329 lines
43 KiB
XML
329 lines
43 KiB
XML
<document id="9FA7BFE35CD5A89DC23BED9BC20CDD7B" ID-ISSN="11755334" ID-ZooBank="2412CB4F-4D29-4988-80C1-205D16767678" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629824082200" checkinUser="felipe" docAuthor="Borkent, Christopher J. & Wheeler, Terry A." docDate="2012" docId="CA3487C862769D0AEECCFBD1FD0CECE9" docLanguage="en" docName="zt03549p117.pdf" docOrigin="Zootaxa 3549" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leptomorphus couturieri Matile" docType="treatment" docVersion="3" lastPageNumber="25" masterDocId="360DFFB062619D12EE5BFF91FFD9EA01" masterDocTitle="Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549" masterLastPageNumber="117" masterPageNumber="1" pageNumber="24" updateTime="1699117448841" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="99DFC1196FBB3AA66E576596688BA7A4">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<mods:namePart id="03F7C1D172D5A1683DBA5B1B1392CBF6">Borkent, Christopher J.</mods:namePart>
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<mods:namePart id="B8F2645C453177E98555EDFF94ABB1E1">Wheeler, Terry A.</mods:namePart>
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<mods:date id="21E415C455623C86A09808BEC6E4E6AA">2012</mods:date>
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<treatment id="CA3487C862769D0AEECCFBD1FD0CECE9" ID-DOI="http://doi.org/10.5281/zenodo.5258333" ID-Zenodo-Dep="5258333" LSID="urn:lsid:plazi:treatment:CA3487C862769D0AEECCFBD1FD0CECE9" httpUri="http://treatment.plazi.org/id/CA3487C862769D0AEECCFBD1FD0CECE9" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<subSubSection id="0A87655562769D05EECCFBD1FDE5EE5B" box="[151,572,1088,1115]" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="422236DE62769D05EECCFBD1FDE5EE5B" blockId="23.[151,572,1088,1148]" box="[151,572,1088,1115]" pageId="23" pageNumber="24">
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<heading id="196A81B262769D05EECCFBD1FDE5EE5B" bold="true" box="[151,572,1088,1115]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="70E9EACC62769D05EECCFBD1FDE5EE5B" bold="true" box="[151,572,1088,1115]" pageId="23" pageNumber="24">
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8.
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<taxonomicName id="859D4D5D62769D05EEEFFBD0FDE5EE5B" ID-CoL="8CJ3H" authority="Matile" authorityName="Matile" box="[180,572,1088,1115]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="couturieri">
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<emphasis id="70E9EACC62769D05EEEFFBD0FE3AEE5A" bold="true" box="[180,483,1089,1115]" italics="true" pageId="23" pageNumber="24">Leptomorphus couturieri</emphasis>
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Matile
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562769D05EECCFBF5FDD7EE7D" box="[151,526,1124,1148]" pageId="23" pageNumber="24" type="description">
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<paragraph id="422236DE62769D05EECCFBF5FDD7EE7D" blockId="23.[151,572,1088,1148]" box="[151,526,1124,1148]" pageId="23" pageNumber="24">
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(
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<figureCitation id="DAA62A5B62769D05EEC5FBF5FED2EE7D" box="[158,267,1124,1148]" captionStart="FIGURES 5–9" captionStartId="84.[151,264,1764,1787]" captionTargetBox="[153,1435,183,1745]" captionTargetId="figure-49@84.[151,1436,181,1746]" captionTargetPageId="84" captionText="FIGURES 5–9. Leptomorphus habitus. 5. L. bifasciatus, neotype male. 6. L. brandiae, paratype male (in ethanol). 7. L. chaseni, holotype female. 8. L. carnevalei, male. 9. L. couturieri, male." pageId="23" pageNumber="24">Figures 9</figureCitation>
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,
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<figureCitation id="DAA62A5B62769D05EF4CFBF5FEEFEE7D" box="[279,310,1124,1148]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="23" pageNumber="24">49</figureCitation>
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,
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<figureCitation id="DAA62A5B62769D05EF1AFBF5FE86EE7D" box="[321,351,1124,1148]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="23" pageNumber="24">97</figureCitation>
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,
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<figureCitation id="DAA62A5B62769D05EF30FBF5FE4EEE7D" box="[363,407,1124,1148]" captionStart="FIGURE 134–136" captionStartId="107.[151,249,1814,1837]" captionTargetBox="[151,1436,193,1793]" captionTargetId="figure-66@107.[151,1436,193,1793]" captionTargetPageId="107" captionText="FIGURE 134–136. Distribution of Afrotropical Leptomorphus. 134. L. grjebinei (red circle), L. mandelai (blue square), L. stigmatus (black triangle). 135. L. aliciae (purple star), L. carnevalei (black square), L. couturieri (black circle), L. gracilis (red triangle). 136. L. crosskeyi (red triangle), L. medleri (blue star), L. obscurus (black circle)." pageId="23" pageNumber="24">135</figureCitation>
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,
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<figureCitation id="DAA62A5B62769D05EFF8FBF5FE16EE7D" box="[419,463,1124,1148]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values>50% are shown in bold below the branches." pageId="23" pageNumber="24">150</figureCitation>
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,
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<figureCitation id="DAA62A5B62769D05EF80FBF5FDDFEE7D" box="[475,518,1124,1148]" captionStart="FIGURE 153" captionStartId="115.[151,250,801,824]" captionTargetBox="[155,1364,186,781]" captionTargetId="figure-100@115.[151,1436,161,836]" captionTargetPageId="115" captionText="FIGURE 153. One of three equally parsimonious trees showing the relationships between species in the L. grjebinei species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="23" pageNumber="24">153</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="0A87655562769D05EECCFB3DFD26EEC0" box="[151,767,1195,1218]" pageId="23" pageNumber="24" type="reference_group">
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||
<paragraph id="422236DE62769D05EECCFB3DFD26EEC0" blockId="23.[151,767,1195,1218]" box="[151,767,1195,1218]" pageId="23" pageNumber="24">
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<treatmentCitationGroup id="628D11F062769D05EECCFB3DFD26EEC0" box="[151,767,1195,1218]" pageId="23" pageNumber="24">
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<treatmentCitation id="C33C10CF62769D05EECCFB3DFD22EEC0" author="Matile, L." box="[151,763,1195,1218]" page="144" pageId="23" pageNumber="24" year="1997">
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<taxonomicName id="859D4D5D62769D05EECCFB3DFD22EEC0" ID-CoL="8CJ3H" authority="Matile, 1997: 144" authorityName="Matile" authorityPageNumber="144" authorityYear="1997" box="[151,763,1195,1218]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="couturieri" subGenus="Gymnoscutum">
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<emphasis id="70E9EACC62769D05EECCFB3DFDE5EEC0" box="[151,572,1196,1218]" italics="true" pageId="23" pageNumber="24">Leptomorphus (Gymnoscutum) couturieri</emphasis>
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<bibRefCitation id="260C4B2F62769D05EC18FB3AFD22EEC0" author="Matile, L." box="[579,763,1195,1217]" pageId="23" pageNumber="24" pagination="143 - 150" refId="ref94722" refString="Matile, L. (1997) Nouvelles donnees sur les Leptomorphus Afrotropicaux (Diptera, Mycetophilidae). Revue Francaise d'Entomologie (Nouvelle Serie) 1996, 18 (4), 143 - 150." type="journal article" year="1997">Matile, 1997: 144</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="0A87655562769D05EECCFB7FFA43EC27" pageId="23" pageNumber="24" type="diagnosis">
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<paragraph id="422236DE62769D05EECCFB7FFD2EEF97" blockId="23.[151,1437,1262,2007]" pageId="23" pageNumber="24">
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||
<emphasis id="70E9EACC62769D05EECCFB7FFEEDEF06" bold="true" box="[151,308,1262,1287]" pageId="23" pageNumber="24">DIAGNOSIS</emphasis>
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: The only extant species of
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<taxonomicName id="859D4D5D62769D05ECEAFB7FFC8AEF07" authorityName=": Edwards" authorityYear="1925" box="[689,851,1262,1286]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus">
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<emphasis id="70E9EACC62769D05ECEAFB7FFC8AEF07" box="[689,851,1262,1286]" italics="true" pageId="23" pageNumber="24">Leptomorphus</emphasis>
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</taxonomicName>
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with the following combination of characters: laterotergite, anepisternum and anepimeron yellow; scutellum yellow; abdominal tergite 6 brown, tergite 7 yellow; male tergite 9 with basal 2/3 of lateral margins bulbous, tapering posteriorly, ventrally directed process at base of tergal evagination a single point, concavity of posterior margin of tergite 9 longer in diameter than width of sternite 9, tergal evagination tapering to blunt points (
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||
<figureCitation id="DAA62A5B62769D05ECCDFAEFFD32EF97" box="[662,747,1406,1430]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="23" pageNumber="24">Fig. 97</figureCitation>
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).
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</paragraph>
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<paragraph id="422236DE62769D05EE9CFA33FA43EC27" blockId="23.[151,1437,1262,2007]" pageId="23" pageNumber="24">
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This species can most easily be confused with other Afrotropical species with a completely brown tergite 6 (
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<emphasis id="70E9EACC62769D05EBDEFA32FE56EFDF" italics="true" pageId="23" pageNumber="24">
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<taxonomicName id="859D4D5D62769D05EBDEFA32FF3FEFDF" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="aliciae">L. aliciae</taxonomicName>
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,
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<taxonomicName id="859D4D5D62769D05EEA8FA56FE52EFDF" box="[243,395,1478,1502]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="carnevalei">L. carnevalei</taxonomicName>
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,
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</emphasis>
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<figureCitation id="DAA62A5B62769D05EFC3FA57FE06EFDF" box="[408,479,1478,1502]" captionStart="FIGURES 1–4" captionStartId="83.[151,263,1566,1589]" captionTargetBox="[153,1436,289,1543]" captionTargetId="figure-45@83.[151,1436,287,1544]" captionTargetPageId="83" captionText="FIGURES 1–4. Leptomorphus habitus. 1. L. aliciae, male. 2. L. babai, paratype male. 3. L. amorimi, paratype female. 4. L. bifasciatus, female." pageId="23" pageNumber="24">Figs 1</figureCitation>
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,
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<figureCitation id="DAA62A5B62769D05EFB4FA57FDC5EFDF" box="[495,540,1478,1502]" captionStart="FIGURES 5–9" captionStartId="84.[151,264,1764,1787]" captionTargetBox="[153,1435,183,1745]" captionTargetId="figure-49@84.[151,1436,181,1746]" captionTargetPageId="84" captionText="FIGURES 5–9. Leptomorphus habitus. 5. L. bifasciatus, neotype male. 6. L. brandiae, paratype male (in ethanol). 7. L. chaseni, holotype female. 8. L. carnevalei, male. 9. L. couturieri, male." pageId="23" pageNumber="24">8, 9</figureCitation>
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). It can be distinguished from these species based solely on the male genitalia (
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<figureCitation id="DAA62A5B62769D05EEC4FA7BFF2BEC03" box="[159,242,1514,1538]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="23" pageNumber="24">Fig. 97</figureCitation>
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) which are as long as wide, with sternite 9 not reaching margins of gonocoxite, and apex of the posterior lobes gradually tapering to a point, rather than having a step like constriction (
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<figureCitation id="DAA62A5B62769D05EA58F99FFB8DEC27" box="[1027,1108,1550,1574]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="23" pageNumber="24">Fig. 96</figureCitation>
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) or a bulbous apex (
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<figureCitation id="DAA62A5B62769D05EB62F99FFA55EC27" box="[1337,1420,1550,1574]" captionStart="FIGURES 91–94" captionStartId="97.[151,263,1788,1811]" captionTargetBox="[192,1391,186,1708]" captionTargetId="figure-96@97.[181,1403,176,1722]" captionTargetPageId="97" captionText="FIGURES 91–94. Leptomorphus male genitalia, ventral. 91. L. aliciae. 92. L. amorimi. 93. L. babai. 94. L. bifasciatus. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gc lb = gonocoxal lobe; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="23" pageNumber="24">Fig. 91</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562769D0AEE9CF9A3FE1BEF39" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
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||
<paragraph id="422236DE62769D0AEE9CF9A3FC2CEEF1" blockId="23.[151,1437,1262,2007]" lastBlockId="24.[151,1437,151,1768]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<emphasis id="70E9EACC62769D05EE9CF9A3FE5FEC4A" bold="true" box="[199,390,1586,1611]" pageId="23" pageNumber="24">DESCRIPTION</emphasis>
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:
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<emphasis id="70E9EACC62769D05EFCFF9A2FE14EC4A" box="[404,461,1587,1611]" italics="true" pageId="23" pageNumber="24">Male</emphasis>
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. (
|
||
<figureCitation id="DAA62A5B62769D05EFBAF9A3FDFBEC4B" box="[481,546,1586,1610]" captionStart="FIGURES 5–9" captionStartId="84.[151,264,1764,1787]" captionTargetBox="[153,1435,183,1745]" captionTargetId="figure-49@84.[151,1436,181,1746]" captionTargetPageId="84" captionText="FIGURES 5–9. Leptomorphus habitus. 5. L. bifasciatus, neotype male. 6. L. brandiae, paratype male (in ethanol). 7. L. chaseni, holotype female. 8. L. carnevalei, male. 9. L. couturieri, male." pageId="23" pageNumber="24">Fig. 9</figureCitation>
|
||
)
|
||
<emphasis id="70E9EACC62769D05EC6AF9A2FDA9EC4A" bold="true" box="[561,624,1587,1611]" pageId="23" pageNumber="24">Head</emphasis>
|
||
: yellow, circular in anterior view. Antenna dark brown; scape yellow, with brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae present; pedicel yellow, with 2–3 large bristles, several setae on apicodorsal margin, number of fine setae on apicoventral margin; flagellomere 1 with tapered base yellow remainder dark brown; flagellomere 6 1.5X as long as broad. Clypeus yellow, dorsoventrally elongate oval; bristles on clypeus light brown, 4–6 strong bristles on ventral margin directed ventrally, remaining setae directed medioventrally, clypeus 2X as long as face. Face yellow; shape a just longer than wide triangle, with few bristles ventrolaterally. Frons yellow; with few bristles medioventrally, frontal furrow running 1/2 distance from dorsal margin towards ventral margin, frontal cleft just anterior of median ocellus. Palpus with segments 1–4 yellow, segment 5 brown; segment 1 hidden behind eye, segments increasing in length, segment 5 2X length of segment 4 with central half thinner than base and apex, segment 3 with apicolateral patch of fine yellow setae encircled by strong dark setae. Labellum yellow. Eye with very few, short inter-ommatidial setulae scattered on surface. Occiput yellow with appressed, anteriorly directed setae. Ocelli in a straight line, space between ocelli less than diameter of laterals, lateral ocelli 2X their own diameter from eye margin, ocellar triangle dark brown/black with electric blue green specks.
