297 lines
38 KiB
XML
297 lines
38 KiB
XML
<document id="344668555B5D60FCEA23D2A780FB4983" ID-ISSN="11755334" ID-ZooBank="2412CB4F-4D29-4988-80C1-205D16767678" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629824082200" checkinUser="felipe" docAuthor="Borkent, Christopher J. & Wheeler, Terry A." docDate="2012" docId="CA3487C862449D34EECCFF06FE50E8CD" docLanguage="en" docName="zt03549p117.pdf" docOrigin="Zootaxa 3549" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Leptomorphus grjebinei Matile" docType="treatment" docVersion="3" lastPageNumber="39" masterDocId="360DFFB062619D12EE5BFF91FFD9EA01" masterDocTitle="Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549" masterLastPageNumber="117" masterPageNumber="1" pageNumber="38" updateTime="1699117448841" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="ED1DA870B2DD8E6D89D8F9AC5E58E5C4">Systematics and Phylogeny of Leptomorphus Curtis (Diptera: Mycetophilidae) 3549</mods:title>
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<mods:namePart id="3FEF5597D7DB185C47AB5364DB8F5D01">Borkent, Christopher J.</mods:namePart>
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<mods:namePart id="0684B28F52E03D15F008C852FAD97376">Wheeler, Terry A.</mods:namePart>
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<mods:title id="CF26A87E78A648A837AC29404E9C1026">Zootaxa</mods:title>
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<mods:date id="39991669975F7D661FD96AC126536118">2012</mods:date>
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<mods:number id="AB5DA25B5B4C6C60E891742BBD02E314">3549</mods:number>
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<treatment id="CA3487C862449D34EECCFF06FE50E8CD" ID-DOI="http://doi.org/10.5281/zenodo.5258345" ID-Zenodo-Dep="5258345" LSID="urn:lsid:plazi:treatment:CA3487C862449D34EECCFF06FE50E8CD" httpUri="http://treatment.plazi.org/id/CA3487C862449D34EECCFF06FE50E8CD" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
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<paragraph id="422236DE62449D37EECCFF06FDE4EAB0" blockId="37.[151,573,151,210]" box="[151,573,151,177]" pageId="37" pageNumber="38">
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<emphasis id="70E9EACC62449D37EECCFF06FDE4EAB0" bold="true" box="[151,573,151,177]" pageId="37" pageNumber="38">
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17.
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<taxonomicName id="859D4D5D62449D37EE98FF06FDE4EAB0" ID-CoL="8CJ3D" authority="Matile" authorityName="Matile" box="[195,573,151,177]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="grjebinei">
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<emphasis id="70E9EACC62449D37EE98FF06FE3CEAB0" bold="true" box="[195,485,151,177]" italics="true" pageId="37" pageNumber="38">Leptomorphus grjebinei</emphasis>
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Matile
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562449D37EECCFF2BFDF3EAD3" box="[151,554,186,210]" pageId="37" pageNumber="38" type="description">
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<paragraph id="422236DE62449D37EECCFF2BFDF3EAD3" blockId="37.[151,573,151,210]" box="[151,554,186,210]" pageId="37" pageNumber="38">
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(
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<figureCitation id="DAA62A5B62449D37EEC5FF2BFEC0EAD3" box="[158,281,186,210]" captionStart="FIGURES 16–19" captionStartId="86.[151,263,1697,1720]" captionTargetBox="[153,1435,194,1675]" captionTargetId="figure-40@86.[151,1436,193,1676]" captionTargetPageId="86" captionText="FIGURES 16–19. Leptomorphus habitus.16. L. forcipatus, male. 17. L. furcatus, paratype male. 18. L. gracilis, holotype male. 19. L. grjebinei, holotype male." pageId="37" pageNumber="38">Figures 19</figureCitation>
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,
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<figureCitation id="DAA62A5B62449D37EF7EFF2BFE9AEAD3" box="[293,323,186,210]" captionStart="FIGURES 56–69" captionStartId="93.[151,263,1811,1834]" captionTargetBox="[176,1418,181,1790]" captionTargetId="figure-173@93.[176,1420,181,1791]" captionTargetPageId="93" captionText="FIGURES 56–69. Leptomorphus wings. 56. L. furcatus paratype male, dorsal. 57. L. gracilis, ventral. 58. L. grjebinei, ventral. 59. L. hyalinus, ventral. 60. L. magnificus, ventral. 61. L. mandelai holotype male, ventral. 62. L. medleri, ventral. 63. L. nebulosus dark form, dorsal. Arrow indicates cell r4 (contains an extra vein in this specimen). 64. L. nebulosus light form, dorsal. 65. L. neivai ventral. 66. L. obscurus ventral. 67. L. ornatus ventral. 68. L. panorpiformis ventral. 69. L. perplexus paratype female, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1,M2,M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="37" pageNumber="38">58</figureCitation>
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,
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<figureCitation id="DAA62A5B62449D37EF14FF2BFEA2EAD3" box="[335,379,186,210]" captionStart="FIGURES 103–106" captionStartId="100.[151,264,1903,1926]" captionTargetBox="[199,1395,189,1831]" captionTargetId="figure-90@100.[168,1453,169,1891]" captionTargetPageId="100" captionText="FIGURES 103–106. Leptomorphus male genitalia, ventral. 103. L. forcipatus. 104. L. furcatus. 105. L. grjebinei. 106. L. gracilis. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="37" pageNumber="38">105</figureCitation>
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,
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<figureCitation id="DAA62A5B62449D37EFDCFF2BFE6AEAD3" box="[391,435,186,210]" captionStart="FIGURE 134–136" captionStartId="107.[151,249,1814,1837]" captionTargetBox="[151,1436,193,1793]" captionTargetId="figure-66@107.[151,1436,193,1793]" captionTargetPageId="107" captionText="FIGURE 134–136. Distribution of Afrotropical Leptomorphus. 134. L. grjebinei (red circle), L. mandelai (blue square), L. stigmatus (black triangle). 135. L. aliciae (purple star), L. carnevalei (black square), L. couturieri (black circle), L. gracilis (red triangle). 136. L. crosskeyi (red triangle), L. medleri (blue star), L. obscurus (black circle)." pageId="37" pageNumber="38">134</figureCitation>
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,
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<figureCitation id="DAA62A5B62449D37EFE4FF2BFE32EAD3" box="[447,491,186,210]" captionStart="FIGURE 150" captionStartId="113.[151,250,1941,1964]" captionTargetBox="[243,1336,190,1897]" captionTargetId="figure-51@113.[220,1371,151,1934]" captionTargetPageId="113" captionText="FIGURE 150. Strict consensus tree of three equally parsimonious trees found in the phylogenetic analysis. Bremer support values are shown above the branches and bootstrap values>50% are shown in bold below the branches." pageId="37" pageNumber="38">150</figureCitation>
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,
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<figureCitation id="DAA62A5B62449D37EFACFF2BFDFBEAD3" box="[503,546,186,210]" captionStart="FIGURE 153" captionStartId="115.[151,250,801,824]" captionTargetBox="[155,1364,186,781]" captionTargetId="figure-100@115.[151,1436,161,836]" captionTargetPageId="115" captionText="FIGURE 153. One of three equally parsimonious trees showing the relationships between species in the L. grjebinei species group. Character state changes are indicated on each branch by a black bar. Uniquely derived synapomorphies are indicated in bold and with an asterisk." pageId="37" pageNumber="38">153</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562449D37EECCFE93FBF8EB5C" pageId="37" pageNumber="38" type="reference_group">
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<paragraph id="422236DE62449D37EECCFE93FCEBEB19" blockId="37.[151,818,258,280]" box="[151,818,258,280]" pageId="37" pageNumber="38">
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<treatmentCitationGroup id="628D11F062449D37EECCFE93FCEBEB19" box="[151,818,258,280]" pageId="37" pageNumber="38">
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<treatmentCitation id="C33C10CF62449D37EECCFE93FCF7EB19" author="Matile, L." box="[151,814,258,280]" page="154" pageId="37" pageNumber="38" year="1977">
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<taxonomicName id="859D4D5D62449D37EECCFE93FCF7EB19" ID-CoL="8CJ3D" authority="Matile, 1977: 154" authorityName="Matile" authorityPageNumber="154" authorityYear="1977" box="[151,814,258,280]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="grjebinei" subGenus="Austroleptomorphus">
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<emphasis id="70E9EACC62449D37EECCFE93FDA9EB19" box="[151,624,258,280]" italics="true" pageId="37" pageNumber="38">Leptomorphus (Austroleptomorphus) grjebinei</emphasis>
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<bibRefCitation id="260C4B2F62449D37EC2DFE93FCF7EB19" author="Matile, L." box="[630,814,258,280]" pageId="37" pageNumber="38" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile, 1977: 154</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="422236DE62449D37EECCFED5FBF8EB5C" blockId="37.[151,1437,324,2034]" box="[151,1057,324,349]" pageId="37" pageNumber="38">
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<emphasis id="70E9EACC62449D37EECCFED5FEC0EB5C" bold="true" box="[151,281,324,349]" pageId="37" pageNumber="38">References</emphasis>
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:
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<bibRefCitation id="260C4B2F62449D37EF73FED4FDCFEB5C" author="Crosskey, R. W." box="[296,534,325,349]" pageId="37" pageNumber="38" refId="ref92665" refString="Crosskey, R. W. (ed.) (1980) Catalogue of the Diptera of the Afrotropical Region. British Museum of Natural History, London, 1437 pp." type="book" year="1980">Crosskey 1980: 1221</bibRefCitation>
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(catalogue appendix);
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<bibRefCitation id="260C4B2F62449D37ED4CFED4FC01EB5C" author="Matile, L." box="[791,984,325,349]" pageId="37" pageNumber="38" pagination="143 - 150" refId="ref94722" refString="Matile, L. (1997) Nouvelles donnees sur les Leptomorphus Afrotropicaux (Diptera, Mycetophilidae). Revue Francaise d'Entomologie (Nouvelle Serie) 1996, 18 (4), 143 - 150." type="journal article" year="1997">Matile 1997: 149</bibRefCitation>
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(key).
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</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562449D37EE9CFEF9FBEBE832" pageId="37" pageNumber="38" type="diagnosis">
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<paragraph id="422236DE62449D37EE9CFEF9FCEFEBED" blockId="37.[151,1437,324,2034]" pageId="37" pageNumber="38">
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<emphasis id="70E9EACC62449D37EE9CFEF9FEBDEB80" bold="true" box="[199,356,360,385]" pageId="37" pageNumber="38">DIAGNOSIS</emphasis>
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: The only extant species of
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<taxonomicName id="859D4D5D62449D37ECFBFEF8FC9BEB80" authorityName=": Edwards" authorityYear="1925" box="[672,834,361,385]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
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<emphasis id="70E9EACC62449D37ECFBFEF8FC9BEB80" box="[672,834,361,385]" italics="true" pageId="37" pageNumber="38">Leptomorphus</emphasis>
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</taxonomicName>
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with the following combination of characters: a thick medial yellow line on the otherwise brown scutum; foretibia with dense row (comb) of short anteroventral bristles; missing wing vein sc-r, male genitalia with gonostylus only a single lobe, apical tergal evaginations present (
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<figureCitation id="DAA62A5B62449D37EB35FE21FF1DEBED" captionStart="FIGURES 103–106" captionStartId="100.[151,264,1903,1926]" captionTargetBox="[199,1395,189,1831]" captionTargetId="figure-90@100.[168,1453,169,1891]" captionTargetPageId="100" captionText="FIGURES 103–106. Leptomorphus male genitalia, ventral. 103. L. forcipatus. 104. L. furcatus. 105. L. grjebinei. 106. L. gracilis. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="37" pageNumber="38">Fig. 105</figureCitation>
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). This is also the only species known from
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<collectingCountry id="3A8A764E62449D37ECF2FE45FCEBEBED" box="[681,818,468,492]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
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.
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</paragraph>
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<paragraph id="422236DE62449D37EE9CFE66FBEBE832" blockId="37.[151,1437,324,2034]" pageId="37" pageNumber="38">This species can be distinguished from other Afrotropical species by the presence of small setae on the entire surface of the scutum (not just restricted to dorsocentral lines and lateral margins).</paragraph>
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</subSubSection>
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<subSubSection id="0A87655562449D34EE9CFDAFFE1BEAF6" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
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<paragraph id="422236DE62449D37EE9CFDAFFA99EDF0" blockId="37.[151,1437,324,2034]" pageId="37" pageNumber="38">
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<emphasis id="70E9EACC62449D37EE9CFDAFFE51E856" bold="true" box="[199,392,574,599]" pageId="37" pageNumber="38">DESCRIPTION</emphasis>
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:
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<emphasis id="70E9EACC62449D37EFC5FDAEFE0EE856" box="[414,471,575,599]" italics="true" pageId="37" pageNumber="38">Male</emphasis>
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. (
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<figureCitation id="DAA62A5B62449D37EFAEFDAEFD94E856" box="[501,589,575,599]" captionStart="FIGURES 16–19" captionStartId="86.[151,263,1697,1720]" captionTargetBox="[153,1435,194,1675]" captionTargetId="figure-40@86.[151,1436,193,1676]" captionTargetPageId="86" captionText="FIGURES 16–19. Leptomorphus habitus.16. L. forcipatus, male. 17. L. furcatus, paratype male. 18. L. gracilis, holotype male. 19. L. grjebinei, holotype male." pageId="37" pageNumber="38">Fig. 19</figureCitation>
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)
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<emphasis id="70E9EACC62449D37EC3FFDAEFD7AE856" bold="true" box="[612,675,575,599]" pageId="37" pageNumber="38">Head</emphasis>
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: yellow, somewhat dorsoventrally compressed in anterior view. Antenna brown; scape yellow, with brown setae in single apical row extending from dorsum laterally into thick patch covering apicoventral process, basal third and entire medial surface bare, anterobasal patch of setulae absent; pedicel yellow, with 1 large bristle, few setae on apicodorsal margin, none ventrally; flagellomere 1 with tapered base light brown remainder light brown; flagellomere 6 1.4X as long as broad. Clypeus yellow, square; bristles on clypeus brown, 4–6 strong bristles on ventral margin directed ventrally, a number of bristles on remainder (almost in rows) all directed medioventrally, clypeus 1.1X as long as face. Face yellow; shape a just longer than wide triangle, bare. Frons yellow; with 3–4 bristles medially, frontal furrow running 1/2 distance from dorsal margin towards ventral margin, frontal cleft running to lateral ocellus. Palpus with segments 1–2 yellow, 3–5 light brown; segment 1 hidden behind eye, segments 2–4 subequal, segment 5 1.5X length of segment 4 with central half thinner than base and apex, segment 3 with apicolateral patch of fine yellow setae weakly encircled by strong dark setae. Labellum yellow. Eye with inter-ommatidial setulae absent. Occiput yellow with appressed, anteriorly directed setae. Ocelli with median slightly in front of laterals, space between ocelli less than diameter of laterals, lateral ocelli 3X their own diameter from eye margin, ocellar triangle dark brown/black.
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<emphasis id="70E9EACC62449D37EDF0FB81FBDAEE29" bold="true" box="[939,1027,1040,1064]" pageId="37" pageNumber="38">Thorax</emphasis>
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: Length
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<quantity id="85659B3B62449D37EA29FB81FB00EE26" box="[1138,1241,1040,1064]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="37" pageNumber="38" unit="mm" value="1.23">1.23 mm</quantity>
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(n = 1). Brown dorsally, yellow laterally. Scutum anterior margin and inverted medial triangle yellow/white as well as posterior corners and central lateral spot, two dark brown spots posterolaterally only reaching lateral margin at wing base; surface of scutum covered with small setae; acrostichal setae absent; single dorsocentral seta present anteriorly; single row of lateral setae present; patch of setae on scutum at wing base small. Scutellum yellow; with 2 large bristles and many small bristles. Prescutum yellow. Mediotergite brown with medial line lighter with 6–12 bristles on posterolateral corners. Laterotergite yellow; anterior margin of laterotergite not reaching katepisternum. Anepimeron yellow. Anepisternum yellow. Katepisternum yellow with small brown spot anterodorsally. Antepronotum and proepisternum yellow. Margin of anterior and posterior spiracles yellow with light brown and yellow trichia respectively. Metepisternum white. Anapleural suture straight but faint. Halter stem brown, knob white.
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<emphasis id="70E9EACC62449D37EEB9FAE4FEC0EF8C" bold="true" box="[226,281,1397,1421]" pageId="37" pageNumber="38">Legs</emphasis>
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: principally yellow; extreme anteroapical corner yellow on forefemur, on other femora dark brown; hind tibia light brown. Midfemur without apical spine-like process. Tibia with covering of brown macrotrichia, foretibia with remnant of comb of short setae along length of anteroventral surface, tibial spurs brown, foretibial spur length 2X apical thickness of foretibia, midtibia with strong, dorsal, bare patch of even thickness for 2/3 of its length, placed centrally, shortest midtibial spur subequal to length of longest, longest midtibial spur 4X apical thickness of midtibia, shortest hind tibial spur subequal to length of longest, longest hind tibial spur 3.5X apical thickness of hind tibia. Foreleg first tarsomere 1.7X length of foretibia.
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<emphasis id="70E9EACC62449D37ED50F9DAFC92EC65" bold="true" box="[779,843,1611,1636]" pageId="37" pageNumber="38">Wing</emphasis>
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(
|
||
<figureCitation id="DAA62A5B62449D37ED05F9DAFC6FEC62" box="[862,950,1611,1635]" captionStart="FIGURES 56–69" captionStartId="93.[151,263,1811,1834]" captionTargetBox="[176,1418,181,1790]" captionTargetId="figure-173@93.[176,1420,181,1791]" captionTargetPageId="93" captionText="FIGURES 56–69. Leptomorphus wings. 56. L. furcatus paratype male, dorsal. 57. L. gracilis, ventral. 58. L. grjebinei, ventral. 59. L. hyalinus, ventral. 60. L. magnificus, ventral. 61. L. mandelai holotype male, ventral. 62. L. medleri, ventral. 63. L. nebulosus dark form, dorsal. Arrow indicates cell r4 (contains an extra vein in this specimen). 64. L. nebulosus light form, dorsal. 65. L. neivai ventral. 66. L. obscurus ventral. 67. L. ornatus ventral. 68. L. panorpiformis ventral. 69. L. perplexus paratype female, dorsal. Abbreviations: A1, A2 = Anal veins; C = costa; CuA = anterior branch of cubitus; CuP = posterior branch of cubitus; h = humeral crossvein; M1,M2,M4, = branches of the media; r-m = radial-medial crossvein; Rs = radial sector vein; R1, R4, R5 = branches of the radius; Sc = subcosta; sc-r = subcostal-radial crossvein." pageId="37" pageNumber="38">Fig. 58</figureCitation>
|
||
): Length
|
||
<quantity id="85659B3B62449D37EA7CF9DAFBA6EC62" box="[1063,1151,1611,1635]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="37" pageNumber="38" unit="mm" value="5.2">5.2 mm</quantity>
|
||
(n = 1). Hyaline; apical macula absent; medial macula absent. Macrotrichia in all cells, though absent from posterobasal margin of cell a. Setae on basal posterior margin of wing (along base of cell a) alternating long, short. Calypter with a few short setae. Vein sc-r absent. R
|
||
<subScript id="DE19349B62449D37EF36F957FEACECD5" attach="left" box="[365,373,1734,1748]" fontSize="6" pageId="37" pageNumber="38">4</subScript>
|
||
absent. R
|
||
<subScript id="DE19349B62449D37EFB9F957FE33ECD5" attach="left" box="[482,490,1734,1748]" fontSize="6" pageId="37" pageNumber="38">5</subScript>
|
||
straight, slight posterior turn near tip. M
|
||
<subScript id="DE19349B62449D37EDE3F957FC19ECD5" attach="left" box="[952,960,1734,1748]" fontSize="6" pageId="37" pageNumber="38">1</subScript>
|
||
reaching apex just before R
|
||
<subScript id="DE19349B62449D37EAA4F957FADEECD5" attach="right" box="[1279,1287,1734,1748]" fontSize="6" pageId="37" pageNumber="38">5</subScript>
|
||
, apices of M veins reaching wing margin. M
|
||
<subScript id="DE19349B62449D37EFADF97DFE27ECFB" attach="left" box="[502,510,1772,1786]" fontSize="6" pageId="37" pageNumber="38">4</subScript>
|
||
-CuA fork arising after apex of r-m. A
|
||
<subScript id="DE19349B62449D37EDF6F97DFC6CECFB" attach="left" box="[941,949,1772,1786]" fontSize="6" pageId="37" pageNumber="38">2</subScript>
|
||
absent.
|
||
<emphasis id="70E9EACC62449D37EA49F94CFB5FECF7" bold="true" box="[1042,1158,1757,1782]" pageId="37" pageNumber="38">Abdomen</emphasis>
|
||
: Tergites 1 brown, T2–6 brown with lateral yellow spots, T7 with anterior 1/2 brown remainder yellow. Tergite 8 smaller than all other abdominal sclerites, with 3–4 bristles on each apicolateral corner.
|
||
<emphasis id="70E9EACC62449D37ED34F8B6FC05ED41" bold="true" box="[879,988,1831,1856]" pageId="37" pageNumber="38">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="DAA62A5B62449D37EDB0F8B6FB95ED3E" box="[1003,1100,1831,1855]" captionStart="FIGURES 103–106" captionStartId="100.[151,264,1903,1926]" captionTargetBox="[199,1395,189,1831]" captionTargetId="figure-90@100.[168,1453,169,1891]" captionTargetPageId="100" captionText="FIGURES 103–106. Leptomorphus male genitalia, ventral. 103. L. forcipatus. 104. L. furcatus. 105. L. grjebinei. 106. L. gracilis. Abbreviations: aed = aedeagus; aed ap = aedeagal apodeme; ce = cercus; gc = gonocoxite, III = section 3; gs = gonostylus; hyp = hypoproct; par = paramere; par ap = parameral apodeme; S9 = sternite 9; tg evg = apical evagination of tergite 9; vd = vas deferens. Scale bar = 0.1mm." pageId="37" pageNumber="38">Fig. 105</figureCitation>
|
||
): yellow. Sternite 9 not clearly visible possibly due to condition of specimen. Tergite 9 as wide as long, widening apically into lateral process bearing two points and medial square-shaped indentaiton. Gonocoxite placed basally on T9, medial margin reaching medial line, bearing gonostylus apically. Gonostylus a single broad-based lobe tapering to a point apically, gonocoxite III fused to dorsolateral margin. Aedeagus 4/5 length of gonocoxite, tapering towards apex (slight central swelling), apodemes 1/3 total length. Parameres a simple taper, apodemes ~4/5 length of parameres.
|
||
</paragraph>
|
||
<paragraph id="422236DE62479D34EE9CFF09FC3AEAD5" blockId="38.[151,1436,151,716]" pageId="38" pageNumber="39">
|
||
<emphasis id="70E9EACC62479D34EE9CFF09FEC2EAB1" box="[199,283,152,176]" italics="true" pageId="38" pageNumber="39">Female</emphasis>
|
||
. As for male, except as follows.
|
||
<emphasis id="70E9EACC62479D34ECC8FF09FD32EAB1" bold="true" box="[659,747,152,176]" pageId="38" pageNumber="39">Thorax</emphasis>
|
||
: Length
|
||
<quantity id="85659B3B62479D34ED08FF06FC6EEAAE" box="[851,951,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="38" pageNumber="39" unit="mm" value="1.19">1.19 mm</quantity>
|
||
(
|
||
<quantity id="85659B3B62479D34ED93FF06FBA9EAAE" box="[968,1136,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" metricValueMax="1.23" metricValueMin="1.15" pageId="38" pageNumber="39" unit="mm" value="1.19" valueMax="1.23" valueMin="1.15">1.15–1.23 mm</quantity>
|
||
, n = 2).
|
||
<emphasis id="70E9EACC62479D34EA8AFF06FACBEAB1" bold="true" box="[1233,1298,151,176]" pageId="38" pageNumber="39">Wing</emphasis>
|
||
: Length 5.0 mm (
|
||
<quantity id="85659B3B62479D34EE8AFF2DFE8CEAD5" box="[209,341,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="5.05" metricValueMax="5.5" metricValueMin="4.6" pageId="38" pageNumber="39" unit="mm" value="5.05" valueMax="5.5" valueMin="4.6">4.6–5.5mm</quantity>
|
||
, n = 2).
|
||
<emphasis id="70E9EACC62479D34EFEAFF2AFDFFEAD5" bold="true" box="[433,550,187,212]" pageId="38" pageNumber="39">Abdomen</emphasis>
|
||
: Sternites white/yellow. Cercus yellow.
|
||
</paragraph>
|
||
<paragraph id="422236DE62479D34EE9CFF70FE1BEAF6" blockId="38.[151,1436,151,716]" box="[199,450,223,248]" pageId="38" pageNumber="39">
|
||
<emphasis id="70E9EACC62479D34EE9CFF70FEE7EAF9" box="[199,318,225,248]" italics="true" pageId="38" pageNumber="39">Immatures</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562479D34EE9CFE92FE16EB1D" box="[199,463,259,284]" pageId="38" pageNumber="39" type="biology_ecology">
|
||
<paragraph id="422236DE62479D34EE9CFE92FE16EB1D" blockId="38.[151,1436,151,716]" box="[199,463,259,284]" pageId="38" pageNumber="39">
|
||
<emphasis id="70E9EACC62479D34EE9CFE92FE93EB1D" bold="true" box="[199,330,259,284]" pageId="38" pageNumber="39">BIOLOGY</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562479D34EE9CFEB6FBBEEB3E" box="[199,1127,295,320]" pageId="38" pageNumber="39" type="distribution">
|
||
<paragraph id="422236DE62479D34EE9CFEB6FBBEEB3E" blockId="38.[151,1436,151,716]" box="[199,1127,295,320]" pageId="38" pageNumber="39">
|
||
<emphasis id="70E9EACC62479D34EE9CFEB6FE4FEB41" bold="true" box="[199,406,295,320]" pageId="38" pageNumber="39">DISTRIBUTION</emphasis>
|
||
: Eastern rainforests of
|
||
<collectingCountry id="3A8A764E62479D34ECC0FEB6FCFBEB3E" box="[667,802,295,319]" name="Madagascar" pageId="38" pageNumber="39">Madagascar</collectingCountry>
|
||
(
|
||
<figureCitation id="DAA62A5B62479D34ED69FEB6FC4AEB3E" box="[818,915,295,319]" captionStart="FIGURE 134–136" captionStartId="107.[151,249,1814,1837]" captionTargetBox="[151,1436,193,1793]" captionTargetId="figure-66@107.[151,1436,193,1793]" captionTargetPageId="107" captionText="FIGURE 134–136. Distribution of Afrotropical Leptomorphus. 134. L. grjebinei (red circle), L. mandelai (blue square), L. stigmatus (black triangle). 135. L. aliciae (purple star), L. carnevalei (black square), L. couturieri (black circle), L. gracilis (red triangle). 136. L. crosskeyi (red triangle), L. medleri (blue star), L. obscurus (black circle)." pageId="38" pageNumber="39">Fig. 134</figureCitation>
|
||
),
|
||
<quantity id="85659B3B62479D34EDFCFEB6FBBAEB3E" box="[935,1123,295,319]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.1" metricValueMin="1.0" pageId="38" pageNumber="39" unit="m" value="1050.0" valueMax="1100.0" valueMin="1000.0">1000–1100 masl</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562479D34EE9CFEDAFC59EBD1" pageId="38" pageNumber="39" type="discussion">
|
||
<paragraph id="422236DE62479D34EE9CFEDAFC59EBD1" blockId="38.[151,1436,151,716]" pageId="38" pageNumber="39">
|
||
<emphasis id="70E9EACC62479D34EE9CFEDAFEABEB65" bold="true" box="[199,370,331,356]" pageId="38" pageNumber="39">DISCUSSION</emphasis>
|
||
: The female specimen from near Ranomafana has less dark brown on its thorax and tergites than the other specimens. As discussed below in the phylogeny section, the placement of
|
||
<taxonomicName id="859D4D5D62479D34EADEFEE1FAA7EB86" box="[1157,1406,367,392]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="70E9EACC62479D34EADEFEE1FAFEEB89" box="[1157,1319,368,392]" italics="true" pageId="38" pageNumber="39">Leptomorphus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
in subgenera (
|
||
<bibRefCitation id="260C4B2F62479D34EF43FE05FE7DEBAD" author="Matile, L." box="[280,420,404,428]" pageId="38" pageNumber="39" pagination="141 - 155" refId="ref94584" refString="Matile, L. (1977) Notes sur le genre Leptomorphus et description de taxa nouveaux de la region ethiopienne (Diptera, Mycetophilidae). Bulletin de l'Institute Fondamental de Afrique Noire (A), 38 (1976), 141 - 155." type="journal article" year="1977">Matile 1977</bibRefCitation>
|
||
) is not supported by our phylogenetic results. This species is therefore removed from the subgenus
|
||
<taxonomicName id="859D4D5D62479D34EF5DFE29FE30EBD1" box="[262,489,440,464]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Austroleptomorphus">
|
||
<emphasis id="70E9EACC62479D34EF5DFE29FE30EBD1" box="[262,489,440,464]" italics="true" pageId="38" pageNumber="39">Austroleptomorphus</emphasis>
|
||
</taxonomicName>
|
||
and placed solely in
|
||
<emphasis id="70E9EACC62479D34EC8CFE29FC59EBD1" box="[727,896,440,464]" italics="true" pageId="38" pageNumber="39">
|
||
<taxonomicName id="859D4D5D62479D34EC8CFE29FCA4EBD1" authorityName=": Edwards" authorityYear="1925" box="[727,893,440,464]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0A87655562479D34EE9CFE4AFE50E8CD" pageId="38" pageNumber="39" type="materials_examined">
|
||
<paragraph id="422236DE62479D34EE9CFE4AFD8BE85E" blockId="38.[151,1436,151,716]" pageId="38" pageNumber="39">
|
||
<materialsCitation id="F2F53C8362479D34EE9CFE4AFD8BE85E" collectionCode="MATERIAL, MNHN" collectorName="B. Stuckenberg" country="Madagascar" county="Ambohitra" determinerName="L. Matile" elevation="1000" location="Madagascar Nord" municipality="Montagne d'Ambre" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<emphasis id="70E9EACC62479D34EE9CFE4AFDDBEBF5" bold="true" box="[199,514,475,500]" pageId="38" pageNumber="39">
|
||
<collectionCode id="248CAE1B62479D34EE9CFE4AFE86EBF5" box="[199,351,475,500]" pageId="38" pageNumber="39">MATERIAL</collectionCode>
|
||
EXAMINED
|
||
</emphasis>
|
||
:
|
||
<emphasis id="70E9EACC62479D34EC4BFE4DFD59EBF5" box="[528,640,476,500]" italics="true" pageId="38" pageNumber="39">
|
||
<typeStatus id="9D26887C62479D34EC4BFE4DFDA3EBF5" box="[528,634,476,500]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
adult male, pinned on double mount minuten, genitalia in glass vial on pin, labelled “[
|
||
<collectingCountry id="3A8A764E62479D34EF1AFE6EFDD2E816" box="[321,523,511,535]" name="Madagascar" pageId="38" pageNumber="39">MADAGASCAR</collectingCountry>
|
||
]
|
||
<location id="4742600562479D34EC40FE6EFD3AE816" LSID="urn:lsid:plazi:treatment:CA3487C862449D34EECCFF06FE50E8CD:4742600562479D34EC40FE6EFD3AE816" box="[539,739,511,535]" country="Madagascar" county="Ambohitra" municipality="Montagne d'Ambre" name="Madagascar Nord" pageId="38" pageNumber="39">Madagascar Nord</location>
|
||
/
|
||
<collectingMunicipality id="A246ACA462479D34ECAEFD91FC0DE816" box="[757,980,511,536]" pageId="38" pageNumber="39">Montagne d’Ambre</collectingMunicipality>
|
||
[
|
||
<collectingCounty id="AB434E5262479D34EDB8FE6EFBBBE816" box="[995,1122,511,535]" pageId="38" pageNumber="39">Ambohitra</collectingCounty>
|
||
]
|
||
<quantity id="85659B3B62479D34EA2AFE6EFB67E816" box="[1137,1214,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="m" value="1000.0">
|
||
<elevation id="C9B0D1ED62479D34EA2AFE6EFB67E816" box="[1137,1214,511,535]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="38" pageNumber="39" unit="m" value="1000.0">1000m</elevation>
|
||
</quantity>
|
||
/ det Diego-Suarez/ 23.XI–4.XII.[19]57,
|
||
<collectorName id="EF68530862479D34EFDAFDB5FDE9E83D" box="[385,560,548,572]" pageId="38" pageNumber="39">B. Stuckenberg</collectorName>
|
||
; [Red label]
|
||
<typeStatus id="9D26887C62479D34EC9BFDB5FC8EE83D" box="[704,855,548,572]" pageId="38" pageNumber="39" type="holotype">HOLOTYPE</typeStatus>
|
||
;
|
||
<taxonomicName id="859D4D5D62479D34ED39FDB5FBDDE83D" authorityName=": Edwards" authorityYear="1925" box="[866,1028,548,572]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="genus">Leptomorphus</taxonomicName>
|
||
/ (
|
||
<taxonomicName id="859D4D5D62479D34EA47FDB5FADCE83D" box="[1052,1285,548,572]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="subGenus" subGenus="Austroleptomorphus">Austroleptomorphus</taxonomicName>
|
||
)/ grjebinei
|
||
<taxonomicNameLabel id="6BDA57B762479D34EBDDFDB5FF6EE85E" pageId="38" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
|
||
<specimenCount id="549BFD5762479D34EEE5FDD9FF0AE85E" box="[190,211,584,607]" pageId="38" pageNumber="39" type="male">♂</specimenCount>
|
||
ht/
|
||
<determinerName id="B7FEDA1A62479D34EEACFDD9FE86E85E" box="[247,351,583,607]" pageId="38" pageNumber="39">L. Matile</determinerName>
|
||
det. 1974” [
|
||
<collectionCode id="248CAE1B62479D34EFB2FDD9FD91E85E" box="[489,584,584,607]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="38" pageNumber="39">MNHN</collectionCode>
|
||
].
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="422236DE62479D34EE9CFDFCFC03E885" blockId="38.[151,1436,151,716]" box="[199,986,620,644]" pageId="38" pageNumber="39">
|
||
<materialsCitation id="F2F53C8362479D34EE9CFDFCFC0FE885" box="[199,982,620,644]" collectionCode="MATERIAL, MNHN" collectorName="B. Stuckenberg" country="Madagascar" county="Ambohitra" determinerName="L. Matile" elevation="1000" location="Madagascar Nord" municipality="Montagne d'Ambre" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Monaghan" typeStatus="paratype">
|
||
<emphasis id="70E9EACC62479D34EE9CFDFCFE98E885" box="[199,321,621,644]" italics="true" pageId="38" pageNumber="39">
|
||
<typeStatus id="9D26887C62479D34EE9CFDFCFEE5E885" box="[199,316,621,644]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
Same data as holotype (
|
||
<specimenCount id="549BFD5762479D34EC0DFDFDFDAEE882" box="[598,631,620,644]" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
|
||
, labelled as
|
||
<typeStatus id="9D26887C62479D34ED58FDFDFCB1E885" box="[771,872,620,644]" pageId="38" pageNumber="39" type="allotype">Allotype</typeStatus>
|
||
,
|
||
<collectingRegion id="8059F83C62479D34ED28FDFDFC17E885" box="[883,974,620,644]" country="Ireland" name="Monaghan" pageId="38" pageNumber="39">MNHN</collectingRegion>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="422236DE62479D34EE9CFD01FE50E8CD" blockId="38.[151,1436,151,716]" pageId="38" pageNumber="39">
|
||
<emphasis id="70E9EACC62479D34EE9CFD01FEA7E8A9" box="[199,382,656,680]" italics="true" pageId="38" pageNumber="39">Other material:</emphasis>
|
||
<materialsCitation id="F2F53C8362479D34EFD7FD1EFE5CE8CD" collectingDate="1988-11-01" collectingDateMax="1988-11-07" collectingDateMin="1988-11-01" collectionCode="USNM" collectorName="W. E. Steiner" country="Madagascar" elevation="1100" location="Ranomafana" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Fianarantsoa">
|
||
<collectingCountry id="3A8A764E62479D34EFD7FD1EFD8FE8A6" box="[396,598,655,679]" name="Madagascar" pageId="38" pageNumber="39">MADAGASCAR</collectingCountry>
|
||
, Prov.
|
||
<collectingRegion id="8059F83C62479D34ECEAFD1EFC9AE8A6" box="[689,835,655,679]" country="Madagascar" name="Fianarantsoa" pageId="38" pageNumber="39">Fianarantsoa</collectingRegion>
|
||
,
|
||
<quantity id="85659B3B62479D34ED0EFD1EFC5EE8A6" box="[853,903,655,679]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="38" pageNumber="39" unit="km" value="7.0">7km</quantity>
|
||
W
|
||
<location id="4742600562479D34EDE5FD1EFB97E8A6" LSID="urn:lsid:plazi:treatment:CA3487C862449D34EECCFF06FE50E8CD:4742600562479D34EDE5FD1EFB97E8A6" box="[958,1102,655,679]" country="Madagascar" name="Ranomafana" pageId="38" pageNumber="39" stateProvince="Fianarantsoa">Ranomafana</location>
|
||
,
|
||
<quantity id="85659B3B62479D34EA3BFD1EFB69E8A6" box="[1120,1200,655,679]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="38" pageNumber="39" unit="m" value="1100.0">
|
||
<elevation id="C9B0D1ED62479D34EA3BFD1EFB69E8A6" box="[1120,1200,655,679]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="38" pageNumber="39" unit="m" value="1100.0">1100m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="3623101E62479D34EA99FD1EFA8BE8A6" box="[1218,1362,655,679]" pageId="38" pageNumber="39" value="1988-11-01" valueMax="1988-11-07" valueMin="1988-11-01">
|
||
<collectingDate id="2667E9F662479D34EA99FD1EFA8BE8A6" box="[1218,1362,655,679]" pageId="38" pageNumber="39" value="1988-11-01" valueMax="1988-11-07" valueMin="1988-11-01">1–7.XI.1988</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="EF68530862479D34EB3FFD01FF3FE8CD" pageId="38" pageNumber="39">W.E. Steiner</collectorName>
|
||
(
|
||
<specimenCount id="549BFD5762479D34EEA3FD25FEC0E8CA" box="[248,281,692,716]" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
|
||
,
|
||
<collectionCode id="248CAE1B62479D34EF7CFD25FEA4E8CD" box="[295,381,692,716]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="38" pageNumber="39" type="Museum">USNM</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |