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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.39.26743" ID-GBIF-Dataset="c6ddb185-aac9-4680-bc6d-4ce38c290bb5" ID-PMC="PMC6160792" ID-Pensoft-Pub="1314-4049-39-41" ID-PubMed="30271257" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-4049-39-41" ModsDocOrigin="MycoKeys 39" ModsDocTitle="The genus Coprinellus (Basidiomycota; Agaricales) in Pakistan with the description of four new species" checkinTime="1555333497602" checkinUser="pensoft" docAuthor="Hussain, Shah, Usman, Muhammad, Afshan, Najam-ul-Sehar, Ahmad, Habib, Khan, Junaid &amp; Khalid, Abdul Nasir" docDate="2018" docId="A5E862E5C7D0E141B1BA87727D8027C7" docLanguage="en" docName="MycoKeys 39: 41-61" docOrigin="MycoKeys 39" docSource="http://dx.doi.org/10.3897/mycokeys.39.26743" docTitle="Coprinellus tenuis Hussain, sp. nov." docType="treatment" docVersion="4" lastPageNumber="49" masterDocId="FFCEFFC7FFCBFFD0AD45F21A736ED27E" masterDocTitle="The genus Coprinellus (Basidiomycota; Agaricales) in Pakistan with the description of four new species" masterLastPageNumber="61" masterPageNumber="41" pageNumber="47" updateTime="1668136053748" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The genus Coprinellus (Basidiomycota; Agaricales) in Pakistan with the description of four new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hussain, Shah</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Usman, Muhammad</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Afshan, Najam-ul-Sehar</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ahmad, Habib</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khan, Junaid</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khalid, Abdul Nasir</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>39</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>41</mods:start>
<mods:end>61</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.39.26743</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.39.26743</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-39-41</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="156202305" LSID="urn:lsid:plazi:treatment:A5E862E5C7D0E141B1BA87727D8027C7" httpUri="http://treatment.plazi.org/id/A5E862E5C7D0E141B1BA87727D8027C7" lastPageId="8" lastPageNumber="49" pageId="6" pageNumber="47">
<subSubSection pageId="6" pageNumber="47" type="nomenclature">
<paragraph pageId="6" pageNumber="47">
<taxonomicName LSID="MB825479" authority="Hussain" class="Agaricomycetes" family="Psathyrellaceae" genus="Coprinellus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coprinellus tenuis" order="Agaricales" pageId="6" pageNumber="47" phylum="Basidiomycota" rank="species" species="tenuis">Coprinellus tenuis Hussain</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="47">sp. nov.</taxonomicNameLabel>
Figures 1
<normalizedToken originalValue="CDand">C-Dand</normalizedToken>
7
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="48" type="diagnosis">
<paragraph pageId="7" pageNumber="48">
<pageBreakToken pageId="7" pageNumber="48" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph pageId="7" pageNumber="48">
The new species
<taxonomicName class="Agaricomycetes" family="Psathyrellaceae" genus="Coprinellus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coprinellus tenuis" order="Agaricales" pageId="7" pageNumber="48" phylum="Basidiomycota" rank="species" species="tenuis">Coprinellus tenuis</taxonomicName>
can be recognised by its thin and membranous pileus, surface glabrous and furred, deeply plicate towards margin; lamellae sinuate to uncinate; basidiospores 10.5-14.5
<normalizedToken originalValue="×">x</normalizedToken>
8.0-9.5
<normalizedToken originalValue="×">x</normalizedToken>
6.5-8.5
<normalizedToken originalValue="µm">µm</normalizedToken>
, in face view, broadly ellipsoid to ovoid, in side view, slightly pyriform to ellipsoid, usually with truncate base, apiculus mostly not visible, with eccentric germ-pore, 1.5-2
<normalizedToken originalValue="µm">µm</normalizedToken>
wide.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="materials_examined">
<paragraph pageId="8" pageNumber="49">
<pageBreakToken pageId="8" pageNumber="49" start="start">Type</pageBreakToken>
.
</paragraph>
<paragraph pageId="8" pageNumber="49">PAKISTAN: Khyber Pakhtunkhwa, Malakand, Qaldara, 430 m alt., solitary on leaf litter, 7 July 2014, S. Hussain, SHP10 (SWAT-SH-P10, holotype); GenBank accession ITS: MH753663.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="etymology">
<paragraph pageId="8" pageNumber="49">Etymology.</paragraph>
<paragraph pageId="8" pageNumber="49">
<normalizedToken originalValue="“tenuis”">&quot;tenuis&quot;</normalizedToken>
(Latin) meaning thin, referring to the membranous pileus of the new species.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="description">
<paragraph pageId="8" pageNumber="49">Macroscopic characters.</paragraph>
<paragraph pageId="8" pageNumber="49">
Pileus 15-20 mm diam, pulvinate to convex to plane, light greyish-brown (7.5YR 5/2) to light brown (5YR 6/4); surface glabrous, furred, deeply plicate from centre towards margin; centre truncately conical, moderate reddish-orange (10R 5/8) to greyish-reddish-orange (2.5YR 5/6); context membranous. Lamellae sinuate to uncinate, distant, with 0-2 series of lamelullae, light greyish-brown (7.5YR 5/2) to light brown (5YR 6/4), lamellae edge blackish and fimbriate to eroded. Stipe 40-60
<normalizedToken originalValue="×">x</normalizedToken>
1 mm, equal, cylindrical, surface scabrous, white, translucent, fragile, context hollow.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="description">
<paragraph pageId="8" pageNumber="49">Microscopic characters.</paragraph>
<paragraph pageId="8" pageNumber="49">
Basidiospores (9.0
<normalizedToken originalValue=")10.514.5(">-)10.5-14.5(-</normalizedToken>
15.5)
<normalizedToken originalValue="×">x</normalizedToken>
(7.5
<normalizedToken originalValue=")8.09.5(">-)8.0-9.5(-</normalizedToken>
10.5)
<normalizedToken originalValue="×">x</normalizedToken>
(5.0
<normalizedToken originalValue=")6.58.5(">-)6.5-8.5(-</normalizedToken>
9.0)
<normalizedToken originalValue="µm">µm</normalizedToken>
, on average 13.1
<normalizedToken originalValue="×">x</normalizedToken>
9.0
<normalizedToken originalValue="×">x</normalizedToken>
7.8
<normalizedToken originalValue="µm">µm</normalizedToken>
; Q1 = 1.25-1.49, Q2 = 1.57-1.63, av. Q = 1.45; in face view, broadly ellipsoid to ovoid, in side view, slightly pyriform to ellipsoid, usually with truncate base, apiculus mostly not visible, germ-pore eccentric, 1.5-2
<normalizedToken originalValue="µm">µm</normalizedToken>
wide, wall 1.5
<normalizedToken originalValue="µm">µm</normalizedToken>
thick, dark brown to almost black. Basidia 22-24
<normalizedToken originalValue="×">x</normalizedToken>
9-12
<normalizedToken originalValue="µm">µm</normalizedToken>
, clavate, 2- to 4-spored, hyaline in KOH. Cheilocystidia 22-30
<normalizedToken originalValue="×">x</normalizedToken>
19-28
<normalizedToken originalValue="µm">µm</normalizedToken>
, rounded to globose, abundant, hyaline. Pleurocystidia absent. Pileocystidia 78-94
<normalizedToken originalValue="×">x</normalizedToken>
10-12
<normalizedToken originalValue="µm">µm</normalizedToken>
, lageniform to cylindrical with rounded apex, elongated rod shape neck with rounded enlarged base, hyaline in KOH. Caulocystidia 50-67
<normalizedToken originalValue="×">x</normalizedToken>
9-11
<normalizedToken originalValue="µm">µm</normalizedToken>
, narrowly clavate to clavate, with rounded to obtuse apex, cylindrical base. Veil comprised of rounded to subglobose cells, arranged in short chain, thick-walled with encrusted walls, dark brown, with terminal cell 17-23
<normalizedToken originalValue="×">x</normalizedToken>
12-15
<normalizedToken originalValue="µm">µm</normalizedToken>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="habitat">
<paragraph pageId="8" pageNumber="49">Habitat and distribution.</paragraph>
<paragraph pageId="8" pageNumber="49">
Scattered on leaf litter under
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia modesta" order="Fabales" pageId="8" pageNumber="49" phylum="Tracheophyta" rank="species" species="modesta">Acacia modesta</taxonomicName>
, so far only known from lowland northern Pakistan.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="materials_examined">
<paragraph pageId="8" pageNumber="49">Additional specimens examined.</paragraph>
<paragraph pageId="8" pageNumber="49">
PAKISTAN. Khyber Pakhtunkhwa: Malakand, Qaldara, on leaf litter under
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia modesta" order="Fabales" pageId="8" pageNumber="49" phylum="Tracheophyta" rank="species" species="modesta">Acacia modesta</taxonomicName>
, 10 July 2014, S. Hussain, SH10 (SWAT SH-10).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="49" type="comments">
<paragraph pageId="8" pageNumber="49">Comments.</paragraph>
<paragraph pageId="8" pageNumber="49">
<taxonomicName class="Agaricomycetes" family="Psathyrellaceae" genus="Coprinellus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coprinellus tenuis" order="Agaricales" pageId="8" pageNumber="49" phylum="Basidiomycota" rank="species" species="tenuis">Coprinellus tenuis</taxonomicName>
with thin membranous pileus, shows similarities with
<taxonomicName lsidName="Co. curtus" pageId="8" pageNumber="49" rank="species" species="curtus">Co. curtus</taxonomicName>
. Both these species can be differentiated on (i) pileus morphology (ii) basidiospore shape and (iii) habitat. Pileus is deeply plicate in both these species, in
<taxonomicName lsidName="Co. tenuis" pageId="8" pageNumber="49" rank="species" species="tenuis">Co. tenuis</taxonomicName>
pileus is glabrous and furred; however, there is no furcation in the pileus of
<taxonomicName lsidName="Co. curtus" pageId="8" pageNumber="49" rank="species" species="curtus">Co. curtus</taxonomicName>
. Spores in
<taxonomicName lsidName="Co. curtus" pageId="8" pageNumber="49" rank="species" species="curtus">Co. curtus</taxonomicName>
are substantially smaller (8.0-10.0
<normalizedToken originalValue="×">x</normalizedToken>
5.5-7.0
<normalizedToken originalValue="µm">µm</normalizedToken>
), ellipsoid to ovoid in face view, narrowly ellipsoid or phaseoliform in side view, apiculus often not visible, with a distinct central to slightly eccentric germ-pore, not truncate. Basidiospores in
<taxonomicName lsidName="Co. tenuis" pageId="8" pageNumber="49" rank="species" species="tenuis">Co. tenuis</taxonomicName>
are larger (10.5-14.5
<normalizedToken originalValue="×">x</normalizedToken>
8.0-9.5
<normalizedToken originalValue="×">x</normalizedToken>
6.5-8.5
<normalizedToken originalValue="µm">µm</normalizedToken>
), in face view broadly ellipsoid to ovoid, in side view slightly pyriform to ellipsoid, usually with truncate base, apiculus mostly not visible, with eccentric germ-pore of 1.5-2
<normalizedToken originalValue="µm">µm</normalizedToken>
diam.
<taxonomicName class="Agaricomycetes" family="Psathyrellaceae" genus="Coprinellus" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Coprinellus curtus" order="Agaricales" pageId="8" pageNumber="49" phylum="Basidiomycota" rank="species" species="curtus">Coprinellus curtus</taxonomicName>
has a substrate preference and is most commonly collected from
<normalizedToken originalValue="herbivores">herbivores'</normalizedToken>
dung as opposed to
<taxonomicName lsidName="Co. tenuis" pageId="8" pageNumber="49" rank="species" species="tenuis">Co. tenuis</taxonomicName>
basidioma on leaf litter (
<bibRefCitation author="Ulje, CB" journalOrPublisher="Persoonia" pageId="12" pageNumber="53" pagination="275 - 339" title="Studies in Coprinus II. Subsection Setulosi of section Pseudocoprinus" volume="14" year="1991">
<normalizedToken originalValue="Uljé">Ulje</normalizedToken>
and Bas 1991
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>