|
||
<emphasis id="70E9EACC62799D0AEEB5FF71FE9CEAF9" bold="true" box="[238,325,224,248]" pageId="24" pageNumber="25">Thorax</emphasis>
|
||
: Length 1.33 ±
|
||
<quantity id="85659B3B62799D0AEFADFF4EFD80EAF6" box="[502,601,223,247]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="25" unit="mm" value="0.09">0.09 mm</quantity>
|
||
(
|
||
<quantity id="85659B3B62799D0AEC33FF4EFCD5EAF6" box="[616,780,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="1.33" metricValueMax="1.39" metricValueMin="1.27" pageId="24" pageNumber="25" unit="mm" value="1.33" valueMax="1.39" valueMin="1.27">1.27–1.39 mm</quantity>
|
||
, n = 8). Dark brown dorsally, yellow laterally. Scutum dark brown/black with blue-green specks, yellow anteromedial triangular spot pointing posteriorly and circle on each posterolateral corner; surface of scutum bare; acrostichal setae absent; dorsocentral setae present as fine setae for most of length; multiple rows of lateral setae present; patch of setae on scutum at wing base small. Scutellum yellow; with 6 large bristles and few small bristles. Prescutum yellow. Mediotergite yellow to light brown with 6 bristles on posterolateral corners, anteromedial patch of small bristles. Laterotergite yellow; anterior margin of laterotergite not reaching katepisternum. Anepimeron yellow. Anepisternum yellow. Katepisternum yellow. Antepronotum and proepisternum yellow. Margin of anterior and posterior spiracles yellow with yellow trichia. Metepisternum yellow. Anapleural suture with anterior portion slightly curved dorsally. Halter stem yellow, knob light brown.
|
||
<emphasis id="70E9EACC62799D0AEB06FD91FA4DE819" bold="true" box="[1373,1428,512,536]" pageId="24" pageNumber="25">Legs</emphasis>
|
||
: principally yellow; hind femur light brown at very apex; extreme anteroapical corner dark brown on all femora; hind tibia light brown. Midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia without comb of short setae along length of anteroventral surface, tibial spurs brown, foretibial spur length 2.5X apical thickness of foretibia, midtibia with faint, dorsal, bare patch of even thickness for 3/4 of its length, placed centrally, shortest midtibial spur 0.9X length of longest, longest midtibial spur 4X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 4.5X apical thickness of hind tibia. Foreleg first tarsomere 1.6X length of foretibia.
|
||
<emphasis id="70E9EACC62799D0AECF1FD6AFD33E915" bold="true" box="[682,746,763,788]" pageId="24" pageNumber="25">Wing</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62799D0AECA2FD6DFC94E915" box="[761,845,764,788]" captionStart="FIGURES 42–55" captionStartId="92.[151,263,1812,1835]" captionTargetBox="[151,1436,177,1803]" captionTargetId="figure-169@92.[151,1436,177,1803]" captionTargetPageId="92" captionText="FIGURES 42–55. Leptomorphus wings. 42. L. aliciae, dorsal. 43. L. amorimi paratype female, dorsal. 44. L. babai holotype male, ventral. 45. L. bifasciatus lectotype, ventral. 46. L. brandiae paratype female, dorsal. 47. L. carnevalei allotype, dorsal. 48. L. chaseni holotype female, dorsal. 49. L. couturieri, ventral. 50 L. crassipilus holotype male, ventral. 51. L. crosskeyi, ventral. 52. L. eberhardi paratype female, ventral. 53. L. fasciculatus paratype male, dorsal. 54. L. femoratus holotype male, ventral. 55. L. forcipatus dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1, M2, M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="24" pageNumber="25">Fig. 49</figureCitation>
|
||
): Length 5.8 ±
|
||
<quantity id="85659B3B62799D0AEDAEFD6DFB90E912" box="[1013,1097,764,788]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="24" pageNumber="25" unit="mm" value="0.4">0.4 mm</quantity>
|
||
(
|
||
<quantity id="85659B3B62799D0AEA02FD6DFB38E912" box="[1113,1249,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="6.2" metricValueMin="5.5" pageId="24" pageNumber="25" unit="mm" value="5.85" valueMax="6.2" valueMin="5.5">5.5–6.2 mm</quantity>
|
||
, n = 7). Hyaline; apical macula light, beginning at apex of R
|
||
<subScript id="DE19349B62799D0AEC2AFCBEFDA0E93C" attach="left" box="[625,633,815,829]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
and M
|
||
<subScript id="DE19349B62799D0AEC9CFCBEFD16E93C" attach="left" box="[711,719,815,829]" fontSize="6" pageId="24" pageNumber="25">4</subScript>
|
||
and reaching wing tip; medial macula absent. Macrotrichia in all cells, though absent from posterobasal margin of cell a. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with group of macrotrichia. Vein sc-r present, apical end joining R at 2–3X its own length prior to origin of Rs. R
|
||
<subScript id="DE19349B62799D0AEF85FC0FFE3FE9AD" attach="left" box="[478,486,926,940]" fontSize="6" pageId="24" pageNumber="25">4</subScript>
|
||
absent. R
|
||
<subScript id="DE19349B62799D0AEC0BFC0FFD81E9AD" attach="left" box="[592,600,926,940]" fontSize="6" pageId="24" pageNumber="25">5</subScript>
|
||
straight, slight posterior turn near tip. M
|
||
<subScript id="DE19349B62799D0AEA4DFC0FFBC7E9AD" attach="left" box="[1046,1054,926,940]" fontSize="6" pageId="24" pageNumber="25">1</subScript>
|
||
reaching apex before R
|
||
<subScript id="DE19349B62799D0AEB7FFC0FFAF5E9AD" attach="left" box="[1316,1324,926,940]" fontSize="6" pageId="24" pageNumber="25">5</subScript>
|
||
, apices of M veins fading before wing margin. M
|
||
<subScript id="DE19349B62799D0AEC03FC54FDB9E9D2" attach="left" box="[600,608,965,979]" fontSize="6" pageId="24" pageNumber="25">4</subScript>
|
||
-CuA fork arising apically of origin of r-m. A
|
||
<subScript id="DE19349B62799D0AEA29FC54FBA3E9D2" attach="left" box="[1138,1146,965,979]" fontSize="6" pageId="24" pageNumber="25">2</subScript>
|
||
faintly present as crease.
|
||
<emphasis id="70E9EACC62799D0AEECCFC4DFED3E9F4" bold="true" box="[151,266,988,1013]" pageId="24" pageNumber="25">Abdomen</emphasis>
|
||
: Tergites principally yellow. T3–5 with posterior 1/4 dark brown, T6 dark brown. Tergite 8 smaller than all other abdominal sclerites, with ~8 bristles on each apicolateral corner.
|
||
<emphasis id="70E9EACC62799D0AED87FB91FB91EE18" bold="true" box="[988,1096,1024,1049]" pageId="24" pageNumber="25">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62799D0AEA06FB91FB6CEE19" box="[1117,1205,1024,1048]" captionStart="FIGURES 95–98" captionStartId="98.[151,263,1768,1791]" captionTargetBox="[196,1387,188,1694]" captionTargetId="figure-108@98.[188,1396,176,1702]" captionTargetPageId="98" captionText="FIGURES 95–98. Leptomorphus male genitalia, ventral. 95. L. brandiae. 96. L. carnevalei. 97. L. couturieri.98. L. crassipilus, arrow indicates dense rows of thick blunt bristles, arrangement of components reconstituted from crushed genitalia of holotype. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="24" pageNumber="25">Fig. 97</figureCitation>
|
||
): yellow. Sternite 9 sclerotized oval, 1/3 the width of genitalia at widest point, just reaching medial margin of gonoxite. Tergite 9 as wide as long, rounded margins tapering into two pointed lobes with medial U-shaped indentation, a ventrally extending thin point at base of posterior lobe. Gonocoxite placed basally on T9, medial margin not reaching medial line, bearing gonostylus on apical 1/3. Gonostylus swelling from base into a triangular lobe with apex pointed, gonocoxite III fused to dorsolateral margin. Aedeagus 4/5 length of gonocoxite, tapering towards apex (slight central swelling), apodemes 1/3 total length. Parameres a simple taper, apodemes ~4/5 length of parameres.
|
||
</paragraph>
|
||
<paragraph id="422236DE62799D0AEE9CFB6DFE47EF15" blockId="24.[151,1437,151,1768]" box="[199,414,1276,1300]" pageId="24" pageNumber="25">
|
||
<emphasis id="70E9EACC62799D0AEE9CFB6DFEC2EF15" box="[199,283,1276,1300]" italics="true" pageId="24" pageNumber="25">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph id="422236DE62799D0AEE9CFAB0FE1BEF39" blockId="24.[151,1437,151,1768]" box="[199,450,1312,1336]" pageId="24" pageNumber="25">
|
||
<emphasis id="70E9EACC62799D0AEE9CFAB0FEE7EF39" box="[199,318,1313,1336]" italics="true" pageId="24" pageNumber="25">Immatures</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562799D0AEE9CFAD5FE16EF5D" box="[199,463,1348,1373]" pageId="24" pageNumber="25" type="biology_ecology">
|
||
<paragraph id="422236DE62799D0AEE9CFAD5FE16EF5D" blockId="24.[151,1437,151,1768]" box="[199,463,1348,1373]" pageId="24" pageNumber="25">
|
||
<emphasis id="70E9EACC62799D0AEE9CFAD5FE93EF5C" bold="true" box="[199,330,1348,1373]" pageId="24" pageNumber="25">BIOLOGY</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562799D0AEE9CFAF9FC00EF81" box="[199,985,1384,1409]" pageId="24" pageNumber="25" type="distribution">
|
||
<paragraph id="422236DE62799D0AEE9CFAF9FC00EF81" blockId="24.[151,1437,151,1768]" box="[199,985,1384,1409]" pageId="24" pageNumber="25">
|
||
<emphasis id="70E9EACC62799D0AEE9CFAF9FE4FEF80" bold="true" box="[199,406,1384,1409]" pageId="24" pageNumber="25">DISTRIBUTION</emphasis>
|
||
:
|
||
<collectingCountry id="3A8A764E62799D0AEFFFFAF8FDF1EF81" box="[420,552,1384,1409]" name="Ivory Coast" pageId="24" pageNumber="25">Ivory Coast</collectingCountry>
|
||
and
|
||
<collectingCountry id="3A8A764E62799D0AEC05FAF9FD73EF81" box="[606,682,1384,1408]" name="Ghana" pageId="24" pageNumber="25">Ghana</collectingCountry>
|
||
, (
|
||
<figureCitation id="DAA62A5B62799D0AECE5FAF9FCC6EF81" box="[702,799,1384,1408]" captionStart="FIGURE 134–136" captionStartId="107.[151,249,1814,1837]" captionTargetBox="[151,1436,193,1793]" captionTargetId="figure-66@107.[151,1436,193,1793]" captionTargetPageId="107" captionText="FIGURE 134–136. Distribution of Afrotropical Leptomorphus. 134. L. grjebinei (red circle), L. mandelai (blue square), L. stigmatus (black triangle). 135. L. aliciae (purple star), L. carnevalei (black square), L. couturieri (black circle), L. gracilis (red triangle). 136. L. crosskeyi (red triangle), L. medleri (blue star), L. obscurus (black circle)." pageId="24" pageNumber="25">Fig. 135</figureCitation>
|
||
),
|
||
<quantity id="85659B3B62799D0AED68FAF9FC0DEF81" box="[819,980,1384,1408]" metricMagnitude="2" metricUnit="m" metricValue="1.6" metricValueMax="2.0" metricValueMin="1.2" pageId="24" pageNumber="25" unit="m" value="160.0" valueMax="200.0" valueMin="120.0">120–200 masl</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562799D0AEE9CFA1DFCE4EFED" pageId="24" pageNumber="25" type="discussion">
|
||
<paragraph id="422236DE62799D0AEE9CFA1DFCE4EFED" blockId="24.[151,1437,151,1768]" pageId="24" pageNumber="25">
|
||
<emphasis id="70E9EACC62799D0AEE9CFA1DFEABEFA4" bold="true" box="[199,370,1420,1445]" pageId="24" pageNumber="25">DISCUSSION</emphasis>
|
||
: As discussed below in the phylogeny section, the placement of
|
||
<taxonomicName id="859D4D5D62799D0AEA27FA1DFAA3EFA5" box="[1148,1402,1420,1444]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="70E9EACC62799D0AEA27FA1DFAC7EFA5" box="[1148,1310,1420,1444]" italics="true" pageId="24" pageNumber="25">Leptomorphus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in subgenera (
|
||
<bibRefCitation id="260C4B2F62799D0AEF43FA21FE7DEFC9" author="Matile, L." box="[280,420,1456,1480]" pageId="24" pageNumber="25" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977</bibRefCitation>
|
||
) is not supported by our phylogenetic results. This species is therefore removed from the subgenus
|
||
<taxonomicName id="859D4D5D62799D0AEF5DFA45FE7FEFED" box="[262,422,1492,1516]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subGenus" subGenus="Gymnoscutum">
|
||
<emphasis id="70E9EACC62799D0AEF5DFA45FE7FEFED" box="[262,422,1492,1516]" italics="true" pageId="24" pageNumber="25">Gymnoscutum</emphasis>
|
||
</taxonomicName>
|
||
and placed solely in
|
||
<emphasis id="70E9EACC62799D0AECC8FA45FCE4EFED" box="[659,829,1492,1516]" italics="true" pageId="24" pageNumber="25">
|
||
<taxonomicName id="859D4D5D62799D0AECC8FA45FCE0EFED" authorityName=": Edwards" authorityYear="1925" box="[659,825,1492,1516]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562799D0AEE9CFA69FD0CECE9" pageId="24" pageNumber="25" type="materials_examined">
|
||
<paragraph id="422236DE62799D0AEE9CFA69FA9DEC59" blockId="24.[151,1437,151,1768]" pageId="24" pageNumber="25">
|
||
<materialsCitation id="F2F53C8362799D0AEE9CFA69FA9DEC59" collectionCode="MATERIAL, FORET, DENSE" determinerName="L. Matile" location="MNHN" pageId="24" pageNumber="25" specimenCount="3" specimenCount-male="1" stateProvince="Monaghan" typeStatus="holotype">
|
||
<emphasis id="70E9EACC62799D0AEE9CFA69FDDBEC10" bold="true" box="[199,514,1528,1553]" pageId="24" pageNumber="25">
|
||
<collectionCode id="248CAE1B62799D0AEE9CFA69FE86EC10" box="[199,351,1528,1553]" pageId="24" pageNumber="25">MATERIAL</collectionCode>
|
||
EXAMINED
|
||
</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC62799D0AEC4BFA69FD59EC11" box="[528,640,1528,1552]" italics="true" pageId="24" pageNumber="25">
|
||
<typeStatus id="9D26887C62799D0AEC4BFA69FDA3EC11" box="[528,634,1528,1552]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
adult male, pinned on double mount minuten, genitalia in glass vial on pin, labelled “COTE-D’IVOIRE/ TAI/ 9-II-[19]85/ G. COUTURIER leg.;
|
||
<collectionCode id="248CAE1B62799D0AEDBDF98DFB78EC35" box="[998,1185,1564,1588]" pageId="24" pageNumber="25">FORET DENSE</collectionCode>
|
||
/ SEMPERVIRENTE; [Red label]
|
||
<typeStatus id="9D26887C62799D0AEF40F9D1FE6BEC59" box="[283,434,1600,1624]" pageId="24" pageNumber="25" type="holotype">HOLOTYPE</typeStatus>
|
||
;
|
||
<taxonomicName id="859D4D5D62799D0AEFE6F9D1FDB9EC59" authorityName=": Edwards" authorityYear="1925" box="[445,608,1600,1624]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
|
||
(g.)/ couturieri
|
||
<taxonomicNameLabel id="6BDA57B762799D0AED49F9D0FC95EC59" box="[786,844,1600,1624]" pageId="24" pageNumber="25" rank="species">n. sp.</taxonomicNameLabel>
|
||
/
|
||
<specimenCount id="549BFD5762799D0AED06F9D0FCABEC59" box="[861,882,1601,1624]" pageId="24" pageNumber="25" type="male">♂</specimenCount>
|
||
<typeStatus id="9D26887C62799D0AED22F9D1FC03EC59" box="[889,986,1600,1624]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
/
|
||
<determinerName id="B7FEDA1A62799D0AEDB1F9D0FB8BEC59" box="[1002,1106,1600,1624]" pageId="24" pageNumber="25">L. Matile</determinerName>
|
||
det. 1995” [
|
||
<collectingRegion id="8059F83C62799D0AEA80F9D0FAE3EC59" box="[1243,1338,1601,1624]" country="Ireland" name="Monaghan" pageId="24" pageNumber="25">MNHN</collectingRegion>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="422236DE62799D0AEE9CF9F4FC7DEC7D" blockId="24.[151,1437,151,1768]" box="[199,932,1636,1660]" pageId="24" pageNumber="25">
|
||
<materialsCitation id="F2F53C8362799D0AEE9CF9F4FC45EC7D" box="[199,924,1636,1660]" collectingDate="1980-05-06" location="MNHN" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1" stateProvince="Monaghan" typeStatus="paratype">
|
||
<emphasis id="70E9EACC62799D0AEE9CF9F4FEEFEC7D" box="[199,310,1637,1660]" italics="true" pageId="24" pageNumber="25">
|
||
<typeStatus id="9D26887C62799D0AEE9CF9F4FEE8EC7D" box="[199,305,1637,1660]" pageId="24" pageNumber="25" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
labelled as for
|
||
<typeStatus id="9D26887C62799D0AEFB8F9F5FD9CEC7D" box="[483,581,1636,1660]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
|
||
except
|
||
<date id="3623101E62799D0AECC7F9F5FD25EC7D" box="[668,764,1636,1660]" pageId="24" pageNumber="25" value="1980-05-06">
|
||
<collectingDate id="2667E9F662799D0AECC7F9F5FD25EC7D" box="[668,764,1636,1660]" pageId="24" pageNumber="25" value="1980-05-06">6.v.1980</collectingDate>
|
||
</date>
|
||
(
|
||
<specimenCount id="549BFD5762799D0AED55F9F5FCE9EC7D" box="[782,816,1636,1660]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
|
||
,
|
||
<collectingRegion id="8059F83C62799D0AED65F9F5FC4FEC7D" box="[830,918,1636,1660]" country="Ireland" name="Monaghan" pageId="24" pageNumber="25">MNHN</collectingRegion>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="422236DE62799D0AEE9CF919FD0CECE9" blockId="24.[151,1437,151,1768]" pageId="24" pageNumber="25">
|
||
<emphasis id="70E9EACC62799D0AEE9CF919FEA6ECA1" box="[199,383,1672,1696]" italics="true" pageId="24" pageNumber="25">Other material:</emphasis>
|
||
<materialsCitation id="F2F53C8362799D0AEFD5F919FA41ECA1" box="[398,1432,1672,1697]" collectingDate="1993-12-09" collectionCode="T" collectorName="J. Kjaerandsen" country="Ghana" county="Reserve" location="Western Region" municipality="Ankasa" pageId="24" pageNumber="25" specimenCount="1">
|
||
<collectingCountry id="3A8A764E62799D0AEFD5F919FE2FECA1" box="[398,502,1672,1696]" name="Ghana" pageId="24" pageNumber="25">GHANA</collectingCountry>
|
||
,
|
||
<location id="4742600562799D0AEC52F918FD1EECA1" LSID="urn:lsid:plazi:treatment:CA3487C862769D0AEECCFBD1FD0CECE9:4742600562799D0AEC52F918FD1EECA1" box="[521,711,1672,1696]" country="Ghana" county="Reserve" municipality="Ankasa" name="Western Region" pageId="24" pageNumber="25">Western Region</location>
|
||
,
|
||
<collectingMunicipality id="A246ACA462799D0AEC81F919FCF7ECA1" box="[730,814,1672,1696]" pageId="24" pageNumber="25">Ankasa</collectingMunicipality>
|
||
game prod.
|
||
<collectingCounty id="AB434E5262799D0AED8AF918FBF7ECA1" box="[977,1070,1673,1696]" pageId="24" pageNumber="25">Reserve</collectingCounty>
|
||
,
|
||
<date id="3623101E62799D0AEA1AF919FB6CECA1" box="[1089,1205,1672,1696]" pageId="24" pageNumber="25" value="1993-12-09">
|
||
<collectingDate id="2667E9F662799D0AEA1AF919FB6CECA1" box="[1089,1205,1672,1696]" pageId="24" pageNumber="25" value="1993-12-09">9.xii.1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="EF68530862799D0AEA92F918FAADECA1" box="[1225,1396,1672,1696]" pageId="24" pageNumber="25">J. Kjaerandsen</collectorName>
|
||
;
|
||
<collectionCode id="248CAE1B62799D0AEBDDF918FA41ECA1" box="[1414,1432,1673,1696]" name="Tavera, Department of Geology and Geophysics" pageId="24" pageNumber="25">T</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="F2F53C8362799D0AEECCF93DFAEBECC5" box="[151,1330,1708,1732]" collectingDate="1994-10-31" collectingDateMax="1994-11-15" collectingDateMin="1994-10-31" collectionCode="ZMUN" country="Ghana" pageId="24" pageNumber="25" specimenCount="6" specimenCount-male="6">
|
||
Andersen. (
|
||
<specimenCount id="549BFD5762799D0AEF7FF93DFE9CECC5" box="[292,325,1708,1732]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="248CAE1B62799D0AEF00F93DFE6BECC5" box="[347,434,1708,1732]" pageId="24" pageNumber="25">ZMUN</collectionCode>
|
||
); same except
|
||
<date id="3623101E62799D0AEC2BF93DFCDEECC5" box="[624,775,1708,1732]" pageId="24" pageNumber="25" value="1994-11-08" valueMax="1994-11-15" valueMin="1994-11-08">
|
||
<collectingDate id="2667E9F662799D0AEC2BF93DFCDEECC5" box="[624,775,1708,1732]" pageId="24" pageNumber="25" value="1994-11-08" valueMax="1994-11-15" valueMin="1994-11-08">8–15.xi.1994</collectingDate>
|
||
</date>
|
||
. (
|
||
<specimenCount id="549BFD5762799D0AED79F93DFC9FECC5" box="[802,838,1708,1732]" pageId="24" pageNumber="25" type="male">2♂</specimenCount>
|
||
,
|
||
<collectionCode id="248CAE1B62799D0AED06F93DFC6CECC5" box="[861,949,1708,1732]" pageId="24" pageNumber="25">ZMUN</collectionCode>
|
||
);
|
||
<date id="3623101E62799D0AED8BF93DFBA2ECC5" box="[976,1147,1708,1732]" pageId="24" pageNumber="25" value="1994-10-31" valueMax="1994-11-08" valueMin="1994-10-31">
|
||
<collectingDate id="2667E9F662799D0AED8BF93DFBA2ECC5" box="[976,1147,1708,1732]" pageId="24" pageNumber="25" value="1994-10-31" valueMax="1994-11-08" valueMin="1994-10-31">31.x–8.xi.1994</collectingDate>
|
||
</date>
|
||
. (
|
||
<specimenCount id="549BFD5762799D0AEAC3F93DFB62ECC5" box="[1176,1211,1708,1732]" pageId="24" pageNumber="25" type="male">3♂</specimenCount>
|
||
,
|
||
<collectionCode id="248CAE1B62799D0AEA89F93DFAF3ECC5" box="[1234,1322,1708,1732]" pageId="24" pageNumber="25">ZMUN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="F2F53C8362799D0AEB1EF93DFD0BECE9" collectingDate="1980-05-06" collectionCode="MNHN" collectorName="G. Couturier." country="Ivory Coast" location="Tay" pageId="24" pageNumber="25" specimenCount="1" specimenCount-male="1">
|
||
<collectingCountry id="3A8A764E62799D0AEB1EF93DFF2DECE9" name="Ivory Coast" pageId="24" pageNumber="25">IVORY COAST</collectingCountry>
|
||
,
|
||
<location id="4742600562799D0AEEA5F940FEFBECE9" LSID="urn:lsid:plazi:treatment:CA3487C862769D0AEECCFBD1FD0CECE9:4742600562799D0AEEA5F940FEFBECE9" box="[254,290,1744,1768]" country="Ivory Coast" name="Tay" pageId="24" pageNumber="25">Taї</location>
|
||
,
|
||
<date id="3623101E62799D0AEF6BF941FE4AECE9" box="[304,403,1744,1768]" pageId="24" pageNumber="25" value="1980-05-06">
|
||
<collectingDate id="2667E9F662799D0AEF6BF941FE4AECE9" box="[304,403,1744,1768]" pageId="24" pageNumber="25" value="1980-05-06">6.v.1980</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="EF68530862799D0AEFC5F941FDF7ECE9" box="[414,558,1744,1768]" pageId="24" pageNumber="25">G. Couturier.</collectorName>
|
||
(
|
||
<specimenCount id="549BFD5762799D0AEC1BF941FDB8ECE9" box="[576,609,1744,1768]" pageId="24" pageNumber="25" type="male">1♂</specimenCount>
|
||
,
|
||
<collectionCode id="248CAE1B62799D0AEC34F941FD13ECE9" box="[623,714,1744,1768]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="25">MNHN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |