279 lines
33 KiB
XML
279 lines
33 KiB
XML
<document id="120AE6F878E3E0777ED0957DC354DCC0" ID-DOI="10.5281/zenodo.183706" ID-GBIF-Dataset="e274d1d3-7bc2-46f8-a22e-e7d99d667298" ID-ISSN="11755326" ID-Zenodo-Dep="183706" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459929192289" checkinUser="plazi" docAuthor="Schwabe, Enrico & Engl, Winfried" docDate="2008" docId="F976879AFFD2E35EFF5EF99EFC1CEB6C" docLanguage="en" docName="zt01866p204.pdf" docOrigin="Zootaxa 1866" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Brookula exquisita Clarke 1961" docType="treatment" docVersion="11" lastPageNumber="193" masterDocId="054FFFE2FFD6E358FFC9FFA3FF89EC0A" masterDocTitle="Description of two new deepwater species of the genus Brookula Iredale, 1912 (Mollusca, Gastropoda, Trochoidea), with a revision of the genus for the Subantarctic and Arctic Sector of the Atlantic Ocean *" masterLastPageNumber="204" masterPageNumber="187" pageNumber="191" updateTime="1698229735954" updateUser="plazi">
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<mods:title id="DC7168363F50D7F43173E166D8A69D9B">Description of two new deepwater species of the genus Brookula Iredale, 1912 (Mollusca, Gastropoda, Trochoidea), with a revision of the genus for the Subantarctic and Arctic Sector of the Atlantic Ocean *</mods:title>
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<mods:namePart id="667AA1E38C6A0C070C05B1584135A738">Schwabe, Enrico</mods:namePart>
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<mods:namePart id="F65FB9BEA99295948B459BA5CDCDB999">Engl, Winfried</mods:namePart>
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<mods:typeOfResource id="DE02EDD06CD5EECE22B47D113A6F7D8C">text</mods:typeOfResource>
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<mods:title id="725CE1C4CA1BEA559F53BC2D0843CB4F">Zootaxa</mods:title>
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<mods:part id="27B7666829FBA7B4A16E46F06E3FF377">
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<mods:date id="3D3FC0149A592BC5823943FB51BE8731">2008</mods:date>
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<mods:number id="113A2E4804572C41DFD6A27DEFCD6538">1866</mods:number>
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<mods:identifier id="1DF920DFADE1D66542C77D81F90060E6" type="DOI">10.5281/zenodo.183706</mods:identifier>
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<mods:identifier id="41CE4039BBA67D42E010A816135EAADC" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="2EFC312F973A97E611D58B192A94548B" type="Zenodo-Dep">183706</mods:identifier>
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<treatment id="F976879AFFD2E35EFF5EF99EFC1CEB6C" ID-DOI="http://doi.org/10.5281/zenodo.5631749" ID-GBIF-Taxon="119374592" ID-Zenodo-Dep="5631749" LSID="urn:lsid:plazi:treatment:F976879AFFD2E35EFF5EF99EFC1CEB6C" httpUri="http://treatment.plazi.org/id/F976879AFFD2E35EFF5EF99EFC1CEB6C" lastPageId="6" lastPageNumber="193" pageId="4" pageNumber="191">
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<subSubSection id="39C56507FFD2E35CFF5EF99EFE98EA70" pageId="4" pageNumber="191" type="nomenclature">
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<paragraph id="7160368CFFD2E35CFF5EF99EFDAAEA5D" blockId="4.[151,547,1597,1658]" box="[151,547,1597,1623]" pageId="4" pageNumber="191">
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<heading id="2A2881E0FFD2E35CFF5EF99EFDAAEA5D" bold="true" box="[151,547,1597,1623]" fontSize="11" level="1" pageId="4" pageNumber="191" reason="1">
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<taxonomicName id="B6DF4D0FFFD2E35CFF5EF99EFDAAEA5D" ID-CoL="6936Q" authority="Clarke, 1961" authorityName="Clarke" authorityYear="1961" box="[151,547,1597,1623]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="4" pageNumber="191" phylum="Mollusca" rank="species" species="exquisita">
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<emphasis id="43ABEA9EFFD2E35CFF5EF99EFDAAEA5D" bold="true" box="[151,547,1597,1623]" pageId="4" pageNumber="191">
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<emphasis id="43ABEA9EFFD2E35CFF5EF99EFEF4EA5D" bold="true" box="[151,381,1597,1623]" italics="true" pageId="4" pageNumber="191">Brookula exquisita</emphasis>
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<bibRefCitation id="154E4B7DFFD2E35CFE4DF99EFDAAEA5D" author="Clarke" box="[388,547,1597,1623]" pageId="4" pageNumber="203" refString="Clarke, A. H. (1961) Abyssal mollusks from the South Atlantic Ocean. Bulletin of the Museum of Comparative Zoology, 125, 345 - 387." type="journal article" year="1961">Clarke, 1961</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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<paragraph id="7160368CFFD2E35CFF5EF9C3FE98EA70" blockId="4.[151,547,1597,1658]" box="[151,273,1632,1658]" pageId="4" pageNumber="191">
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(
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<figureCitation id="E9E42A09FFD2E35CFF56F9C3FF6EEA70" box="[159,231,1632,1658]" captionStart="FIGURE 3" captionStartId="5.[151,244,1588,1609]" captionTargetBox="[151,1435,597,1560]" captionTargetId="figure@5.[151,1436,597,1561]" captionTargetPageId="5" captionText="FIGURE 3: A – D, Brookula exquisita Clarke, 1961 (ZSM Mol 20070894) from Antarctica, South Sandwich Islands (58 ° 15.98 ’ S 24 ° 53.72 ’ W to 58 ° 16.13 ’ S 24 ° 53.87 ’ W), 2947 – 2970 m depth. A, Ventral view of the complete shell. B, View of the shell base showing the wide opening of the umbilicus. C, Apical view. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183708/files/figure.png" pageId="4" pageNumber="191">Figs 3</figureCitation>
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–4)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="39C56507FFD2E35CFF5EF913FBAAEACD" box="[151,1059,1712,1735]" pageId="4" pageNumber="191" type="reference_group">
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<paragraph id="7160368CFFD2E35CFF5EF913FBAAEACD" blockId="4.[151,1059,1712,1735]" box="[151,1059,1712,1735]" pageId="4" pageNumber="191">
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<treatmentCitationGroup id="51CF11A2FFD2E35CFF5EF913FBAAEACD" box="[151,1059,1712,1735]" pageId="4" pageNumber="191">
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<taxonomicName id="B6DF4D0FFFD2E35CFF5EF913FD6DEACD" ID-CoL="6936Q" authority="Clarke 1961: 356" authorityName="Clarke" authorityPageNumber="356" authorityYear="1961" box="[151,740,1712,1735]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="4" pageNumber="191" phylum="Mollusca" rank="species" species="exquisita" subGenus="Benthobrookula">
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<emphasis id="43ABEA9EFFD2E35CFF5EF913FDAAEACD" box="[151,547,1712,1735]" italics="true" pageId="4" pageNumber="191">Brookula (Benthobrookula) exquisita</emphasis>
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<treatmentCitation id="F07E109DFFD2E35CFDE0F913FD6DEACD" author="Clarke" box="[553,740,1712,1735]" page="356" pageId="4" pageNumber="191" year="1961">
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<bibRefCitation id="154E4B7DFFD2E35CFDE0F913FD39EACD" author="Clarke" box="[553,688,1712,1735]" pageId="4" pageNumber="203" refString="Clarke, A. H. (1961) Abyssal mollusks from the South Atlantic Ocean. Bulletin of the Museum of Comparative Zoology, 125, 345 - 387." type="journal article" year="1961">Clarke 1961</bibRefCitation>
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: 356
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</treatmentCitation>
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</taxonomicName>
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–357, pl. 3, fig. 8, pl. 4, fig. 2.
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="39C56507FFD2E35DFF5EF95BFCE1EDF8" lastPageId="5" lastPageNumber="192" pageId="4" pageNumber="191" type="materials_examined">
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<paragraph id="7160368CFFD2E35CFF5EF95BFF65EB30" blockId="4.[151,1436,1784,2010]" pageId="4" pageNumber="191">
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<typeStatus id="AE64882EFFD2E35CFF5EF95BFE80EB18" box="[151,265,1784,1810]" pageId="4" pageNumber="191" type="holotype">
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<emphasis id="43ABEA9EFFD2E35CFF5EF95BFE80EB18" bold="true" box="[151,265,1784,1810]" pageId="4" pageNumber="191">Holotype</emphasis>
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</typeStatus>
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(
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<collectionCode id="17CEAE49FFD2E35CFED2F95BFEE9EB18" LSID="urn:lsid:biocol.org:col:33791" box="[283,352,1784,1810]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="4" pageNumber="191">MCZ</collectionCode>
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: 225964), not examined, SEM images available in
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<bibRefCitation id="154E4B7DFFD2E35CFC01F95BFB4EEB18" box="[968,1223,1784,1810]" pageId="4" pageNumber="203" refString="Absalao, R. S., Miyaji, C. & Pimenta, A. D. (2001) The genus Brookula Iredale, 1912 (Gastropoda, Trochidae) from Brazil: description of a new species, with notes on other South American species. Zoosystema, 23 (4), 675 - 687." type="journal article">
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Absalão
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<emphasis id="43ABEA9EFFD2E35CFBFAF95AFBE0EB18" box="[1075,1129,1785,1810]" italics="true" pageId="4" pageNumber="191">et al</emphasis>
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. (2001)
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</bibRefCitation>
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and
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<bibRefCitation id="154E4B7DFFD2E35CFACCF95BFF60EB30" pageId="4" pageNumber="204" refString="Zelaya, D. G., Absalao, R. S. & Pimenta, A. D. (2006) A revision of Benthobrookula Clarke, 1961 (Gastropoda, Trochoidea) in the southwestern Atlantic Ocean. Journal of Molluscan Studies, 72 (1), 77 - 87." type="journal article">
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Zelaya
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<emphasis id="43ABEA9EFFD2E35CFA96F95AFA15EB18" box="[1375,1436,1785,1810]" italics="true" pageId="4" pageNumber="191">et al.</emphasis>
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(2006)
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</bibRefCitation>
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.
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</paragraph>
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<paragraph id="7160368CFFD2E35CFF0FF8EAFBBDEB68" blockId="4.[151,1436,1784,2010]" box="[198,1076,1864,1890]" pageId="4" pageNumber="191">
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||
<emphasis id="43ABEA9EFFD2E35CFF0FF8EAFED4EB68" box="[198,349,1865,1890]" italics="true" pageId="4" pageNumber="191">
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||
<typeStatus id="AE64882EFFD2E35CFF0FF8EAFF74EB68" box="[198,253,1865,1890]" pageId="4" pageNumber="191">Type</typeStatus>
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locality
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</emphasis>
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:
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<taxonomicName id="B6DF4D0FFFD2E35CFEA5F8EBFE60EB68" box="[364,489,1864,1890]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">
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<collectingCountry id="09C8761CFFD2E35CFEA5F8EBFE60EB68" box="[364,489,1864,1890]" name="Antarctica" pageId="4" pageNumber="191">Antarctica</collectingCountry>
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</taxonomicName>
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,
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<collectingCountry id="09C8761CFFD2E35CFE3DF8EBFD17EB68" box="[500,670,1864,1890]" name="South Georgia and the South Sandwich Islands" pageId="4" pageNumber="191">South Georgia</collectingCountry>
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||
(
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<geoCoordinate id="14EB504BFFD2E35CFD67F8EBFC86EB68" box="[686,783,1864,1890]" direction="south" orientation="latitude" pageId="4" pageNumber="191" precision="925" value="-55.483334">55°29’S</geoCoordinate>
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||
<geoCoordinate id="14EB504BFFD2E35CFCDEF8EBFC0CEB68" box="[791,901,1864,1890]" direction="west" orientation="longitude" pageId="4" pageNumber="191" precision="925" value="-37.95">37°57’W</geoCoordinate>
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||
),
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||
<quantity id="B6279B69FFD2E35CFC52F8EBFC7EEB68" box="[923,1015,1864,1890]" metricMagnitude="3" metricUnit="m" metricValue="3.758" pageId="4" pageNumber="191" unit="m" value="3758.0">3758 m</quantity>
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. OD.
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</paragraph>
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<paragraph id="7160368CFFD2E35DFF0FF8D2FCE1EDF8" blockId="4.[151,1436,1784,2010]" lastBlockId="5.[151,1436,152,498]" lastPageId="5" lastPageNumber="192" pageId="4" pageNumber="191">
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||
<emphasis id="43ABEA9EFFD2E35CFF0FF8D2FE3FEB80" box="[198,438,1905,1930]" italics="true" pageId="4" pageNumber="191">Material examined:</emphasis>
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Four specimens (
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<collectionCode id="17CEAE49FFD2E35CFD56F8D3FD54EB80" box="[671,733,1904,1930]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="4" pageNumber="191">ZSM</collectionCode>
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Mol 2007089220070894),
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<taxonomicName id="B6DF4D0FFFD2E35CFB8BF8D3FB36EB80" box="[1090,1215,1904,1930]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="4" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">
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<collectingCountry id="09C8761CFFD2E35CFB8BF8D3FB36EB80" box="[1090,1215,1904,1930]" name="Antarctica" pageId="4" pageNumber="191">Antarctica</collectingCountry>
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</taxonomicName>
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,
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<collectingCountry id="09C8761CFFD2E35CFB1AF8D3FF64EBB8" name="South Georgia and the South Sandwich Islands" pageId="4" pageNumber="191">South Sandwich Islands</collectingCountry>
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, St. PS 61/140–08 (
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<geoCoordinate id="14EB504BFFD2E35CFE11F83BFDD6EBB8" box="[472,607,1944,1970]" direction="south" orientation="latitude" pageId="4" pageNumber="191" precision="9" value="-58.266335">58°15.98’S</geoCoordinate>
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||
<geoCoordinate id="14EB504BFFD2E35CFDAEF83BFD7EEBB8" box="[615,759,1944,1970]" direction="west" orientation="longitude" pageId="4" pageNumber="191" precision="9" value="-24.895334">24°53.72’W</geoCoordinate>
|
||
to
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||
<geoCoordinate id="14EB504BFFD2E35CFCD6F83BFC2AEBB8" box="[799,931,1944,1970]" direction="south" orientation="latitude" pageId="4" pageNumber="191" precision="9" value="-58.268833">58°16.13’S</geoCoordinate>
|
||
<geoCoordinate id="14EB504BFFD2E35CFC65F83BFBC9EBB8" box="[940,1088,1944,1970]" direction="west" orientation="longitude" pageId="4" pageNumber="191" precision="9" value="-24.897833">24°53.87’W</geoCoordinate>
|
||
),
|
||
<quantity id="B6279B69FFD2E35CFB9FF83BFB75EBB8" box="[1110,1276,1944,1970]" metricMagnitude="3" metricUnit="m" metricValue="2.9585" metricValueMax="2.9699999999999998" metricValueMin="2.947" pageId="4" pageNumber="191" unit="m" value="2958.5" valueMax="2970.0" valueMin="2947.0">2947–2970 m</quantity>
|
||
, leg. Michael Schrödl,
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||
<date id="0561104CFFD2E35CFECAF863FE34EBD0" box="[259,445,1984,2010]" pageId="4" pageNumber="191" value="2002-03-22">March 22 2002</date>
|
||
, ANT XIX/4 (ANDEEP II) expedition. One specimen fixed and preserved in 75% ethanol (
|
||
<collectionCode id="17CEAE49FFD3E35DFECBFF3BFEC9ECB8" box="[258,320,152,178]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="192">ZSM</collectionCode>
|
||
Mol 20070892); remainder dry shells only including SEMmounted specimen (
|
||
<collectionCode id="17CEAE49FFD3E35DFAD7FF3BFAD5ECB8" box="[1310,1372,152,178]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="192">ZSM</collectionCode>
|
||
Mol 20070894). One specimen (
|
||
<collectionCode id="17CEAE49FFD3E35DFE20FF63FDAEECD0" box="[489,551,192,218]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="192">ZSM</collectionCode>
|
||
Mol
|
||
<date id="0561104CFFD3E35DFDB9FF63FD63ECD0" box="[624,746,192,218]" pageId="5" pageNumber="192" value="1582-02-20">20021582</date>
|
||
),
|
||
<taxonomicName id="B6DF4D0FFFD3E35DFCCBFF63FCF6ECD0" box="[770,895,192,218]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="5" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">
|
||
<collectingCountry id="09C8761CFFD3E35DFCCBFF63FCF6ECD0" box="[770,895,192,218]" name="Antarctica" pageId="5" pageNumber="192">Antarctica</collectingCountry>
|
||
</taxonomicName>
|
||
,
|
||
<collectingCountry id="09C8761CFFD3E35DFC59FF63FB3FECD0" box="[912,1206,192,218]" name="South Georgia and the South Sandwich Islands" pageId="5" pageNumber="192">South Sandwich Islands</collectingCountry>
|
||
, St. PS 61/140–07 (
|
||
<geoCoordinate id="14EB504BFFD3E35DFF69FF4BFEACED08" box="[160,293,232,258]" direction="south" orientation="latitude" pageId="5" pageNumber="192" precision="9" value="-58.266666">58°16.00’S</geoCoordinate>
|
||
<geoCoordinate id="14EB504BFFD3E35DFEE6FF4BFE36ED08" box="[303,447,232,258]" direction="west" orientation="longitude" pageId="5" pageNumber="192" precision="9" value="-24.896334">24°53.78’W</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="14EB504BFFD3E35DFE21FF4BFDE5ED08" box="[488,620,232,258]" direction="south" orientation="latitude" pageId="5" pageNumber="192" precision="9" value="-58.273167">58°16.39’S</geoCoordinate>
|
||
<geoCoordinate id="14EB504BFFD3E35DFDBFFF4BFC83ED08" box="[630,778,232,258]" direction="west" orientation="longitude" pageId="5" pageNumber="192" precision="9" value="-24.914167">24°54.85’W</geoCoordinate>
|
||
),
|
||
<quantity id="B6279B69FFD3E35DFCE9FF4BFC41ED08" box="[800,968,232,258]" metricMagnitude="3" metricUnit="m" metricValue="2.943" metricValueMax="2.945" metricValueMin="2.941" pageId="5" pageNumber="192" unit="m" value="2943.0" valueMax="2945.0" valueMin="2941.0">2941–2945 m</quantity>
|
||
, leg. Michael Schrödl,
|
||
<date id="0561104CFFD3E35DFB17FF4BFA11ED08" box="[1246,1432,232,258]" pageId="5" pageNumber="192" value="2002-03-21">March 21 2002</date>
|
||
, ANT XIX/4 (ANDEEP II) expedition, fixed and preserved in 96% ethanol. One dry specimen (
|
||
<collectionCode id="17CEAE49FFD3E35DFAEBFEB3FAE9ED20" box="[1314,1376,272,298]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="192">ZSM</collectionCode>
|
||
Mol 20070901),
|
||
<taxonomicName id="B6DF4D0FFFD3E35DFEE1FE9BFE2CED58" box="[296,421,312,338]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="5" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">
|
||
<collectingCountry id="09C8761CFFD3E35DFEE1FE9BFE2CED58" box="[296,421,312,338]" name="Antarctica" pageId="5" pageNumber="192">Antarctica</collectingCountry>
|
||
</taxonomicName>
|
||
, Northwest Weddell Sea, St. PS 61/131–03 (
|
||
<geoCoordinate id="14EB504BFFD3E35DFC00FE9BFBC6ED58" box="[969,1103,312,338]" direction="south" orientation="latitude" pageId="5" pageNumber="192" precision="9" value="-65.31983">65°19.19’S</geoCoordinate>
|
||
<geoCoordinate id="14EB504BFFD3E35DFB93FE9BFB63ED58" box="[1114,1258,312,338]" direction="west" orientation="longitude" pageId="5" pageNumber="192" precision="9" value="-51.54233">51°32.54’W</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="14EB504BFFD3E35DFADEFE9BFA12ED58" box="[1303,1435,312,338]" direction="south" orientation="latitude" pageId="5" pageNumber="192" precision="9" value="-65.33317">65°19.99’S</geoCoordinate>
|
||
<geoCoordinate id="14EB504BFFD3E35DFF5EFEC3FEA2ED70" box="[151,299,352,378]" direction="west" orientation="longitude" pageId="5" pageNumber="192" precision="9" value="-51.5205">51°31.23’W</geoCoordinate>
|
||
),
|
||
<quantity id="B6279B69FFD3E35DFEF6FEC3FE6DED70" box="[319,484,352,378]" metricMagnitude="3" metricUnit="m" metricValue="3.0524999999999998" metricValueMax="3.055" metricValueMin="3.05" pageId="5" pageNumber="192" unit="m" value="3052.5" valueMax="3055.0" valueMin="3050.0">3050–3055 m</quantity>
|
||
, leg. Michael Schrödl,
|
||
<date id="0561104CFFD3E35DFD39FEC3FC1FED70" box="[752,918,352,378]" pageId="5" pageNumber="192" value="2002-03-05">March 5 2002</date>
|
||
, ANT XIX/4 (ANDEEP II) expedition. Five specimens (
|
||
<collectionCode id="17CEAE49FFD3E35DFEEEFE2BFEECEDA8" box="[295,357,392,418]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="5" pageNumber="192">ZSM</collectionCode>
|
||
Mol
|
||
<date id="0561104CFFD3E35DFE78FE2BFDA3EDA8" box="[433,554,392,418]" pageId="5" pageNumber="192" value="1216-02-20">20021216</date>
|
||
;
|
||
<date id="0561104CFFD3E35DFDF5FE2BFD3CEDA8" box="[572,693,392,418]" pageId="5" pageNumber="192" value="1170-02-20">20021170</date>
|
||
),
|
||
<taxonomicName id="B6DF4D0FFFD3E35DFD06FE2BFCC5EDA8" box="[719,844,392,418]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="5" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">
|
||
<collectingCountry id="09C8761CFFD3E35DFD06FE2BFCC5EDA8" box="[719,844,392,418]" name="Antarctica" pageId="5" pageNumber="192">Antarctica</collectingCountry>
|
||
</taxonomicName>
|
||
, Drake Passage, St. PS 61
|
||
<emphasis id="43ABEA9EFFD3E35DFB56FE24FB2EEDAB" bold="true" box="[1183,1191,391,417]" pageId="5" pageNumber="192">/</emphasis>
|
||
041–03 (
|
||
<geoCoordinate id="14EB504BFFD3E35DFADFFE2BFA15EDA8" box="[1302,1436,392,418]" direction="south" orientation="latitude" pageId="5" pageNumber="192" precision="9" value="-59.370667">59°22.24’S</geoCoordinate>
|
||
<geoCoordinate id="14EB504BFFD3E35DFF5EFE13FEAEEDC0" box="[151,295,432,458]" direction="west" orientation="longitude" pageId="5" pageNumber="192" precision="9" value="-60.067665">60°04.06’W</geoCoordinate>
|
||
to
|
||
<geoCoordinate id="14EB504BFFD3E35DFE87FE13FE5AEDC0" box="[334,467,432,458]" direction="south" orientation="latitude" pageId="5" pageNumber="192" precision="9" value="-59.37583">59°22.55’S</geoCoordinate>
|
||
<geoCoordinate id="14EB504BFFD3E35DFE12FE13FDE6EDC0" box="[475,623,432,458]" direction="west" orientation="longitude" pageId="5" pageNumber="192" precision="9" value="-60.066833">60°04.01’W</geoCoordinate>
|
||
),
|
||
<quantity id="B6279B69FFD3E35DFD4CFE13FCA2EDC0" box="[645,811,432,458]" metricMagnitude="3" metricUnit="m" metricValue="2.367" metricValueMax="2.375" metricValueMin="2.359" pageId="5" pageNumber="192" unit="m" value="2367.0" valueMax="2375.0" valueMin="2359.0">2359–2375 m</quantity>
|
||
, leg. Katrin Linse,
|
||
<date id="0561104CFFD3E35DFBC7FE13FA88EDC0" box="[1038,1281,432,458]" pageId="5" pageNumber="192" value="2002-01-26" valueMax="2002-01-27" valueMin="2002-01-26">January 26–27 2002</date>
|
||
, ANT XIX/3 (ANDEEP I) expedition, fixed and preserved in 96% ethanol.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="25A06604FFD3E35DFF5EF997FEDEEAA3" httpUri="https://zenodo.org/record/183708/files/figure.png" pageId="5" pageNumber="192" targetBox="[151,1435,597,1560]" targetPageId="5">
|
||
<paragraph id="7160368CFFD3E35DFF5EF997FEDEEAA3" blockId="5.[151,1436,1588,1705]" pageId="5" pageNumber="192">
|
||
<emphasis id="43ABEA9EFFD3E35DFF5EF997FEC0EA43" bold="true" box="[151,329,1588,1609]" pageId="5" pageNumber="192">FIGURE 3: A–D</emphasis>
|
||
,
|
||
<taxonomicName id="B6DF4D0FFFD3E35DFE91F997FD10EA43" authority="Clarke, 1961" authorityName="Clarke" authorityYear="1961" box="[344,665,1588,1609]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="5" pageNumber="192" phylum="Mollusca" rank="species" species="exquisita">
|
||
<emphasis id="43ABEA9EFFD3E35DFE91F997FD98EA43" box="[344,529,1588,1609]" italics="true" pageId="5" pageNumber="192">Brookula exquisita</emphasis>
|
||
Clarke, 1961
|
||
</taxonomicName>
|
||
(ZSM Mol 20070894) from
|
||
<taxonomicName id="B6DF4D0FFFD3E35DFC77F997FBACEA43" box="[958,1061,1588,1609]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="5" pageNumber="201" phylum="Mollusca" rank="species" species="antarctica">Antarctica</taxonomicName>
|
||
, South Sandwich Islands (58°15.98’S 24°53.72’W to 58°16.13’S 24°53.87’W), 2947–2970 m depth.
|
||
<emphasis id="43ABEA9EFFD3E35DFD3FF9F7FC8EEA63" bold="true" box="[758,775,1620,1641]" pageId="5" pageNumber="192">A</emphasis>
|
||
, Ventral view of the complete shell.
|
||
<emphasis id="43ABEA9EFFD3E35DFBA3F9F7FBF3EA63" bold="true" box="[1130,1146,1620,1641]" pageId="5" pageNumber="192">B</emphasis>
|
||
, View of the shell base showing the wide opening of the umbilicus.
|
||
<emphasis id="43ABEA9EFFD3E35DFDC7F9D7FD96EA83" bold="true" box="[526,543,1652,1673]" pageId="5" pageNumber="192">C</emphasis>
|
||
, Apical view.
|
||
<emphasis id="43ABEA9EFFD3E35DFD62F9D7FD35EA83" bold="true" box="[683,700,1652,1673]" pageId="5" pageNumber="192">D</emphasis>
|
||
, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="39C56507FFD3E35EFF0FF8A7FAC0EDA8" lastPageId="6" lastPageNumber="193" pageId="5" pageNumber="192" type="description">
|
||
<paragraph id="7160368CFFD3E35DFF0FF8A7FC69EB14" blockId="5.[151,1439,1796,2022]" box="[198,992,1796,1822]" pageId="5" pageNumber="192">
|
||
<emphasis id="43ABEA9EFFD3E35DFF0FF8A7FD5CEB17" box="[198,725,1796,1821]" italics="true" pageId="5" pageNumber="192">Description of a specimen examined by SEM</emphasis>
|
||
(ZSM Mol 20070894)
|
||
</paragraph>
|
||
<paragraph id="7160368CFFD3E35EFF0FF88FFAC0EDA8" blockId="5.[151,1439,1796,2022]" lastBlockId="6.[151,1438,152,738]" lastPageId="6" lastPageNumber="193" pageId="5" pageNumber="192">
|
||
Shell minute,
|
||
<quantity id="B6279B69FFD3E35DFEBBF88FFE54EB4C" box="[370,477,1836,1862]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="5" pageNumber="192" unit="mm" value="1.11">1.11 mm</quantity>
|
||
long,
|
||
<quantity id="B6279B69FFD3E35DFDE6F88FFD12EB4C" box="[559,667,1836,1862]" metricMagnitude="-3" metricUnit="m" metricValue="1.67" pageId="5" pageNumber="192" unit="mm" value="1.67">1.67 mm</quantity>
|
||
wide (
|
||
<figureCitation id="E9E42A09FFD3E35DFD3AF88FFCDCEB4C" box="[755,853,1836,1862]" captionStart="FIGURE 3" captionStartId="5.[151,244,1588,1609]" captionTargetBox="[151,1435,597,1560]" captionTargetId="figure@5.[151,1436,597,1561]" captionTargetPageId="5" captionText="FIGURE 3: A – D, Brookula exquisita Clarke, 1961 (ZSM Mol 20070894) from Antarctica, South Sandwich Islands (58 ° 15.98 ’ S 24 ° 53.72 ’ W to 58 ° 16.13 ’ S 24 ° 53.87 ’ W), 2947 – 2970 m depth. A, Ventral view of the complete shell. B, View of the shell base showing the wide opening of the umbilicus. C, Apical view. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183708/files/figure.png" pageId="5" pageNumber="192">Fig. 3A</figureCitation>
|
||
), SL/SW ratio 0.66, depressed, fragile, white in color, with roundly inflated whorls, clearly expanded laterally, low spired, spire angle 126°. Protoconch (
|
||
<figureCitation id="E9E42A09FFD3E35DFAA4F8F7FF32EB9C" captionStart="FIGURE 3" captionStartId="5.[151,244,1588,1609]" captionTargetBox="[151,1435,597,1560]" captionTargetId="figure@5.[151,1436,597,1561]" captionTargetPageId="5" captionText="FIGURE 3: A – D, Brookula exquisita Clarke, 1961 (ZSM Mol 20070894) from Antarctica, South Sandwich Islands (58 ° 15.98 ’ S 24 ° 53.72 ’ W to 58 ° 16.13 ’ S 24 ° 53.87 ’ W), 2947 – 2970 m depth. A, Ventral view of the complete shell. B, View of the shell base showing the wide opening of the umbilicus. C, Apical view. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183708/files/figure.png" pageId="5" pageNumber="192">Fig. 3D</figureCitation>
|
||
) of one whorl, 320 µm in diameter, 116 µm long, globose, sculptured with fine anastomosing threads, forming reticulate pattern. Teleoconch (
|
||
<figureCitation id="E9E42A09FFD3E35DFDB9F807FD7EEBB4" box="[624,759,1956,1982]" captionStart="FIGURE 3" captionStartId="5.[151,244,1588,1609]" captionTargetBox="[151,1435,597,1560]" captionTargetId="figure@5.[151,1436,597,1561]" captionTargetPageId="5" captionText="FIGURE 3: A – D, Brookula exquisita Clarke, 1961 (ZSM Mol 20070894) from Antarctica, South Sandwich Islands (58 ° 15.98 ’ S 24 ° 53.72 ’ W to 58 ° 16.13 ’ S 24 ° 53.87 ’ W), 2947 – 2970 m depth. A, Ventral view of the complete shell. B, View of the shell base showing the wide opening of the umbilicus. C, Apical view. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183708/files/figure.png" pageId="5" pageNumber="192">Figs 3A, C</figureCitation>
|
||
) of up to 1.75 whorls with convex profile. Whorls connected only for a short distance, more or less at periphery of preceding one. Suture consequently deep and channeled. Last whorl clearly increasing in diameter and laterally expanded. Aperture circular,
|
||
<quantity id="B6279B69FFD0E35EFB31FF3BFAE8ECB8" box="[1272,1377,152,178]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="6" pageNumber="193" unit="mm" value="0.68">0.68 mm</quantity>
|
||
high (SL/AL ratio 1.63), peritreme holostomate, with thin lip. Shell base roundly inflated. Umbilicus (
|
||
<figureCitation id="E9E42A09FFD0E35EFAFEFF63FA1DECD0" box="[1335,1428,192,218]" captionStart="FIGURE 3" captionStartId="5.[151,244,1588,1609]" captionTargetBox="[151,1435,597,1560]" captionTargetId="figure@5.[151,1436,597,1561]" captionTargetPageId="5" captionText="FIGURE 3: A – D, Brookula exquisita Clarke, 1961 (ZSM Mol 20070894) from Antarctica, South Sandwich Islands (58 ° 15.98 ’ S 24 ° 53.72 ’ W to 58 ° 16.13 ’ S 24 ° 53.87 ’ W), 2947 – 2970 m depth. A, Ventral view of the complete shell. B, View of the shell base showing the wide opening of the umbilicus. C, Apical view. D, Detail of fig. C to show the fine protoconch ornamentation. Scale bars: 100 µm." httpUri="https://zenodo.org/record/183708/files/figure.png" pageId="6" pageNumber="193">Fig. 3B</figureCitation>
|
||
) phaneromphalous, circular, wide, surrounded by four slightly elevated spiral cords, two of them entering umbilicus. Shell surface cancellate. Axial sculpture of narrow ribs, 29 on last whorl in the
|
||
<typeStatus id="AE64882EFFD0E35EFB11FEB3FACBED20" box="[1240,1346,272,298]" pageId="6" pageNumber="193" type="holotype">holotype</typeStatus>
|
||
, deeply entering the umbilicus. Ribs regularly spaced, rounded, interspaces about four times rib width. Spiral sculpture of numerous flat threads, five on first teleoconch whorl, about 14 on the last adult whorl, distinctly narrower than axial ribs, partly overlaying axil ribs. Operculum circular, multispiral, with centric nucleus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39C56507FFD0E35EFF0FFE13FC1CEB6C" pageId="6" pageNumber="193" type="discussion">
|
||
<paragraph id="7160368CFFD0E35EFF0FFE13FB9EEDF8" blockId="6.[151,1438,152,738]" pageId="6" pageNumber="193">
|
||
<emphasis id="43ABEA9EFFD0E35EFF0FFE13FEA4EDC3" box="[198,301,432,457]" italics="true" pageId="6" pageNumber="193">Remarks</emphasis>
|
||
: This is the first record of this species from outside the
|
||
<typeStatus id="AE64882EFFD0E35EFC7DFE13FC6EEDC0" box="[948,999,432,458]" pageId="6" pageNumber="193">type</typeStatus>
|
||
locality and the new material considerably extends the geographic and bathymetric range of this species (Fig. 4).
