7A49682F1BFEB8FF2DFE05DAA4" box="[374,422,151,177]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="277" unit="m" value="3.0">3 m</quantity>
, in 
<taxonomicName id="4C374D1C49682F1BFE1BFF2DFCFBDAA4" authority="(Brook, 1892)" baseAuthorityName="Brook" baseAuthorityYear="1892" box="[469,856,151,177]" class="Anthozoa" family="Acroporidae" genus="Acropora" kingdom="Animalia" order="Scleractinia" pageId="2" pageNumber="277" phylum="Cnidaria" rank="species" species="latistella">
<emphasis id="B943EA8D49682F1BFE1BFF2DFD08DAA4" box="[469,683,151,177]" italics="true" pageId="2" pageNumber="277">Acropora latistella</emphasis>
(
<bibRefCitation id="EFA64B6E49682F1BFD72FF2DFCF2DAA4" author="Brook, G." box="[700,849,151,177]" pageId="2" pageNumber="277" pagination="451 - 465" refId="ref5831" refString="Brook, G. (1892) Preliminary descriptions of new species of Madrepora in the collections of the British Museum. Part II. Annals and Magazine of Natural History, Series 6, 10 (60), 451 - 465. https: // doi. org / 10.1080 / 00222939208677447" type="journal article" year="1892">Brook, 1892</bibRefCitation>
)
</taxonomicName>
, NS-YS-2022-1139, coll. Shaobo Ma, Yuli Sun &amp; Ziming Yuan, 
<date id="FF89105F49682F1BFEF9FF01FE73DAC0" box="[311,464,187,213]" pageId="2" pageNumber="277" value="2022-05-13">13 May 2022</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C32D651449682F1BFF09FF65FCD2DB54" pageId="2" pageNumber="277" type="etymology">
<paragraph id="8B88369F49682F1BFF09FF65FCD2DB54" blockId="2.[151,1437,150,753]" pageId="2" pageNumber="277">
<emphasis id="B943EA8D49682F1BFF09FF65FEEDDAEC" bold="true" box="[199,334,223,249]" pageId="2" pageNumber="277">Etymology.</emphasis>
Named after Mazu, a legendary Chinese woman, Mo Lin (A.D. 960–987) from the Song Dynasty, who made significant contributions to the rescue of shipwrecked people. In 
<collectingCountry id="F320760F49682F1BFC26FEB9FB8DDB08" box="[1000,1070,259,285]" name="China" pageId="2" pageNumber="277">China</collectingCountry>
, she is widely worshipped as the sea goddess who protects fishermen and sailors on their voyages.
</paragraph>
</subSubSection>
<subSubSection id="C32D651449682F1BFF09FEF0FD3BDBC4" pageId="2" pageNumber="277" type="diagnosis">
<paragraph id="8B88369F49682F1BFF09FEF0FD3BDBC4" blockId="2.[151,1437,150,753]" pageId="2" pageNumber="277">
<emphasis id="B943EA8D49682F1BFF09FEF0FEE1DB71" bold="true" box="[199,322,330,356]" pageId="2" pageNumber="277">Diagnosis.</emphasis>
Carapace slightly wider than long, dorsal surface smooth, with sharp granules on orbital region, frontal regions, gastric regions, anterolateral regions and posterolateral regions. Chelipeds with strong spines on outer surface of palm; large cheliped fingers totally white, small cheliped fingers white but with pale brown block. Endopodite of maxilliped 1 strongly concave.
</paragraph>
</subSubSection>
<subSubSection id="C32D651449682F1DFF09FE60FF51D948" lastPageId="4" lastPageNumber="279" pageId="2" pageNumber="277" type="description">
<paragraph id="8B88369F49682F1BFF09FE60FDBDD8E4" blockId="2.[151,1437,150,753]" pageId="2" pageNumber="277">
<emphasis id="B943EA8D49682F1BFF09FE60FEFADBE1" bold="true" box="[199,345,474,500]" pageId="2" pageNumber="277">Description.</emphasis>
Carapace (
<figureCitation id="130C2A1A49682F1BFE18FE61FD80DBE0" box="[470,547,475,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1627,1651]" captionTargetBox="[310,1278,810,1601]" captionTargetId="figure-257@2.[310,1278,810,1601]" captionTargetPageId="2" captionText="FIGURE 1. Colour in life, Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10146587" httpUri="https://zenodo.org/record/10146587/files/figure.png" pageId="2" pageNumber="277">Figs. 1</figureCitation>
, 
<figureCitation id="130C2A1A49682F1BFDFEFE61FDD6DBE0" box="[560,629,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1808,1832]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-17@3.[151,1436,181,1784]" captionTargetPageId="3" captionText="FIGURE 2. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035: A, dorsal habitus; B, carapace; C, frontal view; D, thoracic sternites and pleon; E, chelipeds. Scale bar A–C = 2 mm, D–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146589" httpUri="https://zenodo.org/record/10146589/files/figure.png" pageId="2" pageNumber="277">2A, B</figureCitation>
, 
<figureCitation id="130C2A1A49682F1BFD4EFE61FD07DBE0" box="[640,676,475,501]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="2" pageNumber="277">3A</figureCitation>
) subcircular, slightly wider than long, armed with long setae, denser on front regions; dorsal flat and smooth, regions boundaries unclear; sharp granules on front, orbital region, gastric regions, anterolateral and posterolateral surfaces. Front (
<figureCitation id="130C2A1A49682F1BFCD9FD99FCCDD828" box="[791,878,547,573]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="2" pageNumber="277">Fig. 3B</figureCitation>
) approximately 0.4× CW, divided into two lobes by wide v-shaped notch, margin oblique, with small spines, tip more-or-less truncate. Dorsal inner orbital angle with spine, separated from front by notch; dorsal orbital margin with submedian fissure; outer orbital angle and anterolateral margin confluent; infraorbital tooth sharp. Eyestalk short, with small granules. Anterolateral margin convex, lined with spines; posterolateral margin nearly straight, longer than anterolateral margin. Subhepatic and pterygostomial regions granulated.
</paragraph>
<caption id="DF48661749682F1BFF59F9E1FA25DC66" ID-DOI="http://doi.org/10.5281/zenodo.10146587" ID-Zenodo-Dep="10146587" box="[151,1414,1627,1652]" httpUri="https://zenodo.org/record/10146587/files/figure.png" pageId="2" pageNumber="277" startId="2.[151,250,1627,1651]" targetBox="[310,1278,810,1601]" targetPageId="2" targetType="figure">
<paragraph id="8B88369F49682F1BFF59F9E1FA25DC66" blockId="2.[151,1414,1627,1652]" box="[151,1414,1627,1652]" pageId="2" pageNumber="277">
<emphasis id="B943EA8D49682F1BFF59F9E1FEAEDC66" bold="true" box="[151,269,1627,1651]" pageId="2" pageNumber="277">FIGURE 1</emphasis>
. Colour in life, 
<taxonomicName id="4C374D1C49682F1BFE61F9E6FD86DC66" authority="Yuan &amp; Jiang &amp; Sha, 2023" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[431,549,1628,1651]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="277" phylum="Arthropoda" rank="species" species="mazu" status="sp. nov.">
<emphasis id="B943EA8D49682F1BFE61F9E6FD86DC66" box="[431,549,1628,1651]" italics="true" pageId="2" pageNumber="277">Cymo mazu</emphasis>
</taxonomicName>
<emphasis id="B943EA8D49682F1BFDE2F9E6FDDFDC61" bold="true" box="[556,636,1628,1652]" pageId="2" pageNumber="277">
<taxonomicNameLabel id="A27057F649682F1BFDE2F9E6FDDFDC61" box="[556,636,1628,1652]" pageId="2" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035. Scale bar = 2 mm.
</paragraph>
</caption>
<paragraph id="8B88369F49682F1BFF09F97CFE25DD59" blockId="2.[151,1436,1734,1976]" pageId="2" pageNumber="277">
Antennular fossae (
<figureCitation id="130C2A1A49682F1BFE51F97CFE57DCF5" box="[415,500,1734,1760]" captionStart="FIGURE 2" captionStartId="3.[151,250,1808,1832]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-17@3.[151,1436,181,1784]" captionTargetPageId="3" captionText="FIGURE 2. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035: A, dorsal habitus; B, carapace; C, frontal view; D, thoracic sternites and pleon; E, chelipeds. Scale bar A–C = 2 mm, D–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146589" httpUri="https://zenodo.org/record/10146589/files/figure.png" pageId="2" pageNumber="277">Fig. 2C</figureCitation>
) subcircular, antennule situated obliquely. Basal segment of antenna subrectangular, anterolateral angle produced. Endopodite of maxilliped 1 (
<figureCitation id="130C2A1A49682F1BFCE2F950FC21DD11" box="[812,898,1770,1796]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="2" pageNumber="277">Fig. 3C</figureCitation>
) strongly concave. Maxilliped 3 (
<figureCitation id="130C2A1A49682F1BFACFF950FAC7DD11" box="[1281,1380,1770,1796]" captionStart="FIGURE 2" captionStartId="3.[151,250,1808,1832]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-17@3.[151,1436,181,1784]" captionTargetPageId="3" captionText="FIGURE 2. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035: A, dorsal habitus; B, carapace; C, frontal view; D, thoracic sternites and pleon; E, chelipeds. Scale bar A–C = 2 mm, D–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146589" httpUri="https://zenodo.org/record/10146589/files/figure.png" pageId="2" pageNumber="277">Figs. 2C</figureCitation>
, 
<figureCitation id="130C2A1A49682F1BFABFF950FA37DD11" box="[1393,1428,1770,1796]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="2" pageNumber="277">3D</figureCitation>
) completely covering buccal frame; merus subpentagonal, dorsal margin concave, inner margin with prominent lobe; ischium subquadrate.
</paragraph>
<paragraph id="8B88369F49682F1BFF09F8ECFC11DDAD" blockId="2.[151,1436,1734,1976]" pageId="2" pageNumber="277">Male thoracic sternum (Fig. D) smooth; sternites 1, 2 fused; suture between sternites 2, 3 and 6, 7 wide; suture of sternites 3 and 4 present on lateral edges, continuing via transverse depression; sternite 4 with distinct long median groove; press-button located on posterior of thoracic sternite 5.</paragraph>
<caption id="DF48661749692F1AFF59F8AAFBC9DD59" ID-DOI="http://doi.org/10.5281/zenodo.10146589" ID-Zenodo-Dep="10146589" httpUri="https://zenodo.org/record/10146589/files/figure.png" pageId="3" pageNumber="278" startId="3.[151,250,1808,1832]" targetBox="[151,1436,181,1784]" targetPageId="3" targetType="figure">
<paragraph id="8B88369F49692F1AFF59F8AAFBC9DD59" blockId="3.[151,1436,1808,1868]" pageId="3" pageNumber="278">
<emphasis id="B943EA8D49692F1AFF59F8AAFEB7DD3D" bold="true" box="[151,276,1808,1832]" pageId="3" pageNumber="278">FIGURE 2.</emphasis>
<taxonomicName id="4C374D1C49692F1AFED5F8ABFE31DD3D" authority="Yuan &amp; Jiang &amp; Sha, 2023" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[283,402,1809,1832]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="mazu" status="sp. nov.">
<emphasis id="B943EA8D49692F1AFED5F8ABFE31DD3D" box="[283,402,1809,1832]" italics="true" pageId="3" pageNumber="278">Cymo mazu</emphasis>
</taxonomicName>
<emphasis id="B943EA8D49692F1AFE54F8AAFE4FDD3D" bold="true" box="[410,492,1808,1832]" pageId="3" pageNumber="278">
<taxonomicNameLabel id="A27057F649692F1AFE54F8AAFE4FDD3D" box="[410,492,1808,1832]" pageId="3" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035: A, dorsal habitus; B, carapace; C, frontal view; D, thoracic sternites and pleon; E, chelipeds. Scale bar A–C = 2 mm, D–E = 1 mm.
</paragraph>
</caption>
<paragraph id="8B88369F496E2F1DFF09FF2DFEDADB9C" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="279">
Chelipeds (
<figureCitation id="130C2A1A496E2F1DFE87FF2DFE0FDAA4" box="[329,428,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1808,1832]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-17@3.[151,1436,181,1784]" captionTargetPageId="3" captionText="FIGURE 2. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035: A, dorsal habitus; B, carapace; C, frontal view; D, thoracic sternites and pleon; E, chelipeds. Scale bar A–C = 2 mm, D–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146589" httpUri="https://zenodo.org/record/10146589/files/figure.png" pageId="4" pageNumber="279">Figs. 2E</figureCitation>
, 
<figureCitation id="130C2A1A496E2F1DFE74FF2DFE7ADAA4" box="[442,473,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="4" pageNumber="279">3F</figureCitation>
) markedly unequal, robust and covered with long setae; merus short, granulated on ventral and outer surface; carpus subsphaeroidal, with spines on outer and dorsal surface; palm covered with granules evenly arranged, forming larger, sharp spines on dorsal surface and dorsal part of outer surface, mixed with small granules, transitioning to smaller round granules on inner surface, ventral surface and ventral part of outer surface. Fingers deflexed, dorsal surface of dactylus lined with sharp spines. In preservation, finger white but with very pale brown block, more prominent on minor cheliped, almost invisible on major cheliped; with gape when closed, fingertips hollowed.
</paragraph>
<paragraph id="8B88369F496E2F1DFF09FE29FD5BDBE0" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="279">
Ambulatory legs (
<figureCitation id="130C2A1A496E2F1DFE40FE29FE7ADBB8" box="[398,473,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1627,1651]" captionTargetBox="[310,1278,810,1601]" captionTargetId="figure-257@2.[310,1278,810,1601]" captionTargetPageId="2" captionText="FIGURE 1. Colour in life, Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10146587" httpUri="https://zenodo.org/record/10146587/files/figure.png" pageId="4" pageNumber="279">Figs. 1</figureCitation>
, 
<figureCitation id="130C2A1A496E2F1DFE2AFE29FDAADBB8" box="[484,521,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1808,1832]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-17@3.[151,1436,181,1784]" captionTargetPageId="3" captionText="FIGURE 2. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035: A, dorsal habitus; B, carapace; C, frontal view; D, thoracic sternites and pleon; E, chelipeds. Scale bar A–C = 2 mm, D–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146589" httpUri="https://zenodo.org/record/10146589/files/figure.png" pageId="4" pageNumber="279">2A</figureCitation>
, 
<figureCitation id="130C2A1A496E2F1DFDDDFE29FD91DBB8" box="[531,562,403,429]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="4" pageNumber="279">3E</figureCitation>
) short, flat, covered with long setae; merus with spines on anterior and posterior margins; carpus short, spines present on anterior margin; propodus with spines on anterior margin; dactylus strong, hooked, with developed dactylo-propodal articulation.
</paragraph>
<paragraph id="8B88369F496E2F1DFF09FE45FDBED828" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="279">
Male pleon smooth (
<figureCitation id="130C2A1A496E2F1DFE69FE45FDABD80C" box="[423,520,511,537]" captionStart="FIGURE 2" captionStartId="3.[151,250,1808,1832]" captionTargetBox="[151,1436,181,1784]" captionTargetId="figure-17@3.[151,1436,181,1784]" captionTargetPageId="3" captionText="FIGURE 2. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035: A, dorsal habitus; B, carapace; C, frontal view; D, thoracic sternites and pleon; E, chelipeds. Scale bar A–C = 2 mm, D–E = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146589" httpUri="https://zenodo.org/record/10146589/files/figure.png" pageId="4" pageNumber="279">Figs. 2D</figureCitation>
, 
<figureCitation id="130C2A1A496E2F1DFDDFFE45FD96D80C" box="[529,565,511,537]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="4" pageNumber="279">3G</figureCitation>
), pleonite 3 widest; pleonites 3–5 fused with visible suture; telson subtriangular, tip round, slightly wider than long.
</paragraph>
<paragraph id="8B88369F496E2F1DFF09FDFDFF46D8BC" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="279">
G1 (
<figureCitation id="130C2A1A496E2F1DFF32FDFDFE34D874" box="[252,407,583,609]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="4" pageNumber="279">Figs. 3H, I, J</figureCitation>
) slender, slightly sinuous laterally; inner and outer surface with spines, distal lobe short, deflexed. G2 (
<figureCitation id="130C2A1A496E2F1DFEF0FDD1FE3BD890" box="[318,408,619,645]" captionStart="FIGURE 3" captionStartId="5.[151,250,1812,1836]" captionTargetBox="[160,1426,321,1779]" captionTargetId="figure-51@5.[151,1436,309,1788]" captionTargetPageId="5" captionText="FIGURE 3. Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035:A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="4" pageNumber="279">Fig. 3K</figureCitation>
) small, length about 0.4× length of G1, distal part slightly upturned, subdistal part with spines.
</paragraph>
<paragraph id="8B88369F496E2F1DFF09FD08FF51D948" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="279">
<emphasis id="B943EA8D496E2F1DFF09FD08FE27D8D9" bold="true" box="[199,388,690,716]" pageId="4" pageNumber="279">Live coloration.</emphasis>
Body white to ivory, enveloped in long setae ranging from yellow to rufous, intermixed with bright red setae on carapace. Gastric, cardiac and intestinal regions adorned with fluorescent green vermicular pattern, interval being rufous. Cornea blue-green. Coloration of both cheliped fingers similar to that of palm, with no notable differentiation. Preserved in alcohol, almost entirely white, with only faint patches of brown discernible (
<figureCitation id="130C2A1A496E2F1DFF51FCF9FF42D948" box="[159,225,835,861]" captionStart="FIGURE 1" captionStartId="2.[151,250,1627,1651]" captionTargetBox="[310,1278,810,1601]" captionTargetId="figure-257@2.[310,1278,810,1601]" captionTargetPageId="2" captionText="FIGURE 1. Colour in life, Cymo mazu sp. nov., holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035. Scale bar = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.10146587" httpUri="https://zenodo.org/record/10146587/files/figure.png" pageId="4" pageNumber="279">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32D6514496E2F1FFF09FCDDFC9CDD76" lastPageId="6" lastPageNumber="281" pageId="4" pageNumber="279" type="discussion">
<paragraph id="8B88369F496E2F1DFF09FCDDFDC0DE20" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="279">
<emphasis id="B943EA8D496E2F1DFF09FCDDFE98D994" bold="true" box="[199,315,871,897]" pageId="4" pageNumber="279">Remarks.</emphasis>
<taxonomicName id="4C374D1C496E2F1DFE8DFCDDFE6BD995" authority="Yuan &amp; Jiang &amp; Sha, 2023" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[323,456,871,897]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu" status="sp. nov.">
<emphasis id="B943EA8D496E2F1DFE8DFCDDFE6BD995" box="[323,456,871,897]" italics="true" pageId="4" pageNumber="279">Cymo mazu</emphasis>
</taxonomicName>
<emphasis id="B943EA8D496E2F1DFE1EFCDDFD88D994" bold="true" box="[464,555,871,897]" pageId="4" pageNumber="279">
<taxonomicNameLabel id="A27057F6496E2F1DFE1EFCDDFD88D994" box="[464,555,871,897]" pageId="4" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
distinctly differs from its congeners by several unique features: the carapace, despite being smooth, features anterolateral spines, chelipeds armed with well-arranged, potent, sharp spines, a pale carapace embellished with a fluorescent pattern, and the particularly concave distal end of the maxilliped 1 endopodite. Notably, the G1 distal lobe of 
<taxonomicName id="4C374D1C496E2F1DFDBEFC69FD68D9F9" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[624,715,979,1005]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496E2F1DFDBEFC69FD68D9F9" box="[624,715,979,1005]" italics="true" pageId="4" pageNumber="279">C. mazu</emphasis>
</taxonomicName>
does not have the elongation observed in other species of 
<taxonomicName id="4C374D1C496E2F1DFA9DFC69FA37D9F8" authorityName="de Haan" authorityYear="1833" box="[1363,1428,979,1005]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B943EA8D496E2F1DFA9DFC69FA37D9F8" box="[1363,1428,979,1005]" italics="true" pageId="4" pageNumber="279">Cymo</emphasis>
</taxonomicName>
; rather, it is present as a short, deflexed opening lacking subdistal setae. This distinction could also result from incomplete development of the 
<typeStatus id="548C883D496E2F1DFE34FBA1FDC3DE20" box="[506,608,1051,1077]" pageId="4" pageNumber="279" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8B88369F496E2F1DFF09FB85FE93DFF0" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="279">
While uniquely characterized in many respects, the new species also exhibits combined characteristics shared with congeners. 
<taxonomicName id="4C374D1C496E2F1DFE9CFBD9FE74DE69" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[338,471,1123,1149]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496E2F1DFE9CFBD9FE74DE69" box="[338,471,1123,1149]" italics="true" pageId="4" pageNumber="279">Cymo mazu</emphasis>
</taxonomicName>
bears the closest resemblance to 
<taxonomicName id="4C374D1C496E2F1DFC9DFBD9FC53DE68" box="[851,1008,1123,1149]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496E2F1DFC9DFBD9FC53DE68" box="[851,1008,1123,1149]" italics="true" pageId="4" pageNumber="279">C. deplanatus</emphasis>
</taxonomicName>
due to their similar setal distribution, smooth carapace and a spiny front. However, 
<taxonomicName id="4C374D1C496E2F1DFD6CFB3DFD5CDEB5" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[674,767,1159,1185]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496E2F1DFD6CFB3DFD5CDEB5" box="[674,767,1159,1185]" italics="true" pageId="4" pageNumber="279">C. mazu</emphasis>
</taxonomicName>
can be distinguished by: the carapace being slightly wider than long (vs. carapace longer than wide in 
<taxonomicName id="4C374D1C496E2F1DFD4AFB11FC83DED0" box="[644,800,1195,1221]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496E2F1DFD4AFB11FC83DED0" box="[644,800,1195,1221]" italics="true" pageId="4" pageNumber="279">C. deplanatus</emphasis>
</taxonomicName>
) (
<figureCitation id="130C2A1A496E2F1DFCF7FB11FC1DDED0" box="[825,958,1194,1221]" captionStart="FIGURE 5" captionStartId="8.[151,250,1868,1892]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-27@8.[151,1436,181,1845]" captionTargetPageId="8" captionText="FIGURE 5. Diagnostic features of Cymo: A, F, K, P, C. mazu sp. nov.; B, G, L, Q, C. deplanatus; C, H, M, R, C. quadrilobatus; D, I, N, S, C. andreossyi; E, J, O, T, C. melanodactylus; A–E, overall morphology and live coloration; F–G, front; K–O, major cheliped; P–T, maxilliped 1. Scale bars: A, J, L, M = 2 mm; B–D, N, O = 5 mm; E = 10 mm; F, P = 0.5 mm; G–I, K, Q–T = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146595" httpUri="https://zenodo.org/record/10146595/files/figure.png" pageId="4" pageNumber="279">Figs. 5A, B</figureCitation>
); frontal lobes lined with small spines (vs. larger and longer spines adorning frontal lobes in 
<taxonomicName id="4C374D1C496E2F1DFD07FB75FCC6DEFC" box="[713,869,1231,1257]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496E2F1DFD07FB75FCC6DEFC" box="[713,869,1231,1257]" italics="true" pageId="4" pageNumber="279">C. deplanatus</emphasis>
</taxonomicName>
) (
<figureCitation id="130C2A1A496E2F1DFCB0FB75FBA3DEFC" box="[894,1024,1230,1257]" captionStart="FIGURE 5" captionStartId="8.[151,250,1868,1892]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-27@8.[151,1436,181,1845]" captionTargetPageId="8" captionText="FIGURE 5. Diagnostic features of Cymo: A, F, K, P, C. mazu sp. nov.; B, G, L, Q, C. deplanatus; C, H, M, R, C. quadrilobatus; D, I, N, S, C. andreossyi; E, J, O, T, C. melanodactylus; A–E, overall morphology and live coloration; F–G, front; K–O, major cheliped; P–T, maxilliped 1. Scale bars: A, J, L, M = 2 mm; B–D, N, O = 5 mm; E = 10 mm; F, P = 0.5 mm; G–I, K, Q–T = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146595" httpUri="https://zenodo.org/record/10146595/files/figure.png" pageId="4" pageNumber="279">Figs. 5F, G</figureCitation>
); chelipeds armed with large conical spines and hemispherical granules, evenly spaced and neatly arranged on the surface (vs. the irregular, smaller spines and granules with varying sizes and spacing on the chelipeds surface in 
<taxonomicName id="4C374D1C496E2F1DFC14FAADFBDBDF24" box="[986,1144,1303,1329]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496E2F1DFC14FAADFBDBDF24" box="[986,1144,1303,1329]" italics="true" pageId="4" pageNumber="279">C. deplanatus</emphasis>
</taxonomicName>
) (
<figureCitation id="130C2A1A496E2F1DFB5DFAADFAB9DF24" box="[1171,1306,1302,1329]" captionStart="FIGURE 5" captionStartId="8.[151,250,1868,1892]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-27@8.[151,1436,181,1845]" captionTargetPageId="8" captionText="FIGURE 5. Diagnostic features of Cymo: A, F, K, P, C. mazu sp. nov.; B, G, L, Q, C. deplanatus; C, H, M, R, C. quadrilobatus; D, I, N, S, C. andreossyi; E, J, O, T, C. melanodactylus; A–E, overall morphology and live coloration; F–G, front; K–O, major cheliped; P–T, maxilliped 1. Scale bars: A, J, L, M = 2 mm; B–D, N, O = 5 mm; E = 10 mm; F, P = 0.5 mm; G–I, K, Q–T = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146595" httpUri="https://zenodo.org/record/10146595/files/figure.png" pageId="4" pageNumber="279">Figs. 5K, L</figureCitation>
)); a deeply concave endopodite of maxilliped 1 (vs. absence of concavity on the maxilliped 1 endopodite of 
<taxonomicName id="4C374D1C496E2F1DFB3BFA81FA30DF40" box="[1269,1427,1339,1365]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496E2F1DFB3BFA81FA30DF40" box="[1269,1427,1339,1365]" italics="true" pageId="4" pageNumber="279">C. deplanatus</emphasis>
</taxonomicName>
) (
<figureCitation id="130C2A1A496E2F1DFF51FAE5FEBEDF6C" box="[159,285,1374,1401]" captionStart="FIGURE 5" captionStartId="8.[151,250,1868,1892]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-27@8.[151,1436,181,1845]" captionTargetPageId="8" captionText="FIGURE 5. Diagnostic features of Cymo: A, F, K, P, C. mazu sp. nov.; B, G, L, Q, C. deplanatus; C, H, M, R, C. quadrilobatus; D, I, N, S, C. andreossyi; E, J, O, T, C. melanodactylus; A–E, overall morphology and live coloration; F–G, front; K–O, major cheliped; P–T, maxilliped 1. Scale bars: A, J, L, M = 2 mm; B–D, N, O = 5 mm; E = 10 mm; F, P = 0.5 mm; G–I, K, Q–T = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146595" httpUri="https://zenodo.org/record/10146595/files/figure.png" pageId="4" pageNumber="279">Figs. 5P, Q</figureCitation>
). 
<taxonomicName id="4C374D1C496E2F1DFEFEFAE5FE10DF6D" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[304,435,1375,1401]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496E2F1DFEFEFAE5FE10DF6D" box="[304,435,1375,1401]" italics="true" pageId="4" pageNumber="279">Cymo mazu</emphasis>
</taxonomicName>
can also be differentiated from 
<taxonomicName id="4C374D1C496E2F1DFCDCFAE5FC2CDF6C" authorityName="Ho &amp; Ng" authorityYear="2005" box="[786,911,1375,1401]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="barunae">
<emphasis id="B943EA8D496E2F1DFCDCFAE5FC2CDF6C" box="[786,911,1375,1401]" italics="true" pageId="4" pageNumber="279">C. barunae</emphasis>
</taxonomicName>
, a species described from the South 
<collectingCountry id="F320760F496E2F1DFAEAFAE5FAC4DF6C" box="[1316,1383,1375,1401]" name="China" pageId="4" pageNumber="279">China</collectingCountry>
Sea, in features like those of 
<taxonomicName id="4C374D1C496E2F1DFE63FA39FDE9DF88" box="[429,586,1411,1437]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496E2F1DFE63FA39FDE9DF88" box="[429,586,1411,1437]" italics="true" pageId="4" pageNumber="279">C. deplanatus</emphasis>
</taxonomicName>
. Compared with 
<taxonomicName id="4C374D1C496E2F1DFCDFFA39FCCDDF89" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[785,878,1411,1437]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496E2F1DFCDFFA39FCCDDF89" box="[785,878,1411,1437]" italics="true" pageId="4" pageNumber="279">C. mazu</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C374D1C496E2F1DFCB0FA39FBB8DF88" box="[894,1051,1411,1437]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496E2F1DFCB0FA39FBB8DF88" box="[894,1051,1411,1437]" italics="true" pageId="4" pageNumber="279">C. deplanatus</emphasis>
</taxonomicName>
possesses a proportionally longer carapace, longer and more pronounced but fewer frontal spines, and the absence of the concavity on the maxilliped 1 endopodite.
</paragraph>
<paragraph id="8B88369F496E2F1DFF09FA55FB9FDCF4" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="279">
The morphology of the maxilliped 1 endopodite is an effective diagnostic characteristic for species of 
<taxonomicName id="4C374D1C496E2F1DFA95FA55FA3FDC1C" authorityName="de Haan" authorityYear="1833" box="[1371,1436,1519,1545]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="genus">
<emphasis id="B943EA8D496E2F1DFA95FA55FA3FDC1C" box="[1371,1436,1519,1545]" italics="true" pageId="4" pageNumber="279">Cymo</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFA64B6E496E2F1DFF05F9A9FE63DC38" author="Galil, B. &amp; Vannini, M." box="[203,448,1555,1581]" pageId="4" pageNumber="279" pagination="21 - 56" refId="ref6211" refString="Galil, B. &amp; Vannini, M. (1990) Research on the coast of Somalia. Xanthidae, Trapeziidae, Carpiliidae, Menippidae (Crustacea Brachyura). Tropical Zoology, 3, 21 - 56. https: // doi. org / 10.1080 / 03946975.1990.10539447" type="journal article" year="1990">Galil &amp; Vannini 1990</bibRefCitation>
). In this regard, 
<taxonomicName id="4C374D1C496E2F1DFDB8F9A9FD71DC39" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[630,722,1555,1581]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496E2F1DFDB8F9A9FD71DC39" box="[630,722,1555,1581]" italics="true" pageId="4" pageNumber="279">C. mazu</emphasis>
</taxonomicName>
, with its significantly concave maxilliped 1 endopodite, closely resembles 
<taxonomicName id="4C374D1C496E2F1DFEDEF98DFE1FDC44" authorityName="Galil &amp; Vannini" authorityYear="1990" box="[272,444,1591,1617]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="lanatopodus">
<emphasis id="B943EA8D496E2F1DFEDEF98DFE1FDC44" box="[272,444,1591,1617]" italics="true" pageId="4" pageNumber="279">C. lanatopodus</emphasis>
</taxonomicName>
. However, 
<taxonomicName id="4C374D1C496E2F1DFD8CF98DFD3CDC45" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[578,671,1591,1617]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496E2F1DFD8CF98DFD3CDC45" box="[578,671,1591,1617]" italics="true" pageId="4" pageNumber="279">C. mazu</emphasis>
</taxonomicName>
can be differentiated by the following characteristics: the carapace has distinct setae (vs. nearly glabrous in 
<taxonomicName id="4C374D1C496E2F1DFDB7F9E1FC8BDC60" authorityName="Galil &amp; Vannini" authorityYear="1990" box="[633,808,1627,1653]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="lanatopodus">
<emphasis id="B943EA8D496E2F1DFDB7F9E1FC8BDC60" box="[633,808,1627,1653]" italics="true" pageId="4" pageNumber="279">C. lanatopodus</emphasis>
</taxonomicName>
); frontal lobes armed with spines (vs. bifurcated, flat and spineless frontal lobes in 
<taxonomicName id="4C374D1C496E2F1DFE25F9C5FD34DC8C" authorityName="Galil &amp; Vannini" authorityYear="1990" box="[491,663,1663,1689]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="lanatopodus">
<emphasis id="B943EA8D496E2F1DFE25F9C5FD34DC8C" box="[491,663,1663,1689]" italics="true" pageId="4" pageNumber="279">C. lanatopodus</emphasis>
</taxonomicName>
); cheliped surface with notable long spines and granules (vs. rough cheliped surface with small granules in 
<taxonomicName id="4C374D1C496E2F1DFDADF919FCB3DCA8" authorityName="Galil &amp; Vannini" authorityYear="1990" box="[611,784,1699,1725]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="lanatopodus">
<emphasis id="B943EA8D496E2F1DFDADF919FCB3DCA8" box="[611,784,1699,1725]" italics="true" pageId="4" pageNumber="279">C. lanatopodus</emphasis>
</taxonomicName>
). Additionally, compared to 
<taxonomicName id="4C374D1C496E2F1DFB92F919FB19DCA9" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[1116,1210,1699,1725]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496E2F1DFB92F919FB19DCA9" box="[1116,1210,1699,1725]" italics="true" pageId="4" pageNumber="279">C. mazu</emphasis>
</taxonomicName>
, the carapace of 
<taxonomicName id="4C374D1C496E2F1DFA4DF919FE82DCF4" authorityName="Galil &amp; Vannini" authorityYear="1990" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="lanatopodus">
<emphasis id="B943EA8D496E2F1DFA4DF919FE82DCF4" italics="true" pageId="4" pageNumber="279">C. lanatopodus</emphasis>
</taxonomicName>
presents a more elongated and inwardly tapered posterolateral margin.
</paragraph>
<paragraph id="8B88369F496E2F1CFF09F951FF7CDAEC" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1436,151,249]" lastPageId="5" lastPageNumber="280" pageId="4" pageNumber="279">
<emphasis id="B943EA8D496E2F1DFF09F951FEF1DD11" box="[199,338,1771,1797]" italics="true" pageId="4" pageNumber="279">
<taxonomicName id="4C374D1C496E2F1DFF09F951FEEDDD11" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[199,334,1771,1797]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="mazu">Cymo mazu</taxonomicName>
,
</emphasis>
while bearing resemblance to 
<taxonomicName id="4C374D1C496E2F1DFD62F951FCC9DD10" box="[684,874,1771,1797]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496E2F1DFD62F951FCC9DD10" box="[684,874,1771,1797]" italics="true" pageId="4" pageNumber="279">C. quadrilobatus</emphasis>
</taxonomicName>
in the spiny chelipeds and concave endopodite of maxilliped 1, can be distinguished by: carapace spines and granules conspicuously smaller than on the chelipeds, never grouped in clusters (vs. spines and granules on carapace comparable to those on chelipeds, both large and grouped in clusters in 
<taxonomicName id="4C374D1C496E2F1DFE52F8EDFDFEDD64" box="[412,605,1879,1905]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496E2F1DFE52F8EDFDFEDD64" box="[412,605,1879,1905]" italics="true" pageId="4" pageNumber="279">C. quadrilobatus</emphasis>
</taxonomicName>
); a simpler, bilobed front (vs. quadrilobed front in 
<taxonomicName id="4C374D1C496E2F1DFB7AF8EDFAD6DD64" box="[1204,1397,1879,1905]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496E2F1DFB7AF8EDFAD6DD64" box="[1204,1397,1879,1905]" italics="true" pageId="4" pageNumber="279">C. quadrilobatus</emphasis>
</taxonomicName>
); a near-straight carapace posterolateral margin (vs. rounded posterolateral margin contributing to a more circular carapace appearance in 
<taxonomicName id="4C374D1C496E2F1DFE6FF825FDFCDDAC" box="[417,607,1951,1977]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496E2F1DFE6FF825FDFCDDAC" box="[417,607,1951,1977]" italics="true" pageId="4" pageNumber="279">C. quadrilobatus</emphasis>
</taxonomicName>
); and a deeply concave endopodite of maxilliped 1 (vs. a slightly concave endopodite of maxilliped 
<quantity id="4CCF9B7A496E2F1DFE77F879FE47DDC8" box="[441,484,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="279" unit="in" value="1.0">1 in</quantity>
<taxonomicName id="4C374D1C496E2F1DFE25F879FD0ADDC8" box="[491,681,1987,2013]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496E2F1DFE25F879FD0ADDC8" box="[491,681,1987,2013]" italics="true" pageId="4" pageNumber="279">C. quadrilobatus</emphasis>
</taxonomicName>
). Similarly, the present new species can also be distinguished from the lesser-known species, 
<taxonomicName id="4C374D1C496F2F1CFE75FF2DFCB8DAA4" authority="Ortmann, 1893" authorityName="Ortmann" authorityYear="1893" box="[443,795,151,177]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="280" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="B943EA8D496F2F1CFE75FF2DFDC4DAA4" box="[443,615,151,177]" italics="true" pageId="5" pageNumber="280">C. tuberculatus</emphasis>
<bibRefCitation id="EFA64B6E496F2F1CFDA0FF2DFCB8DAA4" author="Ortmann, A." box="[622,795,151,177]" pageId="5" pageNumber="280" pagination="411 - 495" refId="ref7110" refString="Ortmann, A. (1893) Die decapoden-krebse des Strassburger Museums, mit besonderer Ber ¸ cksichtigung der von Herrn Dr. Doderlein bein Japan und bei den Liu-Kiu-Inseln gesammeltern und zur Zeit im Strassburger Museum aufbewahrten Formen. Zoologische Jahrbucher, Abtheilung fur Systematik, Geographie und Biologie der Thiere, 7, 411 - 495, pl. 17." type="journal article" year="1893">Ortmann, 1893</bibRefCitation>
</taxonomicName>
. The latter possesses a carapace and pereopods similar to 
<taxonomicName id="4C374D1C496F2F1CFF59FF01FEF7DAC0" box="[151,340,187,213]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="280" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496F2F1CFF59FF01FEF7DAC0" box="[151,340,187,213]" italics="true" pageId="5" pageNumber="280">C. quadrilobatus</emphasis>
</taxonomicName>
, characterized by nodules formed from aggregated granules, and a carapace length that exceeds its width.
</paragraph>
<caption id="DF486617496F2F1CFF59F8AEFAF7DD66" ID-DOI="http://doi.org/10.5281/zenodo.10146591" ID-Zenodo-Dep="10146591" httpUri="https://zenodo.org/record/10146591/files/figure.png" pageId="5" pageNumber="280" startId="5.[151,250,1812,1836]" targetBox="[160,1426,321,1779]" targetPageId="5" targetType="figure">
<paragraph id="8B88369F496F2F1CFF59F8AEFAF7DD66" blockId="5.[151,1437,1812,1908]" pageId="5" pageNumber="280">
<emphasis id="B943EA8D496F2F1CFF59F8AEFEB0DD39" bold="true" box="[151,275,1812,1836]" pageId="5" pageNumber="280">FIGURE 3.</emphasis>
<taxonomicName id="4C374D1C496F2F1CFED7F8AEFE2CDD39" authority="Yuan &amp; Jiang &amp; Sha, 2023" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[281,399,1812,1836]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="280" phylum="Arthropoda" rank="species" species="mazu" status="sp. nov.">
<emphasis id="B943EA8D496F2F1CFED7F8AEFE2CDD39" box="[281,399,1812,1836]" italics="true" pageId="5" pageNumber="280">Cymo mazu</emphasis>
</taxonomicName>
<emphasis id="B943EA8D496F2F1CFE58F8AEFE45DD39" bold="true" box="[406,486,1812,1836]" pageId="5" pageNumber="280">
<taxonomicNameLabel id="A27057F6496F2F1CFE58F8AEFE45DD39" box="[406,486,1812,1836]" pageId="5" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, CW 4.1 mm, CL 3.8 mm, MBM287035: A, carapace; B, front; C, maxilliped 1; D, maxilliped 3; E, pereopod 5; F, large cheliped; G, pleon; H, left G1, ventral view; I, left G1, distal part, ventral view; J, same, dorsal view; K, right G2, ventral view. Scale bar A, E, F = 1 mm, C, D, G = 0.5 mm, H = 0.2 mm, I–K = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8B88369F496C2F1FFF09FF2DFE13D80C" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="281">
Given the similar carapace contour, small dorsal spines or granules, and pale cheliped fingers, 
<taxonomicName id="4C374D1C496C2F1FFB25FF2DFAE5DAA5" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[1259,1350,151,177]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496C2F1FFB25FF2DFAE5DAA5" box="[1259,1350,151,177]" italics="true" pageId="6" pageNumber="281">C. mazu</emphasis>
</taxonomicName>
bears a superficial similarity to 
<taxonomicName id="4C374D1C496C2F1FFE51FF01FD94DAC0" box="[415,567,187,213]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="andreossyi">
<emphasis id="B943EA8D496C2F1FFE51FF01FD94DAC0" box="[415,567,187,213]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C374D1C496C2F1FFD8DFF01FD66DAC1" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[579,709,187,213]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496C2F1FFD8DFF01FD66DAC1" box="[579,709,187,213]" italics="true" pageId="6" pageNumber="281">Cymo mazu</emphasis>
</taxonomicName>
can be primarily distinguished by: carapace anterolateral margins lined with distinct spines (vs. anterolateral margin without spines in 
<taxonomicName id="4C374D1C496C2F1FFC5EFF65FB8ADAEC" box="[912,1065,223,249]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="andreossyi">
<emphasis id="B943EA8D496C2F1FFC5EFF65FB8ADAEC" box="[912,1065,223,249]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</taxonomicName>
); chelipeds armed with large and acute spines (vs. chelipeds with low and round granules, densely arranged); front margins armed with simple spines (vs. two of the front margins spines larger and bifurcated); and a markedly concave endopodite of maxilliped 1 (vs. endopodite of maxilliped 1 slightly concave in 
<taxonomicName id="4C374D1C496C2F1FFD67FEF1FCE1DB70" box="[681,834,331,357]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="andreossyi">
<emphasis id="B943EA8D496C2F1FFD67FEF1FCE1DB70" box="[681,834,331,357]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</taxonomicName>
). A notable variant is 
<taxonomicName id="4C374D1C496C2F1FFBF5FEF1FEFDDB9C" authority="Klunzinger, 1913" authorityName="Klunzinger" authorityYear="1913" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="variety" species="andreossyi" variety="maculata">
<emphasis id="B943EA8D496C2F1FFBF5FEF1FB5FDB70" box="[1083,1276,331,357]" italics="true" pageId="6" pageNumber="281">Cymo andreossyi</emphasis>
var. 
<emphasis id="B943EA8D496C2F1FFAFAFEF1FA3FDB70" box="[1332,1436,331,357]" italics="true" pageId="6" pageNumber="281">maculata</emphasis>
<bibRefCitation id="EFA64B6E496C2F1FFF59FED5FEFDDB9C" author="Klunzinger, C. B." box="[151,350,367,393]" pageId="6" pageNumber="281" pagination="97 - 402" refId="ref6442" refString="Klunzinger, C. B. (1913) Die Rundkrabben (Cyclometopa) des Roten Meeres. Abhandlungen der Kaiserlichen Leopoldisch- Carolinischen Deutschen Akademie der Naturforscher, 99 (2), 97 - 402, pls. 5 - 11. https: // doi. org / 10.5962 / bhl. title. 13073" type="journal article" year="1913">Klunzinger, 1913</bibRefCitation>
</taxonomicName>
, which is currently listed as a synonym for 
<taxonomicName id="4C374D1C496C2F1FFC9CFED5FC4FDB9C" box="[850,1004,367,393]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="andreossyi">
<emphasis id="B943EA8D496C2F1FFC9CFED5FC4FDB9C" box="[850,1004,367,393]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFA64B6E496C2F1FFBE4FED5FB70DB9C" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1066,1235,367,393]" pageId="6" pageNumber="281" pagination="1 - 286" refId="ref7056" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, Supplement 17, 1 - 286." type="journal article" year="2008">
Ng 
<emphasis id="B943EA8D496C2F1FFB9AFED5FB24DB9C" box="[1108,1159,367,393]" italics="true" pageId="6" pageNumber="281">et al</emphasis>
. 2008
</bibRefCitation>
). According to its original description, this variant exhibits a longer and straighter posterior carapace margin, a feature that aligns with the new species under discussion. However, 
<taxonomicName id="4C374D1C496C2F1FFD1BFE0DFBB6DBC4" authorityName="Klunzinger" authorityYear="1913" box="[725,1045,439,465]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="variety" species="andreossyi" variety="maculata">
<emphasis id="B943EA8D496C2F1FFD1BFE0DFCD3DBC4" box="[725,880,439,465]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
var. 
<emphasis id="B943EA8D496C2F1FFC63FE0DFBB6DBC4" box="[941,1045,439,465]" italics="true" pageId="6" pageNumber="281">maculata</emphasis>
</taxonomicName>
possesses a unique characteristic: a prominent angle at the junction of its anterior and posterior carapace margins. Further study might show it to represent a valid species.
</paragraph>
<paragraph id="8B88369F496C2F1FFF09FD99FEACD9B0" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="281">
The coloration of the cheliped fingers often serves as a diagnostic characteristic within this genus. 
<taxonomicName id="4C374D1C496C2F1FFAD7FD99FA3FD829" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[1305,1436,547,573]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496C2F1FFAD7FD99FA3FD829" box="[1305,1436,547,573]" italics="true" pageId="6" pageNumber="281">Cymo mazu</emphasis>
</taxonomicName>
can be readily differentiated from two morphologically similar species, 
<taxonomicName id="4C374D1C496C2F1FFC14FDFDFB0DD874" box="[986,1198,583,609]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylus">
<emphasis id="B943EA8D496C2F1FFC14FDFDFB0DD874" box="[986,1198,583,609]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C374D1C496C2F1FFB22FDFDFACCD875" authorityName="Morgan" authorityYear="1990" box="[1260,1391,583,609]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="cerasma">
<emphasis id="B943EA8D496C2F1FFB22FDFDFACCD875" box="[1260,1391,583,609]" italics="true" pageId="6" pageNumber="281">C. cerasma</emphasis>
</taxonomicName>
, by its overall lighter finger coloration. Both 
<taxonomicName id="4C374D1C496C2F1FFDBFFDD1FCE7D890" box="[625,836,619,645]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylus">
<emphasis id="B943EA8D496C2F1FFDBFFDD1FCE7D890" box="[625,836,619,645]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C374D1C496C2F1FFCB1FDD1FBA3D891" authorityName="Morgan" authorityYear="1990" box="[895,1024,619,645]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="cerasma">
<emphasis id="B943EA8D496C2F1FFCB1FDD1FBA3D891" box="[895,1024,619,645]" italics="true" pageId="6" pageNumber="281">C. cerasma</emphasis>
</taxonomicName>
present uniform black fingers, only showing white at the margin hollowed tips. Even in faded specimens of 
<taxonomicName id="4C374D1C496C2F1FFC0FFD35FB31D8BC" box="[961,1170,655,681]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylus">
<emphasis id="B943EA8D496C2F1FFC0FFD35FB31D8BC" box="[961,1170,655,681]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</taxonomicName>
observed in the present study, the cheliped fingers retain their black coloration. In contrast, the new species, when alive, possesses white or ivory fingers with no apparent color separation from the palm. In faded specimens, the fingers are white, exhibiting only faint brown patches, a condition more akin to that of 
<taxonomicName id="4C374D1C496C2F1FFCD5FD41FC7BD900" box="[795,984,763,789]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496C2F1FFCD5FD41FC7BD900" box="[795,984,763,789]" italics="true" pageId="6" pageNumber="281">C. quadrilobatus</emphasis>
</taxonomicName>
. It is noteworthy that 
<taxonomicName id="4C374D1C496C2F1FFB02FD41FA3FD900" box="[1228,1436,763,789]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylus">
<emphasis id="B943EA8D496C2F1FFB02FD41FA3FD900" box="[1228,1436,763,789]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C374D1C496C2F1FFF08FCA5FEE5D92D" authorityName="Morgan" authorityYear="1990" box="[198,326,799,825]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="cerasma">
<emphasis id="B943EA8D496C2F1FFF08FCA5FEE5D92D" box="[198,326,799,825]" italics="true" pageId="6" pageNumber="281">C. cerasma</emphasis>
</taxonomicName>
are highly similar, with the primary distinguishing feature seemingly being the broader carapace of the latter. However, a nearly circular carapace outline has also been observed in 
<taxonomicName id="4C374D1C496C2F1FFBC6FCF9FB6FD948" box="[1032,1228,835,861]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylu">
<emphasis id="B943EA8D496C2F1FFBC6FCF9FB6FD948" box="[1032,1228,835,861]" italics="true" pageId="6" pageNumber="281">C. melanodactylu</emphasis>
</taxonomicName>
s in this study, thus further inspection and confirmation of the identity and more detailed distinguishing features of the latter species are warranted.
</paragraph>
<paragraph id="8B88369F496C2F1FFF09FC15FA9DDED0" blockId="6.[151,1437,151,1221]" pageId="6" pageNumber="281">
To further illuminate the relationships between 
<taxonomicName id="4C374D1C496C2F1FFD29FC15FCE6D9DD" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[743,837,943,969]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496C2F1FFD29FC15FCE6D9DD" box="[743,837,943,969]" italics="true" pageId="6" pageNumber="281">C. mazu</emphasis>
</taxonomicName>
and other related species within the genus, we used COI sequences of 
<taxonomicName id="4C374D1C496C2F1FFEA7FC69FDA0D9F8" box="[361,515,979,1005]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="andreossyi">
<emphasis id="B943EA8D496C2F1FFEA7FC69FDA0D9F8" box="[361,515,979,1005]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C374D1C496C2F1FFDDCFC69FD0CD9F8" box="[530,687,979,1005]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496C2F1FFDDCFC69FD0CD9F8" box="[530,687,979,1005]" italics="true" pageId="6" pageNumber="281">C. deplanatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C374D1C496C2F1FFD70FC69FCB8D9F9" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[702,795,979,1005]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496C2F1FFD70FC69FCB8D9F9" box="[702,795,979,1005]" italics="true" pageId="6" pageNumber="281">C. mazu</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C374D1C496C2F1FFCE5FC69FC5ED9F8" box="[811,1021,979,1005]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylus">
<emphasis id="B943EA8D496C2F1FFCE5FC69FC5ED9F8" box="[811,1021,979,1005]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C374D1C496C2F1FFBF2FC69FB58D9F8" box="[1084,1275,979,1005]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496C2F1FFBF2FC69FB58D9F8" box="[1084,1275,979,1005]" italics="true" pageId="6" pageNumber="281">C. quadrilobatus</emphasis>
</taxonomicName>
for molecular analyses. The interspecific genetic distances of COI among these five species were all greater than 12.3%, with intraspecific genetic distances being less than 0.7% (
<tableCitation id="C6B50324496C2F1FFD22FBA1FCE6DE20" box="[748,837,1051,1077]" captionStart="TABLE 2" captionStartId="6.[151,239,1268,1292]" captionTargetId="graphics-539@6.[151,1436,1373,1571]" captionTargetPageId="6" captionText="TABLE 2. Genetic divergence of the COI sequences among five speciesof Cymo calculated using Kimura 2-parameter- corrected distances." httpUri="http://table.plazi.org/id/DF486617496C2F1FFF59FB4EFEC9DF25" pageId="6" pageNumber="281" tableUuid="DF486617496C2F1FFF59FB4EFEC9DF25">Table 2</tableCitation>
). In the phylogenetic trees (
<figureCitation id="130C2A1A496C2F1FFB4CFBA1FB63DE20" box="[1154,1216,1051,1077]" captionStart="FIGURE 4" captionStartId="7.[151,250,1016,1040]" captionTargetBox="[293,1293,182,988]" captionTargetId="figure-17@7.[293,1293,181,992]" captionTargetPageId="7" captionText="FIGURE 4. Phylogenetic relationships among five species of Cymo based on COI sequences. The BI tree with posterior probabilities and ML bootstrap values labeled (PP/BS)." figureDoi="http://doi.org/10.5281/zenodo.10146593" httpUri="https://zenodo.org/record/10146593/files/figure.png" pageId="6" pageNumber="281">Fig.4</figureCitation>
), individuals of the same species clustered together, exhibiting high support values (
<taxonomicName id="4C374D1C496C2F1FFCB3FB85FBBBDE4C" box="[893,1048,1087,1113]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="andreossyi">
<emphasis id="B943EA8D496C2F1FFCB3FB85FBBBDE4C" box="[893,1048,1087,1113]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</taxonomicName>
: 100/91; 
<taxonomicName id="4C374D1C496C2F1FFB47FB85FA84DE4C" box="[1161,1319,1087,1113]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496C2F1FFB47FB85FA84DE4C" box="[1161,1319,1087,1113]" italics="true" pageId="6" pageNumber="281">C. deplanatus</emphasis>
</taxonomicName>
: 100/100; 
<taxonomicName id="4C374D1C496C2F1FFF59FBD9FECADE68" box="[151,361,1123,1149]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylus">
<emphasis id="B943EA8D496C2F1FFF59FBD9FECADE68" box="[151,361,1123,1149]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</taxonomicName>
: 100/100; 
<taxonomicName id="4C374D1C496C2F1FFE2DFBD9FD01DE68" box="[483,674,1123,1149]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496C2F1FFE2DFBD9FD01DE68" box="[483,674,1123,1149]" italics="true" pageId="6" pageNumber="281">C. quadrilobatus</emphasis>
</taxonomicName>
: 100/99). The 
<typeStatus id="548C883D496C2F1FFC84FBD9FC0FDE68" box="[842,940,1123,1149]" pageId="6" pageNumber="281" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C374D1C496C2F1FFC1DFBD9FB8CDE69" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[979,1071,1123,1149]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496C2F1FFC1DFBD9FB8CDE69" box="[979,1071,1123,1149]" italics="true" pageId="6" pageNumber="281">C. mazu</emphasis>
</taxonomicName>
remained distinct from all other species, exhibiting the closest relationship to 
<taxonomicName id="4C374D1C496C2F1FFD4CFB3DFCB8DEB4" box="[642,795,1159,1185]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496C2F1FFD4CFB3DFCB8DEB4" box="[642,795,1159,1185]" italics="true" pageId="6" pageNumber="281">C. deplanatus</emphasis>
</taxonomicName>
(87/60). 
<taxonomicName id="4C374D1C496C2F1FFCB3FB3DFBC3DEB4" box="[893,1120,1159,1185]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496C2F1FFCB3FB3DFBC3DEB4" box="[893,1120,1159,1185]" italics="true" pageId="6" pageNumber="281">Cymo quadrilobatus</emphasis>
</taxonomicName>
clustered with 
<taxonomicName id="4C374D1C496C2F1FFACDFB3DFA3FDEB4" box="[1283,1436,1159,1185]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496C2F1FFACDFB3DFA3FDEB4" box="[1283,1436,1159,1185]" italics="true" pageId="6" pageNumber="281">C. deplanatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C374D1C496C2F1FFF09FB11FE81DED1" authorityName="Yuan &amp; Jiang &amp; Sha" authorityYear="2023" box="[199,290,1195,1221]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="mazu">
<emphasis id="B943EA8D496C2F1FFF09FB11FE81DED1" box="[199,290,1195,1221]" italics="true" pageId="6" pageNumber="281">C. mazu</emphasis>
</taxonomicName>
(78/-). Additionally, 
<taxonomicName id="4C374D1C496C2F1FFDC1FB11FD43DED0" box="[527,736,1195,1221]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylus">
<emphasis id="B943EA8D496C2F1FFDC1FB11FD43DED0" box="[527,736,1195,1221]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C374D1C496C2F1FFCD8FB11FC0CDED0" box="[790,943,1195,1221]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="andreossyi">
<emphasis id="B943EA8D496C2F1FFCD8FB11FC0CDED0" box="[790,943,1195,1221]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</taxonomicName>
formed a separate cluster (100/90).
</paragraph>
<caption id="DF486617496C2F1FFF59FB4EFEC9DF25" ID-Table-UUID="DF486617496C2F1FFF59FB4EFEC9DF25" httpUri="http://table.plazi.org/id/DF486617496C2F1FFF59FB4EFEC9DF25" pageId="6" pageNumber="281" startId="6.[151,239,1268,1292]" targetBox="[163,1412,1346,1564]" targetIsTable="true" targetPageId="6">
<paragraph id="8B88369F496C2F1FFF59FB4EFEC9DF25" pageId="6" pageNumber="281">
<emphasis id="B943EA8D496C2F1FFF59FB4EFEA9DF19" bold="true" box="[151,266,1268,1292]" pageId="6" pageNumber="281">TABLE 2.</emphasis>
Genetic divergence of the COI sequences among five 
<emphasis id="B943EA8D496C2F1FFC98FB4FFCC3DF19" box="[854,864,1269,1292]" italics="true" pageId="6" pageNumber="281">s</emphasis>
peciesof 
<emphasis id="B943EA8D496C2F1FFC0FFB4FFC5CDF19" box="[961,1023,1269,1292]" italics="true" pageId="6" pageNumber="281">Cymo</emphasis>
calculated using Kimura 2-parameter- corrected distances.
</paragraph>
</caption>
<paragraph id="8B88369F496C2F1FFEB7FAF8FAA4DC0E" pageId="6" pageNumber="281">
<table id="F937C43F496CD0E6FF6DFAF8FA27DC09" box="[163,1412,1346,1564]" gridcols="6" gridrows="6" pageId="6" pageNumber="281">
<tr id="350734DD496CD0E6FF6DFAF8FA27DF42" box="[163,1412,1346,1367]" gridrow="0" pageId="6" pageNumber="281" rowspan-0="1">
<th id="76D65DA1496CD0E6FEB7FAF8FE5DDF42" box="[377,510,1346,1367]" gridcol="1" gridrow="0" pageId="6" pageNumber="281">C. deplanatus</th>
<th id="76D65DA1496CD0E6FD81FAF8FD71DF42" box="[591,722,1346,1367]" gridcol="2" gridrow="0" pageId="6" pageNumber="281">C. andreossyi</th>
<th id="76D65DA1496CD0E6FCEBFAF8FC6BDF42" box="[805,968,1346,1367]" gridcol="3" gridrow="0" pageId="6" pageNumber="281">C. quadrilobatus</th>
<th id="76D65DA1496CD0E6FC35FAF8FBE9DF42" box="[1019,1098,1346,1367]" gridcol="4" gridrow="0" pageId="6" pageNumber="281">C. mazu</th>
<th id="76D65DA1496CD0E6FB1FFAF8FA27DF42" box="[1233,1412,1346,1367]" gridcol="5" gridrow="0" pageId="6" pageNumber="281">C. melanodactylus</th>
</tr>
<tr id="350734DD496CD0E6FF6DFAD3FA27DF6B" box="[163,1412,1385,1406]" gridrow="1" pageId="6" pageNumber="281" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76D65DA1496CD0E6FF6DFAD3FEF5DF6B" box="[163,342,1385,1406]" gridcol="0" gridrow="1" pageId="6" pageNumber="281">
<emphasis id="B943EA8D496C2F1FFF6DFAD3FE8BDF6B" box="[163,296,1385,1406]" italics="true" pageId="6" pageNumber="281">C. deplanatus</emphasis>
</th>
<td id="76D65DA1496CD0E6FEB7FAD3FE5DDF6B" box="[377,510,1385,1406]" gridcol="1" gridrow="1" pageId="6" pageNumber="281">0.001</td>
</tr>
<tr id="350734DD496CD0E6FF6DFA2AFA27DFB3" box="[163,1412,1424,1446]" gridrow="2" pageId="6" pageNumber="281" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="76D65DA1496CD0E6FF6DFA2AFEF5DFB3" box="[163,342,1424,1446]" gridcol="0" gridrow="2" pageId="6" pageNumber="281">
<emphasis id="B943EA8D496C2F1FFF6DFA2BFE85DFB0" box="[163,294,1424,1446]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</th>
<td id="76D65DA1496CD0E6FEB7FA2AFE5DDFB3" box="[377,510,1424,1446]" gridcol="1" gridrow="2" pageId="6" pageNumber="281">0.170</td>
<td id="76D65DA1496CD0E6FD81FA2AFD71DFB3" box="[591,722,1424,1446]" gridcol="2" gridrow="2" pageId="6" pageNumber="281">0.007</td>
</tr>
<tr id="350734DD496CD0E6FF6DFA02FA27DFD8" box="[163,1412,1464,1485]" gridrow="3" pageId="6" pageNumber="281" rowspan-4="1" rowspan-5="1">
<th id="76D65DA1496CD0E6FF6DFA02FEF5DFD8" box="[163,342,1464,1485]" gridcol="0" gridrow="3" pageId="6" pageNumber="281">
<emphasis id="B943EA8D496C2F1FFF6DFA02FEE5DFD8" box="[163,326,1464,1485]" italics="true" pageId="6" pageNumber="281">C. quadrilobatus</emphasis>
</th>
<td id="76D65DA1496CD0E6FEB7FA02FE5DDFD8" box="[377,510,1464,1485]" gridcol="1" gridrow="3" pageId="6" pageNumber="281">0.147</td>
<td id="76D65DA1496CD0E6FD81FA02FD71DFD8" box="[591,722,1464,1485]" gridcol="2" gridrow="3" pageId="6" pageNumber="281">0.132</td>
<td id="76D65DA1496CD0E6FCEBFA02FC6BDFD8" box="[805,968,1464,1485]" gridcol="3" gridrow="3" pageId="6" pageNumber="281">0.007</td>
</tr>
<tr id="350734DD496CD0E6FF6DFA65FA27DFE0" box="[163,1412,1503,1525]" gridrow="4" pageId="6" pageNumber="281" rowspan-5="1">
<th id="76D65DA1496CD0E6FF6DFA65FEF5DFE0" box="[163,342,1503,1525]" gridcol="0" gridrow="4" pageId="6" pageNumber="281">
<emphasis id="B943EA8D496C2F1FFF6DFA65FF51DFE0" box="[163,242,1503,1525]" italics="true" pageId="6" pageNumber="281">C. mazu</emphasis>
</th>
<td id="76D65DA1496CD0E6FEB7FA65FE5DDFE0" box="[377,510,1503,1525]" gridcol="1" gridrow="4" pageId="6" pageNumber="281">0.162</td>
<td id="76D65DA1496CD0E6FD81FA65FD71DFE0" box="[591,722,1503,1525]" gridcol="2" gridrow="4" pageId="6" pageNumber="281">0.187</td>
<td id="76D65DA1496CD0E6FCEBFA65FC6BDFE0" box="[805,968,1503,1525]" gridcol="3" gridrow="4" pageId="6" pageNumber="281">0.174</td>
<td id="76D65DA1496CD0E6FC35FA65FBE9DFE0" box="[1019,1098,1503,1525]" gridcol="4" gridrow="4" pageId="6" pageNumber="281">n/c</td>
</tr>
<tr id="350734DD496CD0E6FF6DF9BCFA27DC09" box="[163,1412,1542,1564]" gridrow="5" pageId="6" pageNumber="281">
<th id="76D65DA1496CD0E6FF6DF9BCFEF5DC09" box="[163,342,1542,1564]" gridcol="0" gridrow="5" pageId="6" pageNumber="281">
<emphasis id="B943EA8D496C2F1FFF6DF9BDFEF5DC0E" box="[163,342,1542,1564]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</th>
<td id="76D65DA1496CD0E6FEB7F9BCFE5DDC09" box="[377,510,1542,1564]" gridcol="1" gridrow="5" pageId="6" pageNumber="281">0.167</td>
<td id="76D65DA1496CD0E6FD81F9BCFD71DC09" box="[591,722,1542,1564]" gridcol="2" gridrow="5" pageId="6" pageNumber="281">0.123</td>
<td id="76D65DA1496CD0E6FCEBF9BCFC6BDC09" box="[805,968,1542,1564]" gridcol="3" gridrow="5" pageId="6" pageNumber="281">0.169</td>
<td id="76D65DA1496CD0E6FC35F9BCFBE9DC09" box="[1019,1098,1542,1564]" gridcol="4" gridrow="5" pageId="6" pageNumber="281">0.175</td>
<td id="76D65DA1496CD0E6FB1FF9BCFA27DC09" box="[1233,1412,1542,1564]" gridcol="5" gridrow="5" pageId="6" pageNumber="281">0.000</td>
</tr>
</table>
</paragraph>
<paragraph id="8B88369F496C2F1FFF09F9F6FC9CDD76" blockId="6.[151,1437,1612,1891]" pageId="6" pageNumber="281">
Species of 
<taxonomicName id="4C374D1C496C2F1FFE85F9F6FE2FDC73" authorityName="de Haan" authorityYear="1833" box="[331,396,1612,1638]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="genus">
<emphasis id="B943EA8D496C2F1FFE85F9F6FE2FDC73" box="[331,396,1612,1638]" italics="true" pageId="6" pageNumber="281">Cymo</emphasis>
</taxonomicName>
present superficially similar morphologies, which can sometimes make them difficult to distinguish. However, based on current observations, we propose that the overall morphology of the carapace, appearance of the setae, features of the frontal margin, surface appendages and fingers coloration of the chelipeds, and characteristics of the first maxilliped endopodite serve as compelling identification features. Additionally, the body and corneal coloration of these species when alive may also be considerable diagnostic features. Based on present observations, we have compiled a comparative summary of these diagnostic features for the new species for 
<taxonomicName id="4C374D1C496C2F1FFF59F89EFE91DD2B" box="[151,306,1828,1854]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="deplanatus">
<emphasis id="B943EA8D496C2F1FFF59F89EFE91DD2B" box="[151,306,1828,1854]" italics="true" pageId="6" pageNumber="281">C. deplanatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C374D1C496C2F1FFE8EF89EFE5EDD2B" box="[320,509,1828,1854]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="quadrilobatus">
<emphasis id="B943EA8D496C2F1FFE8EF89EFE5EDD2B" box="[320,509,1828,1854]" italics="true" pageId="6" pageNumber="281">C. quadrilobatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C374D1C496C2F1FFDC5F89EFD00DD2B" box="[523,675,1828,1854]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="andreossyi">
<emphasis id="B943EA8D496C2F1FFDC5F89EFD00DD2B" box="[523,675,1828,1854]" italics="true" pageId="6" pageNumber="281">C. andreossyi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C374D1C496C2F1FFD11F89EFC13DD2B" box="[735,944,1828,1854]" class="Malacostraca" family="Xanthidae" genus="Cymo" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="281" phylum="Arthropoda" rank="species" species="melanodactylus">
<emphasis id="B943EA8D496C2F1FFD11F89EFC13DD2B" box="[735,944,1828,1854]" italics="true" pageId="6" pageNumber="281">C. melanodactylus</emphasis>
</taxonomicName>
(
<figureCitation id="130C2A1A496C2F1FFC70F89EFBA1DD2A" box="[958,1026,1828,1855]" captionStart="FIGURE 5" captionStartId="8.[151,250,1868,1892]" captionTargetBox="[151,1436,181,1845]" captionTargetId="figure-27@8.[151,1436,181,1845]" captionTargetPageId="8" captionText="FIGURE 5. Diagnostic features of Cymo: A, F, K, P, C. mazu sp. nov.; B, G, L, Q, C. deplanatus; C, H, M, R, C. quadrilobatus; D, I, N, S, C. andreossyi; E, J, O, T, C. melanodactylus; A–E, overall morphology and live coloration; F–G, front; K–O, major cheliped; P–T, maxilliped 1. Scale bars: A, J, L, M = 2 mm; B–D, N, O = 5 mm; E = 10 mm; F, P = 0.5 mm; G–I, K, Q–T = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10146595" httpUri="https://zenodo.org/record/10146595/files/figure.png" pageId="6" pageNumber="281">Fig. 5</figureCitation>
, 
<tableCitation id="C6B50324496C2F1FFBC3F89EFBF0DD2B" box="[1037,1107,1828,1854]" captionStart="TABLE 3" captionStartId="7.[151,239,1124,1148]" captionTargetPageId="7" captionText="TABLE 3. Morphological comparison between five species of Cymo." httpUri="http://table.plazi.org/id/DF486617496D2F1EFF59FBDEFCDCDE6E" pageId="6" pageNumber="281" tableUuid="DF486617496D2F1EFF59FBDEFCDCDE6E">Tab. 3</tableCitation>
), and we provide updated the key for species within this genus, based on 
<bibRefCitation id="EFA64B6E496C2F1FFDB3F8F3FC99DD76" author="Ho, P. H. &amp; Ng, P. K. L." box="[637,826,1864,1891]" pageId="6" pageNumber="281" pagination="31 - 38" refId="ref6329" refString="Ho, P. H. &amp; Ng, P. K. L. (2005) On a new species of coral-symbiont crab of the genus Cymo de Haan, 1833 (Crustacea: Decapoda: Brachyura: Xanthidae), from the South China Sea. Zootaxa, 1029 (1), 31 - 38. https: // doi. org / 10.11646 / zootaxa. 1029.1.2" type="journal article" year="2005">Ho &amp; Ng (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>