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<document id="524AD20EBD717481291DA8C3745533A6" ID-CLB-Dataset="3952" ID-DOI="10.5281/zenodo.1196048" ID-GBIF-Dataset="fed4a06f-7511-4cb0-82de-218f7a6dd132" ID-ISSN="0035-418" ID-Zenodo-Dep="1196048" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1634783961719" checkinUser="plazi" docAuthor="Kurbatov, Sergey A., Cuccodoro, Giulio &amp; Sabella, Giorgio" docDate="2018" docId="A7582020FF8CFFF3FEF3FA8A52F8EFA2" docLanguage="en" docName="13_Kurbatov_etal.pdf" docOrigin="Revue suisse de Zoologie 125 (1)" docStyle="DocumentStyle:225056F0A23C51DDC06AC27EAEA15D5B.2:RevSuisseZool.2015-2019.journal_article" docStyleId="225056F0A23C51DDC06AC27EAEA15D5B" docStyleName="RevSuisseZool.2015-2019.journal_article" docStyleVersion="2" docTitle="Achilia caracolana Jeannel 1962" docType="treatment" docVersion="7" lastPageNumber="176" masterDocId="5B615858FF87FFF8FFFFFFAC5639E96C" masterDocTitle="A revision of the Chilean Brachyglutini – Part 3. Revision of Achilia Reitter, 1890: A. frontalis species group (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="188" masterPageNumber="165" pageNumber="176" updateTime="1699464010858" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="D904F4164E2685B16E28EC4982A2F483" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="10B9F56ACCC84002B93E50651093DB6C">
<mods:title id="B188CA67607EBBE0A602206AA8E7002F">A revision of the Chilean Brachyglutini – Part 3. Revision of Achilia Reitter, 1890: A. frontalis species group (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
</mods:titleInfo>
<mods:name id="94EBFA9F2E9BF777921D2B7123299E13" type="personal">
<mods:role id="6C3B116A6C9C21ECB6453FE13BBBA7E3">
<mods:roleTerm id="6461A0AA8BE0DED806C127164E1FD90B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="17F08ADEF733CD84CEE4CDD9D8B16022">Kurbatov, Sergey A.</mods:namePart>
<mods:affiliation id="573D12DE0A69AAF9450B639F6534BE59">Museum of Entomology, All-Russian Plant Quarantine Center, Pogranichnaya 32, Bykovo 140150, Russia. E-mail:</mods:affiliation>
</mods:name>
<mods:name id="FC25E0D1E4B298E7A70BBFD2ADFA9EA4" type="personal">
<mods:role id="3587C547766C6EEEC92F49A05B8536A2">
<mods:roleTerm id="74BE8482B3887EDD6427F3FA82E43A94">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6D915D5423D4DFDBD88A9CCC21318CE2">Cuccodoro, Giulio</mods:namePart>
<mods:affiliation id="675D501D22BC27A390CF43A92CE89903">Muséum d’histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland.</mods:affiliation>
<mods:nameIdentifier id="748763DA0053743BA4DE19507A09B8D3" type="email">giulio.cuccodoro@ville-ge.ch</mods:nameIdentifier>
</mods:name>
<mods:name id="03604843DDBBCBCEE4975EE99A361A8F" type="personal">
<mods:role id="A21BBD6C8F37239EF8F968C7163D5D17">
<mods:roleTerm id="BD801A700678D221DDC1DE14DAC67F39">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="98602607C806CBD68FB37BC17610CE03">Sabella, Giorgio</mods:namePart>
<mods:nameIdentifier id="B81D352BB6C2CD546DD43686D0F0357A" type="ORCID">0000-0002-2234-1901</mods:nameIdentifier>
<mods:affiliation id="88B72D0D10DE809E765A94E2618FB62E">Dipartimento di Scienze Biologiche, Geologiche ed Ambientali dell’Università - sezione Biologia Animale, via</mods:affiliation>
</mods:name>
<mods:typeOfResource id="09249712D1E6BED33A157F617282C980">text</mods:typeOfResource>
<mods:relatedItem id="E6F86E9BCD944AAC1B82928FFD4D2DA7" type="host">
<mods:titleInfo id="8E9FB1B9BF98C36E30EB336AC360E9A8">
<mods:title id="5BA8072F3C9C1B3EEB304434C52358EC">Revue suisse de Zoologie</mods:title>
</mods:titleInfo>
<mods:part id="42E4FD52E9158EFE751F5CF4F2A5B523">
<mods:date id="06607EA29AB6F91D9976D817F5F5E48A">2018</mods:date>
<mods:detail id="41701A3D11955C03E001D8E4434949DB" type="pubDate">
<mods:number id="6159A779C658AC0B7C63AECC2F74F276">2018-03-31</mods:number>
</mods:detail>
<mods:detail id="ED5B9D57E2F7342A9B789DC4A949C682" type="volume">
<mods:number id="57122F86DBE66BB0966B212C56F14E1C">125</mods:number>
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<mods:number id="5B24DEDC83A356EE322005F10426FC19">1</mods:number>
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<mods:extent id="DA288E026B34C288CA21CBD421081821" unit="page">
<mods:start id="9734C222EA6FB8B8305803D6010F7AB8">165</mods:start>
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<mods:classification id="7D0EBD299E475F4F69BCFD58A5BE4812">journal article</mods:classification>
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<mods:identifier id="4EDD42E4C7818AFFC4AB2306193782F0" type="DOI">10.5281/zenodo.1196048</mods:identifier>
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<mods:identifier id="6E8F2FAD77DC3945E0967960C051BB59" type="ISSN">0035-418</mods:identifier>
<mods:identifier id="0967E4713ECF7276083212D261734A13" type="Zenodo-Dep">1196048</mods:identifier>
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<treatment id="A7582020FF8CFFF3FEF3FA8A52F8EFA2" ID-DOI="http://doi.org/10.5281/zenodo.5589388" ID-GBIF-Taxon="189149714" ID-Zenodo-Dep="5589388" LSID="urn:lsid:plazi:treatment:A7582020FF8CFFF3FEF3FA8A52F8EFA2" httpUri="http://treatment.plazi.org/id/A7582020FF8CFFF3FEF3FA8A52F8EFA2" lastPageNumber="176" pageId="11" pageNumber="176">
<subSubSection id="67EBC2BDFF8CFFF3FEF3FA8A5447EC52" box="[268,638,1318,1342]" pageId="11" pageNumber="176" type="nomenclature">
<paragraph id="2F4E9136FF8CFFF3FEF3FA8A5447EC52" blockId="11.[234,674,1318,1375]" box="[268,638,1318,1342]" pageId="11" pageNumber="176">
<heading id="7406265AFF8CFFF3FEF3FA8A5447EC52" bold="true" box="[268,638,1318,1342]" centered="true" level="2" pageId="11" pageNumber="176" reason="2">
<taxonomicName id="E8F1EAB5FF8CFFF3FEF3FA8A5447EC52" ID-CoL="9778" authority="Jeannel, 1962" authorityName="Jeannel" authorityYear="1962" box="[268,638,1318,1342]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="176" phylum="Arthropoda" rank="species" species="caracolana">
<emphasis id="1D854D24FF8CFFF3FEF3FA8A5447EC52" bold="true" box="[268,638,1318,1342]" pageId="11" pageNumber="176">
<emphasis id="1D854D24FF8CFFF3FEF3FA8A57E7EC52" bold="true" box="[268,478,1318,1342]" italics="true" pageId="11" pageNumber="176">Achilia caracolana</emphasis>
<bibRefCitation id="4B60ECC7FF8CFFF3FE1AFA8B5447EC52" author="Jeannel R." box="[485,638,1319,1342]" pageId="11" pageNumber="176" pagination="295 - 479" refId="ref11488" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D. &amp; Rapoport E. (eds.), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF8CFFF3FF15FAE4549BEC33" box="[234,674,1352,1375]" pageId="11" pageNumber="176" type="description">
<paragraph id="2F4E9136FF8CFFF3FF15FAE4549BEC33" blockId="11.[234,674,1318,1375]" box="[234,674,1352,1375]" pageId="11" pageNumber="176">
<figureCitation id="B7CA8DB3FF8CFFF3FF15FAE45705EC33" box="[234,316,1352,1375]" captionStart="Figs 24-29" captionStartId="4.[151,192,1401,1422]" captionTargetBox="[216,1403,263,1340]" captionTargetId="figure-245@4.[162,1413,254,1361]" captionTargetPageId="4" captionText="Figs 24-29. Aedeagi of Achilia. (24) A.caracolana, specimen from Hualpèn, prov. Concepción. (25) A. auriculata, specimen from Peniquillo, prov. Concepción. (26) A. ovallensis, specimen from Ovalle, prov. Limarí [paratype]. (27) A. frontalis, specimen from Aguas Calientes to Puyehue National Park, prov. Osorno. (28) A. trauco, specimen from 34 km WNW La Union, prov. Ranco [paratype]. (29) A. longispina, specimen from Mina de la Disputada, prov. Valparaíso." figureDoi="http://doi.org/10.5281/zenodo.5589962" httpUri="https://zenodo.org/record/5589962/files/figure.png" pageId="11" pageNumber="176">Figs 24</figureCitation>
,
<figureCitation id="B7CA8DB3FF8CFFF3FEB8FAE4575DEC33" box="[327,356,1352,1375]" captionStart="Figs 30-34" captionStartId="5.[151,192,1388,1409]" captionTargetBox="[189,1397,257,1335]" captionTargetId="figure-284@5.[148,1409,254,1359]" captionTargetPageId="5" captionText="Figs 30-34. Male antennae of Achilia. (30) A. caracolana. (31) A. auriculata. (32) A. ovallensis. (33) A. frontalis. (34) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589966" httpUri="https://zenodo.org/record/5589966/files/figure.png" pageId="11" pageNumber="176">30</figureCitation>
,
<figureCitation id="B7CA8DB3FF8CFFF3FE90FAE4578DEC33" box="[367,436,1352,1375]" captionStart="Figs 35-43" captionStartId="6.[151,192,1343,1364]" captionTargetBox="[158,1412,284,1297]" captionTargetId="figure-278@6.[151,1422,257,1320]" captionTargetPageId="6" captionText="Figs 35-43. Male profemur (35), metafemur (36), the variability of metafemur (38-40), metaleg (37) and mesotrochanter (41-43) of Achilia. (35, 41) A. caracolana. (36) A. trauco. (37) A. frontalis, specimen from Chillàn. (38) A. frontalis, specimen from Cordillera de Nahuelbuta. (39) A. frontalis, specimen from Frutillar. (40) A. frontalis, specimen from Aguas Calientes to Puyehue National Park. (42) A. ovallensis [paratype]. (43) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589968" httpUri="https://zenodo.org/record/5589968/files/figure.png" pageId="11" pageNumber="176">35, 41</figureCitation>
,
<figureCitation id="B7CA8DB3FF8CFFF3FE40FAE4543DEC33" box="[447,516,1352,1375]" captionStart="Figs 44-52" captionStartId="7.[151,192,997,1018]" captionTargetBox="[163,1419,254,945]" captionTargetId="figure-488@7.[151,1431,248,968]" captionTargetPageId="7" captionText="Figs 44-52. Male protibia (44-46) and mesotibia (47-52) of Achilia. (44, 47) A. caracolana. (45, 50) A. frontalis. (46, 51) A. trauco. (48) A. ovallensis. (49) A. auriculata. (52) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589970" httpUri="https://zenodo.org/record/5589970/files/figure.png" pageId="11" pageNumber="176">44, 47</figureCitation>
, 65, 67, 69, 84
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF8CFFF3FF68FAD954F6ECE7" box="[151,719,1397,1419]" pageId="11" pageNumber="176" type="reference_group">
<paragraph id="2F4E9136FF8CFFF3FF68FAD954F6ECE7" blockId="11.[151,719,1397,1419]" box="[151,719,1397,1419]" pageId="11" pageNumber="176">
<treatmentCitationGroup id="0FE1B618FF8CFFF3FF68FAD954F6ECE7" box="[151,719,1397,1419]" pageId="11" pageNumber="176">
<treatmentCitation id="AE50B727FF8CFFF3FF68FAD95433ECE7" author="Jeannel R." box="[151,522,1397,1419]" page="409" pageId="11" pageNumber="176" year="1962">
<taxonomicName id="E8F1EAB5FF8CFFF3FF68FAD95433ECE7" ID-CoL="9778" authority="Jeannel, 1962: 409" authorityName="Jeannel" authorityPageNumber="409" authorityYear="1962" box="[151,522,1397,1419]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="176" phylum="Arthropoda" rank="species" species="caracolana">
<emphasis id="1D854D24FF8CFFF3FF68FAD95769ECE6" box="[151,336,1397,1418]" italics="true" pageId="11" pageNumber="176">Achilia caracolana</emphasis>
<bibRefCitation id="4B60ECC7FF8CFFF3FEA9FADA5433ECE7" author="Jeannel R." box="[342,522,1398,1419]" pageId="11" pageNumber="176" pagination="295 - 479" refId="ref11488" refString="Jeannel R. 1962. Les Pselaphides de la Paleantarctide Occidentale [pp. 295 - 479]. In: Deboutteville C. D. &amp; Rapoport E. (eds.), Biologie de l'Amerique Australe. Vol. I. Etude sur la Faune du Sol. Centre National de la Recherche Scientifique, Paris." type="book chapter" year="1962">Jeannel, 1962: 409</bibRefCitation>
</taxonomicName>
</treatmentCitation>
fig. 163 (aedeagus).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF8CFFF3FF68FA0F54A7EF06" pageId="11" pageNumber="176" type="materials_examined">
<paragraph id="2F4E9136FF8CFFF3FF68FA0F5466EF57" blockId="11.[151,756,1443,1595]" pageId="11" pageNumber="176">
<emphasis id="1D854D24FF8CFFF3FF68FA0F579EECD6" bold="true" box="[151,423,1443,1466]" pageId="11" pageNumber="176">
Type material (
<specimenCount id="39F75ABFFF8CFFF3FEA8FA0F57A0ECD6" box="[343,409,1443,1466]" pageId="11" pageNumber="176" type="generic">7 ex.</specimenCount>
):
</emphasis>
<materialsCitation id="9F999B6BFF8CFFF3FE49FA0F57A9EF77" ID-GBIF-Occurrence="3391021309" collectingDate="1957-05-23" collectingDateMax="1957-05-24" collectingDateMin="1957-05-23" collectionCode="MHNS, MNHN" collectorName="G. Kuschel" country="Chile" county="Cerro Caracol" latitude="-36.833332" location="Region Bio Bio" municipality="Concepcion prov." pageId="11" pageNumber="176" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="holotype">
CENTRAL
<collectingCountry id="57E6D1A6FF8CFFF3FDBFFA0F54ADECD6" box="[576,660,1443,1466]" name="Chile" pageId="11" pageNumber="176">CHILE</collectingCountry>
:
<location id="2A2EC7EDFF8CFFF3FD59FA0F56CDECB6" LSID="urn:lsid:plazi:treatment:A7582020FF8CFFF3FEF3FA8A52F8EFA2:2A2EC7EDFF8CFFF3FD59FA0F56CDECB6" country="Chile" county="Cerro Caracol" latitude="-36.833332" municipality="Concepcion prov." name="Region Bio Bio" pageId="11" pageNumber="176">Región Bío Bío</location>
:
<collectingMunicipality id="CF2A0B4CFF8CFFF3FEFDFA6F57F1ECB6" box="[258,456,1475,1498]" pageId="11" pageNumber="176">Concepción prov.</collectingMunicipality>
:
<collectionCode id="49E009F3FF8CFFF3FE2AFA6F541FECB6" box="[469,550,1475,1498]" pageId="11" pageNumber="176">MHNS</collectionCode>
;
<specimenCount id="39F75ABFFF8CFFF3FDCAFA6F5458ECB6" box="[565,609,1475,1498]" pageId="11" pageNumber="176" type="male">1 ♂</specimenCount>
(
<typeStatus id="F04A2F94FF8CFFF3FD8CFA6F54EAECB6" box="[627,723,1475,1498]" pageId="11" pageNumber="176" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E8F1EAB5FF8CFFF3FF68FA4F575CEC97" authorityName="Jeannel" authorityYear="1962" box="[151,357,1507,1531]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="176" phylum="Arthropoda" rank="species" species="caracolana">
<emphasis id="1D854D24FF8CFFF3FF68FA4F575CEC97" box="[151,357,1507,1531]" italics="true" pageId="11" pageNumber="176">Achilia caracolana</emphasis>
</taxonomicName>
n° 1685);
<collectingCounty id="C62FE9BAFF8CFFF3FE26FA48544DEC97" box="[473,628,1508,1531]" pageId="11" pageNumber="176">Cerro Caracol</collectingCounty>
;
<date id="5B4FB7F6FF8CFFF3FD80FA4854C9EC97" box="[639,752,1508,1531]" pageId="11" pageNumber="176" value="1957-05-24">
<collectingDate id="4B0B4E1EFF8CFFF3FD80FA4854C9EC97" box="[639,752,1508,1531]" pageId="11" pageNumber="176" value="1957-05-24">24.V.1957</collectingDate>
</date>
;
<collectorName id="8204F4E0FF8CFFF3FF68F9A85720EF77" box="[151,281,1540,1563]" pageId="11" pageNumber="176">G. Kuschel.</collectorName>
–
<collectionCode id="49E009F3FF8CFFF3FEC8F9A857B5EF77" box="[311,396,1540,1563]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="176">MNHN</collectionCode>
;
</materialsCitation>
<materialsCitation id="9F999B6BFF8CFFF3FE66F9A85466EF57" ID-GBIF-Occurrence="3391021326" collectingDate="1957-05-23" collectingDateMax="1957-05-24" collectingDateMin="1957-05-23" collectionCode="MHNS, MNHN" collectorName="G. Kuschel" country="Chile" county="Cerro Caracol" latitude="-36.833332" location="Region Bio Bio" municipality="Concepcion prov." pageId="11" pageNumber="176" specimenCount="7" specimenCount-female="4" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="39F75ABFFF8CFFF3FE66F9A857FAEF76" box="[409,451,1540,1563]" pageId="11" pageNumber="176" type="male">2 ♂</specimenCount>
and
<specimenCount id="39F75ABFFF8CFFF3FE03F9A8541FEF76" box="[508,550,1540,1563]" pageId="11" pageNumber="176" type="female">4 ♀</specimenCount>
(
<typeStatus id="F04A2F94FF8CFFF3FDC9F9A8549BEF77" box="[566,674,1540,1563]" pageId="11" pageNumber="176" type="paratype">paratypes</typeStatus>
);
<collectorName id="8204F4E0FF8CFFF3FD48F9A856D4EF57" pageId="11" pageNumber="176">Cerro Caracol</collectorName>
; 36° 50’S;
<date id="5B4FB7F6FF8CFFF3FE9AF98857ECEF57" box="[357,469,1572,1595]" pageId="11" pageNumber="176" value="1957-05-23">
<collectingDate id="4B0B4E1EFF8CFFF3FE9AF98857ECEF57" box="[357,469,1572,1595]" pageId="11" pageNumber="176" value="1957-05-23">23.V.1957</collectingDate>
</date>
;
<collectorName id="8204F4E0FF8CFFF3FE1FF9885462EF57" box="[480,603,1572,1595]" pageId="11" pageNumber="176">G. Kuschel</collectorName>
.
</materialsCitation>
</paragraph>
<paragraph id="2F4E9136FF8CFFF3FF68F9FF54A7EF06" blockId="11.[151,670,1619,1642]" box="[151,670,1619,1642]" pageId="11" pageNumber="176">
<heading id="7406265AFF8CFFF3FF68F9FF54A7EF06" box="[151,670,1619,1642]" fontSize="10" level="3" pageId="11" pageNumber="176" reason="3">
<emphasis id="1D854D24FF8CFFF3FF68F9FF57D1EF06" bold="true" box="[151,488,1619,1642]" pageId="11" pageNumber="176">
Additional material (
<specimenCount id="39F75ABFFF8CFFF3FE79F9FF57E3EF06" box="[390,474,1619,1642]" pageId="11" pageNumber="176" type="generic">318 ex.</specimenCount>
):
</emphasis>
See Appendix 1.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF8CFFF3FF68F92F524FEF31" pageId="11" pageNumber="176" type="description">
<paragraph id="2F4E9136FF8CFFF3FF68F92F579DEE17" blockId="11.[151,756,1667,2077]" pageId="11" pageNumber="176">
<emphasis id="1D854D24FF8CFFF3FF68F92F571CEFF6" bold="true" box="[151,293,1667,1690]" pageId="11" pageNumber="176">Description:</emphasis>
Body
<quantity id="E8093CD3FF8CFFF3FE88F92F57C0EFF6" box="[375,505,1667,1690]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="11" pageNumber="176" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5-1.8 mm</quantity>
long, entirely reddish or reddish brown, with palpi yellowish. Head with eyes longer than temples. Pronotum slightly wider than head, with maximal width on anterior half; median antebasal fovea smaller than lateral ones. First abdominal tergite with basal striae parallel, extending to about quarter of paratergal length, and separated at base by more than one-third of tergal width.
</paragraph>
<paragraph id="2F4E9136FF8CFFF3FF68F8285221EC26" blockId="11.[151,756,1667,2077]" lastBlockId="11.[831,1437,258,1451]" pageId="11" pageNumber="176">
<emphasis id="1D854D24FF8CFFF3FF68F82856F7EEF0" box="[151,206,1924,1948]" italics="true" pageId="11" pageNumber="176">Male</emphasis>
: Head as in
<figureCitation id="B7CA8DB3FF8CFFF3FEA3F829543BEEF0" box="[348,514,1925,1948]" captionStart="Figs 65-70" captionStartId="12.[151,192,1998,2019]" captionTargetBox="[151,1431,257,1958]" captionTargetId="figure-10@12.[151,1431,257,1958]" captionTargetPageId="12" captionText="Figs 65-70. (65, 67, 69) Achilia caracolana. (66, 68, 70) A. auriculata. Male head in (65-66) dorsal and (67-68) lateral views. (69-70) male supraocular area in lateral view. Scale bar left for (65, 67) and right for (66, 68)." figureDoi="http://doi.org/10.5281/zenodo.5589982" httpUri="https://zenodo.org/record/5589982/files/figure.png" pageId="11" pageNumber="176">Figs 65 and 67</figureCitation>
, with deep lateral pits surmounted on each side of eyes by small tooth apically truncated (
<figureCitation id="B7CA8DB3FF8CFFF3FEF1F869575AEEB0" box="[270,355,1989,2012]" captionStart="Figs 65-70" captionStartId="12.[151,192,1998,2019]" captionTargetBox="[151,1431,257,1958]" captionTargetId="figure-10@12.[151,1431,257,1958]" captionTargetPageId="12" captionText="Figs 65-70. (65, 67, 69) Achilia caracolana. (66, 68, 70) A. auriculata. Male head in (65-66) dorsal and (67-68) lateral views. (69-70) male supraocular area in lateral view. Scale bar left for (65, 67) and right for (66, 68)." figureDoi="http://doi.org/10.5281/zenodo.5589982" httpUri="https://zenodo.org/record/5589982/files/figure.png" pageId="11" pageNumber="176">Fig. 69</figureCitation>
); large frontal protuberance convex with distinct V-shaped median impression reaching anteriorly to vertexal sulcus. Antennae (
<figureCitation id="B7CA8DB3FF8CFFF3FDA2F7AA5488E171" box="[605,689,2054,2077]" captionStart="Figs 30-34" captionStartId="5.[151,192,1388,1409]" captionTargetBox="[189,1397,257,1335]" captionTargetId="figure-284@5.[148,1409,254,1359]" captionTargetPageId="5" captionText="Figs 30-34. Male antennae of Achilia. (30) A. caracolana. (31) A. auriculata. (32) A. ovallensis. (33) A. frontalis. (34) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589966" httpUri="https://zenodo.org/record/5589966/files/figure.png" pageId="11" pageNumber="176">Fig. 30</figureCitation>
) with scape and pedicel longer than wide; antennomere III as long as wide; antennomere IV slightly wider than long; antennomere V slightly longer than wide; antennomeres VI-VIII distinctly wider than long; antennomere IX transverse with protruding mesal margin pointed at middle and bearing two long setae; antennomere X barely longer than wide, wider and longer than IX, with mesal margin apically hollowed, basally projecting and bearing two long and large setae as well as a tuft of shorter setae; antennomere XI elongate, longer than VIII-X combined, its medial margin bearing long subbasal setae. Metasternum with deep and broad ovoidal median impression, posterior edges of the latter densely pubescent. Legs with ventral margin of mesotrochanters (
<figureCitation id="B7CA8DB3FF8CFFF3FBF9FD655264EB8C" box="[1030,1117,713,736]" captionStart="Figs 35-43" captionStartId="6.[151,192,1343,1364]" captionTargetBox="[158,1412,284,1297]" captionTargetId="figure-278@6.[151,1422,257,1320]" captionTargetPageId="6" captionText="Figs 35-43. Male profemur (35), metafemur (36), the variability of metafemur (38-40), metaleg (37) and mesotrochanter (41-43) of Achilia. (35, 41) A. caracolana. (36) A. trauco. (37) A. frontalis, specimen from Chillàn. (38) A. frontalis, specimen from Cordillera de Nahuelbuta. (39) A. frontalis, specimen from Frutillar. (40) A. frontalis, specimen from Aguas Calientes to Puyehue National Park. (42) A. ovallensis [paratype]. (43) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589968" httpUri="https://zenodo.org/record/5589968/files/figure.png" pageId="11" pageNumber="176">Fig. 41</figureCitation>
) bearing two short spines; profemora (
<figureCitation id="B7CA8DB3FF8CFFF3FC3AFD465222EA6D" box="[965,1051,746,769]" captionStart="Figs 35-43" captionStartId="6.[151,192,1343,1364]" captionTargetBox="[158,1412,284,1297]" captionTargetId="figure-278@6.[151,1422,257,1320]" captionTargetPageId="6" captionText="Figs 35-43. Male profemur (35), metafemur (36), the variability of metafemur (38-40), metaleg (37) and mesotrochanter (41-43) of Achilia. (35, 41) A. caracolana. (36) A. trauco. (37) A. frontalis, specimen from Chillàn. (38) A. frontalis, specimen from Cordillera de Nahuelbuta. (39) A. frontalis, specimen from Frutillar. (40) A. frontalis, specimen from Aguas Calientes to Puyehue National Park. (42) A. ovallensis [paratype]. (43) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589968" httpUri="https://zenodo.org/record/5589968/files/figure.png" pageId="11" pageNumber="176">Fig. 35</figureCitation>
) strongly thickened, with ventral margin finely denticulate and bearing short modified setae on basal third; mesofemora slightly thickened; protibiae (
<figureCitation id="B7CA8DB3FF8CFFF3FCB8FCE755A0EA0E" box="[839,921,843,866]" captionStart="Figs 44-52" captionStartId="7.[151,192,997,1018]" captionTargetBox="[163,1419,254,945]" captionTargetId="figure-488@7.[151,1431,248,968]" captionTargetPageId="7" captionText="Figs 44-52. Male protibia (44-46) and mesotibia (47-52) of Achilia. (44, 47) A. caracolana. (45, 50) A. frontalis. (46, 51) A. trauco. (48) A. ovallensis. (49) A. auriculata. (52) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589970" httpUri="https://zenodo.org/record/5589970/files/figure.png" pageId="11" pageNumber="176">Fig. 44</figureCitation>
) slightly thickened with mesal margin bearing small subapical spine; mesotibiae (
<figureCitation id="B7CA8DB3FF8CFFF3FB33FCC05319EAEF" box="[1228,1312,876,899]" captionStart="Figs 44-52" captionStartId="7.[151,192,997,1018]" captionTargetBox="[163,1419,254,945]" captionTargetId="figure-488@7.[151,1431,248,968]" captionTargetPageId="7" captionText="Figs 44-52. Male protibia (44-46) and mesotibia (47-52) of Achilia. (44, 47) A. caracolana. (45, 50) A. frontalis. (46, 51) A. trauco. (48) A. ovallensis. (49) A. auriculata. (52) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589970" httpUri="https://zenodo.org/record/5589970/files/figure.png" pageId="11" pageNumber="176">Fig. 47</figureCitation>
) thickened with mesal margin bulging at middle and bearing apically pubescent subbasal spur; metatibiae slightly sinuate on distal half. Abdominal tergites unmodified; first abdominal ventrites with large median impression, all other ventrites flattened at middle; ventrite I with short median carina extending from posterior margin to posterior edge of median impression. Aedeagus (
<figureCitation id="B7CA8DB3FF8CFFF3FCB8FBDC55A0EDEB" box="[839,921,1136,1159]" captionStart="Figs 24-29" captionStartId="4.[151,192,1401,1422]" captionTargetBox="[216,1403,263,1340]" captionTargetId="figure-245@4.[162,1413,254,1361]" captionTargetPageId="4" captionText="Figs 24-29. Aedeagi of Achilia. (24) A.caracolana, specimen from Hualpèn, prov. Concepción. (25) A. auriculata, specimen from Peniquillo, prov. Concepción. (26) A. ovallensis, specimen from Ovalle, prov. Limarí [paratype]. (27) A. frontalis, specimen from Aguas Calientes to Puyehue National Park, prov. Osorno. (28) A. trauco, specimen from 34 km WNW La Union, prov. Ranco [paratype]. (29) A. longispina, specimen from Mina de la Disputada, prov. Valparaíso." figureDoi="http://doi.org/10.5281/zenodo.5589962" httpUri="https://zenodo.org/record/5589962/files/figure.png" pageId="11" pageNumber="176">Fig. 24</figureCitation>
)
<quantity id="E8093CD3FF8CFFF3FC55FBDC5268EDEB" box="[938,1105,1136,1159]" metricMagnitude="-4" metricUnit="m" metricValue="3.125" metricValueMax="3.25" metricValueMin="3.0" pageId="11" pageNumber="176" unit="mm" value="0.3125" valueMax="0.325" valueMin="0.3">0.30-0.325 mm</quantity>
long; dorsal plate ovoid with dorsal longitudinal struts divergent; copulatory pieces consisting of pair of long medial sclerites recurved and strongly sclerotised at base, and apically bifid. Parameres wide with very long seta on poorly developed outer lobe; tips strongly recurved posteriorly and bearing thin and short subapical seta.
</paragraph>
<paragraph id="2F4E9136FF8CFFF3FCC0FAFF53A3ECC7" blockId="11.[831,1437,258,1451]" pageId="11" pageNumber="176">
<emphasis id="1D854D24FF8CFFF3FCC0FAFF55B6EC07" box="[831,911,1363,1387]" italics="true" pageId="11" pageNumber="176">Female</emphasis>
: Similar to male except: head, metasternum, abdominal ventrites, and legs unmodified; antennomeres IX and specially X shorter and less thickened than male.
</paragraph>
<paragraph id="2F4E9136FF8CFFF3FCC0FA68524FEF31" blockId="11.[831,1437,1476,1629]" pageId="11" pageNumber="176">
<emphasis id="1D854D24FF8CFFF3FCC0FA6855C0ECB7" bold="true" box="[831,1017,1476,1499]" pageId="11" pageNumber="176">Collecting data:</emphasis>
Collected from September to May in
<taxonomicName id="E8F1EAB5FF8CFFF3FCC0FA485582EC90" box="[831,955,1508,1532]" class="Magnoliopsida" family="Nothofagaceae" genus="Nothofagus" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="176" phylum="Tracheophyta" rank="genus">
<emphasis id="1D854D24FF8CFFF3FCC0FA485582EC90" box="[831,955,1508,1532]" italics="true" pageId="11" pageNumber="176">Nothofagus</emphasis>
</taxonomicName>
,
<taxonomicName id="E8F1EAB5FF8CFFF3FC33FA485202EC97" box="[972,1083,1508,1531]" class="Pinopsida" family="Cupressaceae" genus="Cupressus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="176" phylum="Tracheophyta" rank="genus">
<emphasis id="1D854D24FF8CFFF3FC33FA485202EC97" box="[972,1083,1508,1531]" italics="true" pageId="11" pageNumber="176">Cupressus</emphasis>
</taxonomicName>
and
<taxonomicName id="E8F1EAB5FF8CFFF3FB88FA4852D7EC90" box="[1143,1262,1508,1532]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="11" pageNumber="176" phylum="Tracheophyta" rank="genus">
<emphasis id="1D854D24FF8CFFF3FB88FA4852D7EC90" box="[1143,1262,1508,1532]" italics="true" pageId="11" pageNumber="176">Eucalyptus</emphasis>
</taxonomicName>
forests; also in remnants, where it was found from sea level to about
<quantity id="E8093CD3FF8CFFF3FCC0F98A55A2EF51" box="[831,923,1574,1597]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="176" unit="m" value="1200.0">1200 m</quantity>
. Most specimens were collected from sifted samples of leaf and log litter.
</paragraph>
</subSubSection>
<subSubSection id="67EBC2BDFF8CFFF3FCC0F9DA52F8EFA2" pageId="11" pageNumber="176" type="distribution">
<paragraph id="2F4E9136FF8CFFF3FCC0F9DA52F8EFA2" blockId="11.[831,1436,1653,1742]" pageId="11" pageNumber="176">
<emphasis id="1D854D24FF8CFFF3FCC0F9DA55EEEFE1" bold="true" box="[831,983,1654,1677]" pageId="11" pageNumber="176">Distribution:</emphasis>
<taxonomicName id="E8F1EAB5FF8CFFF3FC13F9D952F5EFE1" authorityName="Jeannel" authorityYear="1962" box="[1004,1228,1653,1677]" class="Insecta" family="Staphylinidae" genus="Achilia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="176" phylum="Arthropoda" rank="species" species="caracolana">
<emphasis id="1D854D24FF8CFFF3FC13F9D952F5EFE1" box="[1004,1228,1653,1677]" italics="true" pageId="11" pageNumber="176">Achilia caracolana</emphasis>
</taxonomicName>
is known from Central
<collectingCountry id="57E6D1A6FF8CFFF3FC65F93A55EEEFC1" box="[922,983,1686,1709]" name="Chile" pageId="11" pageNumber="176">Chile</collectingCountry>
, ranging from Chiloé Province to Talca Province (
<figureCitation id="B7CA8DB3FF8CFFF3FC52F91B55C4EFA2" box="[941,1021,1719,1742]" captionStart="Fig" captionStartId="17.[831,866,1968,1989]" captionTargetBox="[880,1355,257,1924]" captionTargetId="figure-12@17.[913,1355,257,1924]" captionTargetPageId="17" captionText="Fig. 84. Distribution map. (● red circles) Achilia testacea. (▲ blue triangles) A. frontalis. (◆ green diamonds) A. caracolana. (■ pink squares) A. trauco. (✷ yellow stars) A. longispina." figureDoi="http://doi.org/10.5281/zenodo.5589993" httpUri="https://zenodo.org/record/5589993/files/figure.png" pageId="11" pageNumber="176">Fig. 84</figureCitation>
: green diamonds).
</paragraph>
</subSubSection>
</treatment>
</document>
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