|
||
</paragraph>
|
||
<paragraph id="7160368CFFD0E35EFF0FFDA3FEEEEEE8" blockId="6.[151,1438,152,738]" pageId="6" pageNumber="193">
|
||
There are minor differences to the description of the
|
||
<typeStatus id="AE64882EFFD0E35EFCE6FDA3FC1CEE10" box="[815,917,512,538]" pageId="6" pageNumber="193" type="holotype">holotype</typeStatus>
|
||
presented by
|
||
<bibRefCitation id="154E4B7DFFD0E35EFBF1FDA3FAAAEE10" box="[1080,1315,512,538]" pageId="6" pageNumber="204" refString="Zelaya, D. G., Absalao, R. S. & Pimenta, A. D. (2006) A revision of Benthobrookula Clarke, 1961 (Gastropoda, Trochoidea) in the southwestern Atlantic Ocean. Journal of Molluscan Studies, 72 (1), 77 - 87." type="journal article">
|
||
Zelaya
|
||
<emphasis id="43ABEA9EFFD0E35EFB46FDA3FB43EE13" box="[1167,1226,512,537]" italics="true" pageId="6" pageNumber="193">et al.</emphasis>
|
||
(2006)
|
||
</bibRefCitation>
|
||
. Although somewhat smaller (
|
||
<quantity id="B6279B69FFD0E35EFEB4FD8BFE6EEE48" box="[381,487,552,578]" metricMagnitude="-3" metricUnit="m" metricValue="1.11" pageId="6" pageNumber="193" unit="mm" value="1.11">1.11 mm</quantity>
|
||
vs
|
||
<quantity id="B6279B69FFD0E35EFDD8FD8BFDE7EE48" box="[529,622,552,578]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="193" unit="mm" value="1.5">1.5 mm</quantity>
|
||
) our specimen has a slightly larger protoconch (320 µm vs 300 µm). The spire angle in our specimen is greater (126° vs 105°) and although the teleoconch whorl number is smaller (1.75 vs 2.25), the number of axial ribs on the last body whorl is similar (29 vs 28). However, too little is known about the intraspecific variation of this species to warrant the description of a new species on these differences alone.
|
||
</paragraph>
|
||
<paragraph id="7160368CFFD0E35EFF5EF889FC1CEB6C" blockId="6.[151,1436,1834,1894]" pageId="6" pageNumber="193">
|
||
<emphasis id="43ABEA9EFFD0E35EFF5EF889FF7FEB35" bold="true" box="[151,246,1834,1855]" pageId="6" pageNumber="193">Figure 4:</emphasis>
|
||
The geographic distribution of
|
||
<taxonomicName id="B6DF4D0FFFD0E35EFDF9F888FCE6EB4A" authority="Clarke, 1961" authorityName="Clarke" authorityYear="1961" box="[560,879,1835,1856]" class="Gastropoda" family="Liotiidae" genus="Brookula" higherTaxonomySource="CoL" kingdom="Animalia" order="Seguenziida" pageId="6" pageNumber="193" phylum="Mollusca" rank="species" species="exquisita">
|
||
<emphasis id="43ABEA9EFFD0E35EFDF9F888FD6FEB4A" box="[560,742,1835,1856]" italics="true" pageId="6" pageNumber="193">Brookula exquisita</emphasis>
|
||
<bibRefCitation id="154E4B7DFFD0E35EFD27F888FCE6EB4A" author="Clarke" box="[750,879,1835,1856]" pageId="6" pageNumber="203" refString="Clarke, A. H. (1961) Abyssal mollusks from the South Atlantic Ocean. Bulletin of the Museum of Comparative Zoology, 125, 345 - 387." type="journal article" year="1961">Clarke, 1961</bibRefCitation>
|
||
</taxonomicName>
|
||
. The circle marks the literature data (
|
||
<typeStatus id="AE64882EFFD0E35EFB28F888FA85EB4A" box="[1249,1292,1835,1856]" pageId="6" pageNumber="193">type</typeStatus>
|
||
locality only), while the triangles show the new records. A
|
||
<quantity id="B6279B69FFD0E35EFD8BF8F2FD02EB6C" box="[578,651,1873,1894]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="6" pageNumber="193" unit="m" value="2500.0">2500 m</quantity>
|
||
depth contour in light grey.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |