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<document id="5F44A4ECF6BE1E33D8E768BC8C1A6647" ID-DOI="10.5281/zenodo.193406" ID-GBIF-Dataset="5e8043a5-e685-4a21-b696-0f7a706ab223" ID-ISSN="1175-5326" ID-Zenodo-Dep="193406" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460138703176" checkinUser="plazi" docAuthor="Toledo, Mario, Spangler, Paul J. &amp; Balke, Michael" docDate="2010" docId="11578788FE5BFFCA7DB5FC2414F3D9C8" docLanguage="en" docName="zt02347p058.pdf" docOrigin="Zootaxa 2347" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Laccodytes rondonia Toledo, Spangler &amp; Balke, 2010, sp.n." docType="treatment" docVersion="9" lastPageNumber="45" masterDocId="ED6EFFF0FE5CFFC27D22FFF3160ED852" masterDocTitle="Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)" masterLastPageNumber="58" masterPageNumber="37" pageNumber="44" updateTime="1698235333712" updateUser="plazi">
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<mods:title id="575D70EB2D93089ABC7B8EA53C2F4920">Taxonomic revision of the Neotropical diving beetles genus Laccodytes Régimbart, 1895 (Coleoptera: Dytiscidae)</mods:title>
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<mods:namePart id="0145DD8F7117D0FAB37A9DE5122435A6">Toledo, Mario</mods:namePart>
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<mods:namePart id="7BCBBBAC08385A2BE13243C2064E436A">Spangler, Paul J.</mods:namePart>
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<mods:namePart id="7295A76F8DF85735160A9FA7C6540E41">Balke, Michael</mods:namePart>
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<mods:title id="A2E7C0147EC75CD937CD268921AB4333">Zootaxa</mods:title>
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<mods:date id="0A488883DB299F268C1500FA50D922A1">2010</mods:date>
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<treatment id="11578788FE5BFFCA7DB5FC2414F3D9C8" ID-DOI="http://doi.org/10.5281/zenodo.6198366" ID-GBIF-Taxon="124955824" ID-Zenodo-Dep="6198366" LSID="urn:lsid:plazi:treatment:11578788FE5BFFCA7DB5FC2414F3D9C8" httpUri="http://treatment.plazi.org/id/11578788FE5BFFCA7DB5FC2414F3D9C8" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44">
<subSubSection id="D1E46515FE5BFFC57DB5FC2417F4DC46" pageId="7" pageNumber="44" type="nomenclature">
<paragraph id="9941369EFE5BFFC57DB5FC2417DBDBA3" blockId="7.[151,506,983,1044]" box="[151,469,983,1009]" pageId="7" pageNumber="44">
<heading id="C20981F2FE5BFFC57DB5FC2417DBDBA3" bold="true" box="[151,469,983,1009]" fontSize="11" level="1" pageId="7" pageNumber="44" reason="1">
<emphasis id="AB8AEA8CFE5BFFC57DB5FC2417DBDBA3" bold="true" box="[151,469,983,1009]" pageId="7" pageNumber="44">
<taxonomicName id="5EFE4D1DFE5BFFC57DB5FC24179DDBA3" ID-CoL="3RLZN" box="[151,403,983,1009]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="rondonia" status="sp. nov.">
<emphasis id="AB8AEA8CFE5BFFC57DB5FC24179DDBA3" bold="true" box="[151,403,983,1009]" italics="true" pageId="7" pageNumber="44">Laccodytes rondonia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B0B957F7FE5BFFC57CB8FC2417DBDBA3" box="[410,469,983,1009]" pageId="7" pageNumber="44" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9941369EFE5BFFC57DB5FC0917F4DC46" blockId="7.[151,506,983,1044]" box="[151,506,1018,1044]" pageId="7" pageNumber="44">Figs (1, 26, 29, 42, 45, 55, 62)</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE5BFFC57DB5FBB41392DCFD" pageId="7" pageNumber="44" type="materials_examined">
<paragraph id="9941369EFE5BFFC57DB5FBB41527DC30" blockId="7.[151,1437,1095,2011]" box="[151,809,1095,1122]" pageId="7" pageNumber="44">
<emphasis id="AB8AEA8CFE5BFFC57DB5FBB41733DC33" bold="true" box="[151,317,1095,1121]" pageId="7" pageNumber="44">
<typeStatus id="4645883CFE5BFFC57DB5FBB416DADC33" box="[151,212,1095,1121]" pageId="7" pageNumber="44">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="E1E9760EFE5BFFC57C66FBBB179EDC30" box="[324,400,1096,1122]" name="Brazil" pageId="7" pageNumber="44">Brazil</collectingCountry>
:
<taxonomicName id="5EFE4D1DFE5BFFC57CB8FBBB1401DC30" box="[410,527,1096,1122]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="rondonia">Rondônia</taxonomicName>
,
<quantity id="5E069B7BFE5BFFC57F39FBBB146BDC30" box="[539,613,1096,1122]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="44" unit="km" value="62.0">62 km</quantity>
SW Ariquemes.
</paragraph>
<paragraph id="9941369EFE5BFFC57DE4FB9D1392DCFD" blockId="7.[151,1437,1095,2011]" pageId="7" pageNumber="44">
<emphasis id="AB8AEA8CFE5BFFC57DE4FB9D1774DCDA" bold="true" box="[198,378,1134,1160]" pageId="7" pageNumber="44">
<typeStatus id="4645883CFE5BFFC57DE4FB9D170FDCDA" box="[198,257,1134,1160]" pageId="7" pageNumber="44">Type</typeStatus>
material.
</emphasis>
<typeStatus id="4645883CFE5BFFC57CA7FB9D17FCDCDA" box="[389,498,1134,1160]" pageId="7" pageNumber="44" type="holotype">Holotype</typeStatus>
3 (
<collectionCode id="FFEFAE5BFE5BFFC57F02FB9D147CDCDA" box="[544,626,1134,1160]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="44">NMW</collectionCode>
):
<collectingCountry id="E1E9760EFE5BFFC57FA4FB9D14DFDCDA" box="[646,721,1134,1160]" name="Brazil" pageId="7" pageNumber="44">Brazil</collectingCountry>
,
<taxonomicName id="5EFE4D1DFE5BFFC57FFFFB9D155CDCDA" box="[733,850,1134,1160]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="57" phylum="Arthropoda" rank="species" species="rondonia">Rondônia</taxonomicName>
,
<quantity id="5E069B7BFE5BFFC57E7DFB9D15A4DCDA" box="[863,938,1134,1160]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="7" pageNumber="44" unit="km" value="62.0">62 km</quantity>
SW Ariquemes, near Fz. Rancho Grande,
<date id="ED40105EFE5BFFC57DB5FB66174ADCFD" box="[151,324,1173,1199]" pageId="7" pageNumber="44" value="1994-11-08" valueMax="1994-11-20" valueMin="1994-11-08">8.20.XI.1994</date>
, Eger &amp; O´Brien.
<typeStatus id="4645883CFE5BFFC57F3BFB661481DCFD" box="[537,655,1173,1199]" pageId="7" pageNumber="44" type="paratype">Paratypes</typeStatus>
: same data as
<typeStatus id="4645883CFE5BFFC57E16FB661594DCFD" box="[820,922,1173,1199]" pageId="7" pageNumber="44" type="holotype">holotype</typeStatus>
(15 exs
<collectionCode id="FFEFAE5BFE5BFFC57EDCFB661232DCFD" box="[1022,1084,1173,1199]" pageId="7" pageNumber="44">CLH</collectionCode>
,
<collectionCode id="FFEFAE5BFE5BFFC57964FB661289DCFD" box="[1094,1159,1173,1199]" pageId="7" pageNumber="44">CMT</collectionCode>
,
<collectionCode id="FFEFAE5BFE5BFFC579B0FB6612D0DCFD" box="[1170,1246,1173,1199]" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="7" pageNumber="44">NMW</collectionCode>
,
<collectionCode id="FFEFAE5BFE5BFFC579CAFB66134DDCFD" box="[1256,1347,1173,1199]" pageId="7" pageNumber="44">BMNH</collectionCode>
,
<collectionCode id="FFEFAE5BFE5BFFC5786CFB661381DCFD" box="[1358,1423,1173,1199]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="44">ZSM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE5BFFCA7DE4FB4817D5D974" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44" type="diagnosis">
<paragraph id="9941369EFE5BFFC57DE4FB48131FDD71" blockId="7.[151,1437,1095,2011]" pageId="7" pageNumber="44">
<emphasis id="AB8AEA8CFE5BFFC57DE4FB481748DC87" bold="true" box="[198,326,1211,1237]" pageId="7" pageNumber="44">Diagnosis.</emphasis>
Habitus (
<figureCitation id="01C52A1BFE5BFFC57C95FB4F140FDC84" box="[439,513,1212,1238]" captionStart="FIGURES 1 2" captionStartId="2.[151,270,2000,2024]" captionTargetBox="[168,1422,904,1961]" captionTargetId="figure@2.[151,1436,896,1975]" captionTargetPageId="2" captionText="FIGURES 1 2. Habitus of: 1) Laccodytes rondonia sp. n.; 2) L. bassignanii sp. n." httpUri="https://zenodo.org/record/193407/files/figure.png" pageId="7" pageNumber="44">Figs 1</figureCitation>
,
<figureCitation id="01C52A1BFE5BFFC57F2DFB4F143EDC84" box="[527,560,1212,1238]" captionStart="FIGURES 28 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="7" pageNumber="44">29</figureCitation>
). Close to the preceding species, from which differs principally for the less extended and sharper dark patterns and aedeagal features. Body length 1.7 mm. Body slightly drop-shaped, tip of elytron truncated.
<collectingCountry id="E1E9760EFE5BFFC57CAEFAFA17C9DD71" box="[396,455,1289,1315]" name="India" pageId="7" pageNumber="44">Hind</collectingCountry>
angle of pronotum slightly produced backwards, faintly needle-shaped.
</paragraph>
<paragraph id="9941369EFE5BFFC57DE4FAC314C2DDC5" blockId="7.[151,1437,1095,2011]" pageId="7" pageNumber="44">
Color. Head and pronotum yellow, middle of base a bit darker. Elytron black with contrasting yellow pattern as in
<figureCitation id="01C52A1BFE5BFFC57C14FAA5179EDD22" box="[310,400,1366,1392]" captionStart="FIGURES 28 39" captionStartId="8.[151,272,1875,1899]" captionTargetBox="[154,1429,467,1844]" captionTargetId="figure@8.[151,1436,467,1848]" captionTargetPageId="8" captionText="FIGURES 28 39. Subschematic body shape and colour patterns: 28) Laccodytes apalodes; 29) L. rondonia; 30) L. americanus; 31) L. phalacroides; 32) L. takutuanus; 33 a 33 b) L. neblinae (variation of elytral patterns); 34) L. bassignanii; 35) L. obscuratus; 36) L. olibroides; 37) L. androginus; 38) Laccodytes sp. 1.; 39) Laccodytes sp. 2." httpUri="https://zenodo.org/record/193411/files/figure.png" pageId="7" pageNumber="44">Fig. 29</figureCitation>
, entire marginal area yellow except for tiny darker area at base, below pronotal hind angle. Appendages and venter yellow to orange.
</paragraph>
<paragraph id="9941369EFE5BFFC57DE4FA571729DEF4" blockId="7.[151,1437,1095,2011]" pageId="7" pageNumber="44">Sculpture. Dorsal and ventral surface of the beetle with a faint MR, except for base of metaventrite as well as posterior part of metacoxal process. Rather regular, honeycomb shaped on head and pronotum; more irregular, slightly elongate and horizontally orientated on elytron, apparently with very small punctures (100x). Metaventrite with MR of larger polygonal meshes posteriorly and few punctures; metaxcoxal process with MR anteriorly, posteriorly only with punctures; metacoxa with numerous diagonally orientated longish cells, giving the impression of a dense diagonal striation or dense punctation depending on angle of observation.</paragraph>
<paragraph id="9941369EFE5BFFC57DE4F9411769DFDC" blockId="7.[151,1437,1095,2011]" pageId="7" pageNumber="44">
Structures. Pronotum with narrow lateral bead; posterior angle produced backwards, acute, but less developed than in
<taxonomicName id="5EFE4D1DFE5BFFC57C5EF92A1402DEA0" box="[380,524,1753,1778]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="44" phylum="Arthropoda" rank="species" species="apalodes">
<emphasis id="AB8AEA8CFE5BFFC57C5EF92A1402DEA0" box="[380,524,1753,1778]" italics="true" pageId="7" pageNumber="44">L. apalodes</emphasis>
</taxonomicName>
. Prosternum with ridge, prosternal process not ridged; prosternal process carinate and with a very long, needle shaped tip (
<figureCitation id="01C52A1BFE5BFFC57FD3F8F31543DF48" box="[753,845,1792,1818]" captionStart="FIGURES 40 43" captionStartId="9.[151,276,898,922]" captionTargetBox="[156,1431,199,863]" captionTargetId="figure@9.[151,1436,194,874]" captionTargetPageId="9" captionText="FIGURES 40 43. Line drawings of prosternal process: 40) Laccodytes takutuanus; 41) L. phalacroides; 42) L. rondonia; 43) L. olibroides." httpUri="https://zenodo.org/record/193412/files/figure.png" pageId="7" pageNumber="44">Fig. 42</figureCitation>
). Epipleuron broad up to level of sternite 6 or 7. Metatibia very short and compact, only about 1/2 as long as metafemur. Metatarsomeres 14 with apicolateral angle distinctly lobed.
<collectingCountry id="E1E9760EFE5BFFC57CD5F8BE143CDF35" box="[503,562,1869,1895]" name="India" pageId="7" pageNumber="44">Hind</collectingCountry>
lobes of metacoxal process wing shaped, with a large and wide V-shaped notch in between.
</paragraph>
<paragraph id="9941369EFE5BFFCA7DE4F86915F3D88A" blockId="7.[151,1437,1095,2011]" lastBlockId="8.[151,1437,152,410]" lastPageId="8" lastPageNumber="45" pageId="7" pageNumber="44">
Male. Pro- and mesotarsi slightly dilated.
<collectingCountry id="E1E9760EFE5BFFC57F90F86914E3DFE6" box="[690,749,1946,1972]" name="India" pageId="7" pageNumber="44">Hind</collectingCountry>
margin of sternite 7 truncate, middle slightly convex (
<figureCitation id="01C52A1BFE5BFFC5784EF86916B7DF89" captionStart="FIGURES 54 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="7" pageNumber="44">Fig. 55</figureCitation>
). Aedeagus (
<figureCitation id="01C52A1BFE5BFFC57C74F83217BCDF89" box="[342,434,1985,2011]" captionStart="FIGURES 44 47" captionStartId="10.[151,270,1823,1847]" captionTargetBox="[180,1399,210,1765]" captionTargetId="figure@10.[171,1415,197,1798]" captionTargetPageId="10" captionText="FIGURES 44 47. Male genitals in Laccodytes species: 44) L. apalodes: a) median lobe of aedeagus in lateral view, b c) right and left parameres; 45) L. rondonia: idem; 46) L. americanus: idem; 47) L. phalacroides: a) apical half of median lobe of aedeagus in dorsal view, b) median lobe in lateral view, c d) right and left parameres;" httpUri="https://zenodo.org/record/193413/files/figure.png" pageId="7" pageNumber="44">Fig. 45</figureCitation>
): median lobe similar to the preceding species, but larger, shorter and more robust, straight; the widened central portion is more extended and a rounded membraneous process emerges. Parameres different in size, both with more than one apical very long seta.
</paragraph>
<paragraph id="9941369EFE54FFCA7DE4FF1617D5D974" blockId="8.[151,1437,152,410]" pageId="8" pageNumber="45">
Female. Sternite 7 (
<figureCitation id="01C52A1BFE54FFCA7C98FF16141AD8AD" box="[442,532,229,255]" captionStart="FIGURES 54 61" captionStartId="12.[151,273,1107,1131]" captionTargetBox="[153,1431,196,1081]" captionTargetId="figure@12.[151,1436,194,1083]" captionTargetPageId="12" captionText="FIGURES 54 61. Last abdominal segment (left male; right female) in Laccodytes species: 54) L. apalodes; 55) L. rondonia; 56) L. americanus; 57) L. phalacroides; 58) L. takutuanus; 59) L. neblinae; 60) L. bassignanii; 61) Laccodytes sp. 1 female." httpUri="https://zenodo.org/record/193415/files/figure.png" pageId="8" pageNumber="45">Fig. 55</figureCitation>
) rounded on both sides, hind margin with a small and shallow V-shaped emargination in the middle.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE54FFCA7DE4FEC115BDD91E" box="[198,947,306,332]" pageId="8" pageNumber="45" type="distribution">
<paragraph id="9941369EFE54FFCA7DE4FEC115BDD91E" blockId="8.[151,1437,152,410]" box="[198,947,306,332]" pageId="8" pageNumber="45">
<emphasis id="AB8AEA8CFE54FFCA7DE4FEC11751D91E" bold="true" box="[198,351,306,332]" pageId="8" pageNumber="45">Distribution</emphasis>
(
<figureCitation id="01C52A1BFE54FFCA7C4DFEC117C9D91E" box="[367,455,306,332]" captionStart="FIGURE 62" captionStartId="18.[151,255,1941,1965]" captionText="FIGURE 62. Known distribution of Laccodytes species." pageId="8" pageNumber="45">Fig. 62</figureCitation>
). Southwestern
<collectingCountry id="E1E9760EFE54FFCA7FA0FEC114C3D91E" box="[642,717,306,332]" name="Brazil" pageId="8" pageNumber="45">Brazil</collectingCountry>
, State of
<taxonomicName id="5EFE4D1DFE54FFCA7E1BFEC115A1D91E" box="[825,943,306,332]" class="Insecta" family="Dytiscidae" genus="Laccodytes" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="57" phylum="Arthropoda" rank="species" species="rondonia">Rondônia</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE54FFCA7DE4FEAB1402D921" box="[198,524,344,371]" pageId="8" pageNumber="45" type="biology_ecology">
<paragraph id="9941369EFE54FFCA7DE4FEAB1402D921" blockId="8.[151,1437,152,410]" box="[198,524,344,371]" pageId="8" pageNumber="45">
<emphasis id="AB8AEA8CFE54FFCA7DE4FEAB1724D920" bold="true" box="[198,298,344,370]" pageId="8" pageNumber="45">Biology.</emphasis>
Found in a stream.
</paragraph>
</subSubSection>
<subSubSection id="D1E46515FE54FFCA7DE4FE8C14F3D9C8" box="[198,765,383,410]" pageId="8" pageNumber="45" type="etymology">
<paragraph id="9941369EFE54FFCA7DE4FE8C14F3D9C8" blockId="8.[151,1437,152,410]" box="[198,765,383,410]" pageId="8" pageNumber="45">
<emphasis id="AB8AEA8CFE54FFCA7DE4FE8C17A3D9CB" bold="true" box="[198,429,383,409]" pageId="8" pageNumber="45">Derivatio nominis.</emphasis>
Named for the
<typeStatus id="4645883CFE54FFCA7F47FE731496D9C8" box="[613,664,384,410]" pageId="8" pageNumber="45">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>