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<document ID-CLB-Dataset="108060" ID-DOI="http://dx.doi.org/10.3897/zookeys.1115.83266" ID-GBIF-Dataset="310d67f4-8b17-48e3-84e1-e7f6fd328045" ID-PMC="PMC9848874" ID-Pensoft-Pub="1313-2970-1115-1" ID-Pensoft-UUID="1AB765B8065B5504A31014D2B0937E01" ID-PubMed="36761071" ID-ZooBank="C3E233F10EF74D2DBD4AA32AE7C4DF5E" ModsDocID="1313-2970-1115-1" checkinTime="1659044149331" checkinUser="pensoft" docAuthor="Lee, Jimin, Chang, Cheon Young &amp; Kim, Il-Hoi" docDate="2022" docId="EDC2C5724DF35AF8BA49927F82CAB8A0" docLanguage="en" docName="ZooKeys 1115: 1-71" docOrigin="ZooKeys 1115" docPubDate="2022-07-28" docSource="http://dx.doi.org/10.3897/zookeys.1115.83266" docTitle="Acontiophorus estivalis Lee &amp; Chang &amp; Kim 2022, sp. nov." docType="treatment" docUuid="5654B945-0217-4D32-A0EB-2BF6D204BD4E" docUuidSource="ZooBank" docVersion="5" id="1AB765B8065B5504A31014D2B0937E01" lastPageNumber="1" masterDocId="1AB765B8065B5504A31014D2B0937E01" masterDocTitle="Symbiotic copepods (Cyclopoida and Siphonostomatoida) collected by light trap from Korea" masterLastPageNumber="71" masterPageNumber="1" pageNumber="1" updateTime="1732655501609" updateUser="ExternalLinkService">
<mods:mods id="6C85EDDFCAD95421642B1D3C08901FE9" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="DD3744F35C7FA171A32E565972D0F0C1">Symbiotic copepods (Cyclopoida and Siphonostomatoida) collected by light trap from Korea</mods:title>
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<mods:namePart id="36EB8D6F526F86C4EBBAE54CDF3FE5AD">Lee, Jimin</mods:namePart>
<mods:nameIdentifier id="2930B9EFFF07B35326566E12E2D65B85" type="ORCID">https://orcid.org/0000-0001-9004-8275</mods:nameIdentifier>
<mods:affiliation id="CE0028B5A20BE3BCEEB53D2833B63C5A">Marine Ecosystem and Biological Research Center, Korea Institute of Ocean Science &amp; Technology, Busan 49111, Republic of Korea</mods:affiliation>
</mods:name>
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<mods:namePart id="2BFF9133E0C07B86CEB2E34F40DDA847">Chang, Cheon Young</mods:namePart>
<mods:nameIdentifier id="27D40C6951B0DA7A8D6E5C67B0A9D5CB" type="ORCID">https://orcid.org/0000-0001-5557-7120</mods:nameIdentifier>
<mods:affiliation id="7953F273A26CC2B856ABC33E2CA992B1">Department of Biological Science, Daegu University, Gyeongsan 38453, Republic of Korea</mods:affiliation>
</mods:name>
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<mods:namePart id="453A320103156678B9CB9B0F9D01AF00">Kim, Il-Hoi</mods:namePart>
<mods:nameIdentifier id="36138139B16CEF532CC2BF79651293D8" type="ORCID">https://orcid.org/0000-0002-7332-0043</mods:nameIdentifier>
<mods:affiliation id="1F72239C03EA2EF813FC0EAFF43A39F6">Korea Institute of Coastal Ecology, 302 - 802, Seokcheon-ro 397, Bucheon 14449, Republic of Korea</mods:affiliation>
<mods:nameIdentifier id="3E184E47EF77D8B4D098505D7A101ECA" type="email">ihkim@gwnu.ac.kr</mods:nameIdentifier>
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<mods:date id="3ED0D00196308D97D07ECE3EFDA6E54D">2022</mods:date>
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<mods:number id="D2E7352DFB4850BC8BE78F0052CD550F">2022-07-28</mods:number>
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<mods:identifier id="64CD4DDE1F8BAFECA570DBBC64CDC8E9" type="DOI">http://dx.doi.org/10.3897/zookeys.1115.83266</mods:identifier>
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<treatment id="EDC2C5724DF35AF8BA49927F82CAB8A0" ID-GBIF-Taxon="197867680" LSID="urn:lsid:zoobank.org:act:5654B945-0217-4D32-A0EB-2BF6D204BD4E" httpUri="http://treatment.plazi.org/id/EDC2C5724DF35AF8BA49927F82CAB8A0" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Asterocheridae" scope_order="Cyclopoida">
<subSubSection id="0C2CD25B5EF3D07FA93D1630864414FA" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="8E5B33C9E85515311A1195F11F71166B" pageId="0" pageNumber="1">
<taxonomicName id="A577C546FB3F55D4F677081EDCEA89FC" ID-CoL="9GDKP" LSID="https://zoobank.org/5654B945-0217-4D32-A0EB-2BF6D204BD4E" authority="Lee &amp; Chang &amp; Kim, 2022" authorityName="Lee &amp; Chang &amp; Kim" authorityYear="2022" class="Copepoda" family="Asterocheridae" genus="Acontiophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acontiophorus estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis" status="sp. nov.">Acontiophorus estivalis</taxonomicName>
<taxonomicNameLabel id="3A1D0E6943736ACA6B0CA4A0E279D332" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="34694BA0DDC5B6CC622DD66FB5F04232" pageId="0" pageNumber="1" type="description">
<paragraph id="32357D0BD6D64D876B624CE7AE794A8F" pageId="0" pageNumber="1">
<figureCitation id="6D8D2E9DE8B93925DA3C8902A1403C66" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">Figs 25</figureCitation>
<figureCitation id="1BEFBF67694D831E6113B2136AC611FD" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Acontiophorus estivalis sp. nov., female A maxilla B maxilliped C leg 1 D leg 2 E leg 3 F leg 4 G right genital aperture, dorsal. Scale bars: 0.05 mm (A-F); 0.02 mm (G)." figureDoi="10.3897/zookeys.1115.83266.figure26" httpUri="https://binary.pensoft.net/fig/720746" pageId="0" pageNumber="1">, 26</figureCitation>
<figureCitation id="D73608B957E8CC730A7AD26606A97EC8" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Acontiophorus estivalis sp. nov., female A leg 5, dorsal. Male B habitus, dorsal C antennule (setae omitted) D leg 5, dorsal E leg 6. Scale bars: 0.02 mm (A, C-E); 0.1 mm (B)." figureDoi="10.3897/zookeys.1115.83266.figure27" httpUri="https://binary.pensoft.net/fig/720747" pageId="0" pageNumber="1">, 27</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="55E285048B89F8B54CC2F0140A7A7B95" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="1011689E2DCF8BEA8363E73AE34DC77B" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="CEC3F9F09305CEA2F3395376EA81E40B" pageId="0" pageNumber="1">
<emphasis id="F954C584C5B9612B5A0B85BFEDE5F288" bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus id="A9DEF6DD321265D8101BEA209EDBD2D7">Holotype</typeStatus>
</emphasis>
<specimenCount id="69F50A788EF933A3D704F849573797B6"></specimenCount>
(MABIK CR00250115) and
<emphasis id="4482DF4BDCB1AF67CB508D0A6D3A2DFC" bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus id="FD4E9B490AED801971251606032F57A5">paratypes</typeStatus>
</emphasis>
<specimenCount id="30CCCB0D11678FEC42212A51AD4E04D2" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="EC2C9A6869453CB108AA94D40808C9DF" type="male">2 ♂♂</specimenCount>
(MABIK CR00250116) preserved in 90% alcohol, and
<emphasis id="6AE9A16DCCC89C3257D350E37654D5F0" bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus id="92A99EDD567114E4C1F0F269C58AB5D8">paratypes</typeStatus>
</emphasis>
<specimenCount id="F3DD903CF684F547379153F2BEBDBCEA" type="female">1 ♀</specimenCount>
,
<specimenCount id="3C52CBDBF6B786EB8B6F9317ED6A56A9" type="male">1 ♂</specimenCount>
dissected and mounted on a slide, Site 11 (Yeongdo, Pusan,
<geoCoordinate id="46CFCF98E1D667C7DC4E2C8F80E2E1E6" degrees="35" direction="north" minutes="04" orientation="latitude" precision="1" seconds="31.0" value="35.075275">35°04'31.0&quot;N</geoCoordinate>
,
<geoCoordinate id="727455FB6D92734724BCFA166F8254D6" degrees="129" direction="east" minutes="05" orientation="longitude" precision="1" seconds="08.7" value="129.08574">129°05'08.7&quot;E</geoCoordinate>
),
<date id="439463B553A376792A620BEC0A74B3FD" value="2020-07-07">07 Jul. 2020</date>
, leg. J. G. Kim. Dissected
<typeStatus id="AA5FF1292B18558232A25C394C6F508C">paratypes</typeStatus>
(
<specimenCount id="33F20A63292824FFAB378D40D006572D" type="female">1 ♀</specimenCount>
,
<specimenCount id="914792DA7F84F12908442FA325727AC3" type="male">1 ♂</specimenCount>
) are retained in the collection of I.-H. Kim.
</paragraph>
</subSubSection>
<subSubSection id="055EB6D62D21BA9B72B9AE331BD55D2B" pageId="0" pageNumber="1" type="description">
<paragraph id="360E872EC5848CCC13468FA20C33E107" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="67C6FF41811CEBCCB559F4CC35ED48EA" pageId="0" pageNumber="1">
<emphasis id="D308E8166D29133A3D1287B9C9B16077" bold="true" pageId="0" pageNumber="1">Female.</emphasis>
Body (Fig.
<figureCitation id="8934352E7A784FFC354F9A9E45A24929" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25A</figureCitation>
) stout, 938
<normalizedToken id="995F573C6B7B0BAAE62AF19F76F7A5EF" originalValue="μm">μm</normalizedToken>
long in dissected and figured paratype (length range 893-945
<normalizedToken id="6FE4046D84A4FF298274B7C74FF1E860" originalValue="μm">μm</normalizedToken>
, holotype 945
<normalizedToken id="81E7FA5321599FA953C7C3649C76A9A2" originalValue="μm">μm</normalizedToken>
). Prosome 625
<normalizedToken id="99294D3308AE0EB87A79D7F20529A4A1" originalValue="×">x</normalizedToken>
425
<normalizedToken id="9509E354EA80C7C827F811B86D5AD038" originalValue="μm">μm</normalizedToken>
, occupying 67% of body length, consisting of cephalothorax and second to fourth pedigerous somites. Cephalothorax 425
<normalizedToken id="EBED8B2CD10C39A1F637412410E8B10B" originalValue="μm">μm</normalizedToken>
long, as long as wide, without any dorsal suture line delimiting cephalosome and first pedigerous somite. Cephalothorax and second to third pedigerous somites with membranous fringe along posterodorsal margin. Fourth pedigerous somite with deeply concave posterior margin. Urosome (Fig.
<figureCitation id="C66268D0DA99DB11CB43FA3C0CA49BC9" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25B</figureCitation>
) four-segmented. Fifth pedigerous somite 133
<normalizedToken id="ADFCC2268E28447F9F86D122BF98BC02" originalValue="μm">μm</normalizedToken>
wide, with round lateral margins. Genital double-somite wider than long (115
<normalizedToken id="F101F32613CF97AB5AE8F94E2EB066DA" originalValue="×">x</normalizedToken>
132
<normalizedToken id="E79DE1532BEBB890DF207E22E1E10777" originalValue="μm">μm</normalizedToken>
), consisting of laterally expanded anterior third and narrower posterior two-thirds, with pointed posterolateral corners; genital apertures positioned dorsolaterally at expanded anterior region. Two free abdominal somites 39
<normalizedToken id="482D48D8CF95F4A054F5AA6B7CD44DD8" originalValue="×">x</normalizedToken>
77
<normalizedToken id="12047C9A72379E854F767EAB66F7F7E5" originalValue="μm">μm</normalizedToken>
and 45
<normalizedToken id="31D7ED1AF150BE7D373F4C418B48596B" originalValue="×">x</normalizedToken>
75
<normalizedToken id="008412087DB3CCD673F7136BA641760F" originalValue="μm">μm</normalizedToken>
. Anal somite (Fig.
<figureCitation id="C36607C4641BB12DFD0A85223327D2D7" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25C</figureCitation>
) ornamented on ventral surface with two groups of several large setules on medial region, scattered fine setules on lateral regions, and several spinules at medial posterior margin near bases of caudal rami. Caudal ramus (Fig.
<figureCitation id="31162BD730C584315462D50EB176A184" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25C</figureCitation>
) rectangular, 2.12
<normalizedToken id="1992252BBF9424DD0597047F0436B606" originalValue="×">x</normalizedToken>
longer than wide (70
<normalizedToken id="910B7AB2AAB19DD02D3375F5CAA3B1C7" originalValue="×">x</normalizedToken>
33
<normalizedToken id="22940812F4691E8E3B4B7A20E75D2C23" originalValue="μm">μm</normalizedToken>
), armed with six setae plus one aesthetasc-like element (indicated by arrowhead in Fig.
<figureCitation id="C843F9F954F5BACAAD1F10A55BE9E631" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25C</figureCitation>
), and ornamented with fine setules on ventral surface and four transverse rows of minute spinules on inner margin; dorsal setae (setae VI and VII) naked, other setae pinnate; outer lateral seta (seta II) with long setules along outer margin but spinulose (or with short setules) along inner margin; seta VI inserted on prolongation of ramus.
</paragraph>
<caption id="6DC6F29BAEBD997B0950816691D97EE7" doi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1" start="Figure 25" startId="F25">
<paragraph id="A3F1DDE783A6F1B4350EAD3AF9851A91" pageId="0" pageNumber="1">
<emphasis id="BEC7EBF837A01AB793710A9B9334C0F7" bold="true" pageId="0" pageNumber="1">Figure 25.</emphasis>
<taxonomicName id="471066C740EAD4CD6C58264DFE3B3784" authorityName="Lee &amp; Chang &amp; Kim" authorityYear="2022" class="Copepoda" family="Asterocheridae" genus="Acontiophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acontiophorus estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="78C22B78103A621A0EB13103C27B8AAC" italics="true" pageId="0" pageNumber="1">Acontiophorus estivalis</emphasis>
</taxonomicName>
sp. nov., female
<emphasis id="7D89C064F4F7520634E07D03DC7E48E4" bold="true" pageId="0" pageNumber="1">A</emphasis>
habitus, dorsal
<emphasis id="8545D4C417FD25A30C3CB622458BC8F7" bold="true" pageId="0" pageNumber="1">B</emphasis>
urosome, dorsal
<emphasis id="90A60E49B0BB93E4C9D7A0FA6B00BA58" bold="true" pageId="0" pageNumber="1">C</emphasis>
anal somite and caudal rami, ventral
<emphasis id="291A5E4B8C30E9FC8EEF10B67E723BD8" bold="true" pageId="0" pageNumber="1">D</emphasis>
antennule
<emphasis id="59AB47E6026105396C9CE41B32F5872E" bold="true" pageId="0" pageNumber="1">E</emphasis>
antenna
<emphasis id="172357685178D377F144E73FA06954DD" bold="true" pageId="0" pageNumber="1">F</emphasis>
oral siphon
<emphasis id="487816B36D1B06F5EB2821DA7B6689AB" bold="true" pageId="0" pageNumber="1">G</emphasis>
mandible
<emphasis id="7C9EA221F8CB4EB0DD34334795CD4547" bold="true" pageId="0" pageNumber="1">H</emphasis>
inner lobe of maxillule
<emphasis id="1D223693509FBB7549ADA6854C5CF511" bold="true" pageId="0" pageNumber="1">I</emphasis>
outer lobe of maxillule. Scale bars: 0.1 mm (
<emphasis id="B733AEAFF7B3F94E246D59DCFDC9C131" bold="true" pageId="0" pageNumber="1">A</emphasis>
); 0.05 mm (
<emphasis id="C6F7EA848009ABF374B72F01E848855B" bold="true" pageId="0" pageNumber="1">B, C, F, G</emphasis>
); 0.02 mm (
<emphasis id="4B4E8A1203D28C6F6CE839CBD27C89F1" bold="true" pageId="0" pageNumber="1">D, E, H, I</emphasis>
).
</paragraph>
</caption>
<paragraph id="4A6738629440DD2004778CD9B5335D74" pageId="0" pageNumber="1">
Rostrum absent. Antennule (Fig.
<figureCitation id="59D00240966B68B96F2394C03371815D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25D</figureCitation>
) short, 147
<normalizedToken id="8DE7D7FAE19F102397E100AB8899156F" originalValue="μm">μm</normalizedToken>
long, 11-segmented; armature formula 2, 14, 4, 2, 2, 8, 2, 1+aesthetasc, 2, 4, and 7; aesthetasc on 8th segment large; setae densely arranged, difficult to distinguish from one another. Antenna (Fig.
<figureCitation id="2ECDD8B7E2E187F9A22CAB9F0EB018DC" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25E</figureCitation>
) consisting of coxa, basis, one-segmented exopod, and two-segmented endopod; coxa short, unarmed; basis longest segment, narrowed in mid-region, with tuft of long setules at inner distal corner; exopod elongate, 6.0
<normalizedToken id="F70ABF0F2EA14A619EDF1930912D5D9E" originalValue="×">x</normalizedToken>
longer than wide (54
<normalizedToken id="CAEEA26B2784752EF9877D90D49A6E81" originalValue="×">x</normalizedToken>
9
<normalizedToken id="5B530620A3AEE02FB3EE03DA06A96291" originalValue="μm">μm</normalizedToken>
), extending to middle of second endopodal segment, armed with one small seta in middle, one minute seta subdistally, and one large, unilaterally pinnate seta (97
<normalizedToken id="1F0E67297B7581FC6BCEF8DF32BC6FF5" originalValue="μm">μm</normalizedToken>
long) distally; first endopodal segment unarmed, 32
<normalizedToken id="4622199A3B06270C9C24ADD23F108A9E" originalValue="×">x</normalizedToken>
22
<normalizedToken id="1B3C00F34A18D7F2376CB47F789CF3B2" originalValue="μm">μm</normalizedToken>
; second endopodal segment 2.2
<normalizedToken id="7E7F7DDB7BFF25B178665F0D644447AD" originalValue="×">x</normalizedToken>
longer than wide (40
<normalizedToken id="9FC759BF002B5A320FC219461A8A9FDC" originalValue="×">x</normalizedToken>
18
<normalizedToken id="00AE4E06F737821F9B191AD542FC8721" originalValue="μm">μm</normalizedToken>
), armed with six setae consisting of one large proximal seta, three unequal subdistal setae (one minute, setule-like), and two broad apical setae 75 and 43
<normalizedToken id="B11601E18502E4BAA00FC3115CFCD52A" originalValue="μm">μm</normalizedToken>
long, and ornamented with several rows of fine spinules or setules.
</paragraph>
<paragraph id="36346CE1A0EC992E7700302EED4F6B12" pageId="0" pageNumber="1">
Oral siphon (Fig.
<figureCitation id="351C07AC42ABEBCCE0B8F4A5B994D5C5" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25F</figureCitation>
) consisting of conical proximal part (maximum width 67
<normalizedToken id="AAFAFEDF147CC56D1C8BA82463B3FE81" originalValue="μm">μm</normalizedToken>
) and thin distal part, extending to middle of genital double-somite. Mandible (Fig.
<figureCitation id="FE0ADDA7BB7E61A318474B1597CFC0A6" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25G</figureCitation>
) consisting of thread-like stylet and palp; palp short, tapering, armed with one large, heavily pinnate seta and one minute, setule-like seta distally. Maxillule (Fig.
<figureCitation id="5878A4010671EE73994D4E09F26C3CA1" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Acontiophorus estivalis sp. nov., female A habitus, dorsal B urosome, dorsal C anal somite and caudal rami, ventral D antennule E antenna F oral siphon G mandible H inner lobe of maxillule I outer lobe of maxillule. Scale bars: 0.1 mm (A); 0.05 mm (B, C, F, G); 0.02 mm (D, E, H, I)." figureDoi="10.3897/zookeys.1115.83266.figure25" httpUri="https://binary.pensoft.net/fig/720745" pageId="0" pageNumber="1">25H, I</figureCitation>
) bilobed; larger inner lobe armed with four large (two feebly pinnate and two plumose) and one small setae distally; smaller inner lobe armed with three pinnate and one small, naked setae. Maxilla (Fig.
<figureCitation id="52E1D9335698ADFD47CA459263023DD4" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Acontiophorus estivalis sp. nov., female A maxilla B maxilliped C leg 1 D leg 2 E leg 3 F leg 4 G right genital aperture, dorsal. Scale bars: 0.05 mm (A-F); 0.02 mm (G)." figureDoi="10.3897/zookeys.1115.83266.figure26" httpUri="https://binary.pensoft.net/fig/720746" pageId="0" pageNumber="1">26A</figureCitation>
) slender, two-segmented; proximal segment (syncoxa) unarmed, basally with short tube of maxillary gland; distal segment (basis) forming long claw, longer than proximal segment, ornamented with rows of small spinules and one tuft of few setules. Maxilliped (Fig.
<figureCitation id="59A84921D7907FC31D4B320E4FC2CA25" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Acontiophorus estivalis sp. nov., female A maxilla B maxilliped C leg 1 D leg 2 E leg 3 F leg 4 G right genital aperture, dorsal. Scale bars: 0.05 mm (A-F); 0.02 mm (G)." figureDoi="10.3897/zookeys.1115.83266.figure26" httpUri="https://binary.pensoft.net/fig/720746" pageId="0" pageNumber="1">26B</figureCitation>
) five-segmented, consisting of syncoxa, basis, three-segmented endopod, and terminal claw; syncoxa with trace of articulation delimiting praecoxal and coxal regions, coxal region with one seta on inner margin and row of spinules along outer margin; basis with one rudimentary seta at distal third of inner margin and row of spinules along outer margin; three endopodal segments armed with two, two, and one setae, respectively; terminal claw weakly curved, 66
<normalizedToken id="2CC129D0A61D3A271B5404B5033E0B23" originalValue="μm">μm</normalizedToken>
long, more than twice longer than third endopodal segment (31
<normalizedToken id="F365C2AD6E3C78CDA28AB8638564308D" originalValue="μm">μm</normalizedToken>
long).
</paragraph>
<caption id="5673514B58E9F52BCE3C9BD4F11BE0C3" doi="10.3897/zookeys.1115.83266.figure26" httpUri="https://binary.pensoft.net/fig/720746" pageId="0" pageNumber="1" start="Figure 26" startId="F26">
<paragraph id="978BD191B897F7154A329749ECC60996" pageId="0" pageNumber="1">
<emphasis id="51CFF949D05A7254296B71A4B794A13B" bold="true" pageId="0" pageNumber="1">Figure 26.</emphasis>
<taxonomicName id="91773A0225E2A106A1B5AAE31B82EA66" authorityName="Lee &amp; Chang &amp; Kim" authorityYear="2022" class="Copepoda" family="Asterocheridae" genus="Acontiophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acontiophorus estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="2DDEECC5BB5DD28FECEEC65FB3BF6692" italics="true" pageId="0" pageNumber="1">Acontiophorus estivalis</emphasis>
</taxonomicName>
sp. nov., female
<emphasis id="EA9A93E74AAC5685F7FEEBAE33D8DD0E" bold="true" pageId="0" pageNumber="1">A</emphasis>
maxilla
<emphasis id="6A4DA993A5BD7B22661A735D414B1667" bold="true" pageId="0" pageNumber="1">B</emphasis>
maxilliped
<emphasis id="E2E06A7500F0D97C227D57B8C0608BC8" bold="true" pageId="0" pageNumber="1">C</emphasis>
leg 1
<emphasis id="1434A1306B8B4E331AFE2A71BC7735C4" bold="true" pageId="0" pageNumber="1">D</emphasis>
leg 2
<emphasis id="CE687C1E7EB9DB50E73DF56EAE3E08CE" bold="true" pageId="0" pageNumber="1">E</emphasis>
leg 3
<emphasis id="E8CC79AA696CB093EA28B4048D54846D" bold="true" pageId="0" pageNumber="1">F</emphasis>
leg 4
<emphasis id="7B5033812954F8CD3644E02086F99F1C" bold="true" pageId="0" pageNumber="1">G</emphasis>
right genital aperture, dorsal. Scale bars: 0.05 mm (
<emphasis id="769D8F1ACEED5844B1004776E4E47169" bold="true" pageId="0" pageNumber="1">A-F</emphasis>
); 0.02 mm (
<emphasis id="59236F43B6D3BF7A7794060E3F92EE5D" bold="true" pageId="0" pageNumber="1">G</emphasis>
).
</paragraph>
</caption>
<paragraph id="69D0FFB3F6CA94CF962514C86F910A39" pageId="0" pageNumber="1">
Legs 1-4 (Fig.
<figureCitation id="935218DBD92B3EDEB84753BBD1AFEC03" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Acontiophorus estivalis sp. nov., female A maxilla B maxilliped C leg 1 D leg 2 E leg 3 F leg 4 G right genital aperture, dorsal. Scale bars: 0.05 mm (A-F); 0.02 mm (G)." figureDoi="10.3897/zookeys.1115.83266.figure26" httpUri="https://binary.pensoft.net/fig/720746" pageId="0" pageNumber="1">26C-F</figureCitation>
) biramous, with three-segmented rami. Inner coxal seta well-developed in legs 1-4; outer seta on basis small in legs 1-3, but markedly large in leg 4. Inner distal spine on basis of leg 1 extending to middle of second endopodal segment. Second endopodal segment of legs 1-4 with bicuspid outer distal corner. Inner distal process of third endopodal segment of leg 1 acutely pointed. Armature formula for legs 1-4 as follows:
</paragraph>
<paragraph id="B4F5077125B0C88297FA59755A0F298C" pageId="0" pageNumber="1">
<table id="0097386822CBAEF496D19990A27B98CF" pageId="0" pageNumber="1">
<tr id="E836F2A18DC5E0D8FF6268DEB0546FD3" pageId="0" pageNumber="1">
<td id="F101656808705EBCA7E3322489034E28" colspan="1" isEmpty="true" pageId="0" pageNumber="1" rowspan="1">-</td>
<td id="DE241DD54E6B6EA4096E6525C036D3E4" colspan="1" pageId="0" pageNumber="1" rowspan="1">Coxa</td>
<td id="57D031D29617D0A54D8859DF4A1D1E41" colspan="1" pageId="0" pageNumber="1" rowspan="1">Basis</td>
<td id="37125B02DA80DF7F875CC6BB03F40082" colspan="1" pageId="0" pageNumber="1" rowspan="1">Exopod</td>
<td id="1ABBBF2D36F9BDE05295A4E75EF79D8D" colspan="1" pageId="0" pageNumber="1" rowspan="1">Endopod</td>
</tr>
<tr id="D55FDFFF4DE134EB19520BC4F2284333" pageId="0" pageNumber="1">
<td id="881A59F1DBEFEDB02D02F10B7970FF58" colspan="1" pageId="0" pageNumber="1" rowspan="1">Leg 1</td>
<td id="4FE65FB995BB9638D42B147948F18254" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="85B68CA0B17F124BED509F2483A1254A" colspan="1" pageId="0" pageNumber="1" rowspan="1">1-I</td>
<td id="B3B7BD850B3B0F34CD8F27C974715658" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1; I-1; II, I, 5</td>
<td id="8C4C1D87A5AA30FF8F9ADEBF41A23FBF" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1; 0-2; 1, 2, 3</td>
</tr>
<tr id="61B5788866CA97111E309D4DD20F3A84" pageId="0" pageNumber="1">
<td id="D77E330C52CEA511E33C759C2487801B" colspan="1" pageId="0" pageNumber="1" rowspan="1">Leg 2</td>
<td id="C912FC2B34CED4782B28298A897E9A18" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="AB226B8E9883D24CFA2B9FFE27B0420C" colspan="1" pageId="0" pageNumber="1" rowspan="1">1-0</td>
<td id="FFB242B034E553D247800994F488BEEF" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1; I-1; III, I, 4</td>
<td id="CEF9F5E36B7CDC4550E8E6CCD15150CB" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1; 0-2; 1, 1+I, 3</td>
</tr>
<tr id="797376CE4CB024CCFED48AF4CE9AE691" pageId="0" pageNumber="1">
<td id="A1C5F760A575F755201B0890AF432D3F" colspan="1" pageId="0" pageNumber="1" rowspan="1">Leg 3</td>
<td id="5F58816CF71A14019A22ACF1A51B02BC" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="64CACC78DE19284488D0523FE91A7A26" colspan="1" pageId="0" pageNumber="1" rowspan="1">1-0</td>
<td id="4799B0655927F5EAAFBBD8244180EA60" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1; I-1; III, I, 3</td>
<td id="A0ACF0F495F387A98C5C04DCC07EF328" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1; 0-2; 1, I, 3</td>
</tr>
<tr id="4F94E36A6543B7EFCF52E96206250009" pageId="0" pageNumber="1">
<td id="C4B59CB83C635E0000DFA4ED18C493DF" colspan="1" pageId="0" pageNumber="1" rowspan="1">Leg 4</td>
<td id="D8A5EBF5B61B2CBB9B3B0D0076EF92CB" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1</td>
<td id="617E621CD995AD71F37050C25ED4D3D6" colspan="1" pageId="0" pageNumber="1" rowspan="1">1-0</td>
<td id="68E20D4809F4C8BEDFC2275D66AB7E92" colspan="1" pageId="0" pageNumber="1" rowspan="1">I-1; I-1; III, I, 3</td>
<td id="9FFD5AE3657BFE4C28EC7A936A491C7A" colspan="1" pageId="0" pageNumber="1" rowspan="1">0-1; 0-2; 1, I, 2</td>
</tr>
</table>
</paragraph>
<paragraph id="70B9C98F8789C3E179ACC7D38238D0EF" pageId="0" pageNumber="1">
Leg 5 (Fig.
<figureCitation id="D5F4B0B814CD8339AE67A37330DBDDFE" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Acontiophorus estivalis sp. nov., female A leg 5, dorsal. Male B habitus, dorsal C antennule (setae omitted) D leg 5, dorsal E leg 6. Scale bars: 0.02 mm (A, C-E); 0.1 mm (B)." figureDoi="10.3897/zookeys.1115.83266.figure27" httpUri="https://binary.pensoft.net/fig/720747" pageId="0" pageNumber="1">27A</figureCitation>
) two-segmented. First segment (protopod) broad, not articulated from somite, armed with large, naked outer distal seta and small, naked inner distal seta. Distal segment (exopod) 1.12
<normalizedToken id="4016CB74BCB13D3EBFD180BA5471FF55" originalValue="×">x</normalizedToken>
longer than wide (28
<normalizedToken id="A73BFB412375D0CB6C12696E72FF2FD5" originalValue="×">x</normalizedToken>
25
<normalizedToken id="C0F71013163DCB197F30B3F78FC15C7B" originalValue="μm">μm</normalizedToken>
), armed with five setae, and ornamented with fine setules on outer surface; two smaller setae on inner margin pinnate, terminal seta naked, two outer setae feebly pinnate. Leg 6 (Fig.
<figureCitation id="F6726909356D55721CDEE2E93270775B" captionStart="Figure 26" captionStartId="F26" captionText="Figure 26. Acontiophorus estivalis sp. nov., female A maxilla B maxilliped C leg 1 D leg 2 E leg 3 F leg 4 G right genital aperture, dorsal. Scale bars: 0.05 mm (A-F); 0.02 mm (G)." figureDoi="10.3897/zookeys.1115.83266.figure26" httpUri="https://binary.pensoft.net/fig/720746" pageId="0" pageNumber="1">26G</figureCitation>
) represented by one pinnate seta and two minute setules on genital operculum.
</paragraph>
<caption id="6899E84B4CF2FF414C1FD0E70DD63BE5" doi="10.3897/zookeys.1115.83266.figure27" httpUri="https://binary.pensoft.net/fig/720747" pageId="0" pageNumber="1" start="Figure 27" startId="F27">
<paragraph id="58C98E4A250FEA1EAA42F410650CE837" pageId="0" pageNumber="1">
<emphasis id="EFBCC5F4F0956EB9DEE5D04F591AEC67" bold="true" pageId="0" pageNumber="1">Figure 27.</emphasis>
<taxonomicName id="8686BB03539303E4901EC2B9BF2E4CFD" authorityName="Lee &amp; Chang &amp; Kim" authorityYear="2022" class="Copepoda" family="Asterocheridae" genus="Acontiophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acontiophorus estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="18473A95518F1CF4130C4DB75308955D" italics="true" pageId="0" pageNumber="1">Acontiophorus estivalis</emphasis>
</taxonomicName>
sp. nov., female
<emphasis id="9E5B792C16CC8520B4F0063E81E8532F" bold="true" pageId="0" pageNumber="1">A</emphasis>
leg 5, dorsal. Male
<emphasis id="534730A9F7733DC6D933E8B91FDD8B41" bold="true" pageId="0" pageNumber="1">B</emphasis>
habitus, dorsal
<emphasis id="D2D24F72D70E70CC8E7B41862E1F0AA1" bold="true" pageId="0" pageNumber="1">C</emphasis>
antennule (setae omitted)
<emphasis id="3807E7D4E9AC716688EC0AFBEE08D8DE" bold="true" pageId="0" pageNumber="1">D</emphasis>
leg 5, dorsal
<emphasis id="1CC1EDE72219DECF4FAC63D40B7735AD" bold="true" pageId="0" pageNumber="1">E</emphasis>
leg 6. Scale bars: 0.02 mm (
<emphasis id="3228BE13E73E94F1F22CAE6772A5F3F2" bold="true" pageId="0" pageNumber="1">A, C-E</emphasis>
); 0.1 mm (
<emphasis id="1E98C8EB616622E945F87603969FA40B" bold="true" pageId="0" pageNumber="1">B</emphasis>
).
</paragraph>
</caption>
<paragraph id="AE9CBE524AE97303B2AADA1118E82786" pageId="0" pageNumber="1">
<emphasis id="B8C8172E6BE076DFC822D7C88323A3A5" bold="true" pageId="0" pageNumber="1">Male.</emphasis>
Body form (Fig.
<figureCitation id="6EB1CA0D6CE179E5992C4CA24BC74B63" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Acontiophorus estivalis sp. nov., female A leg 5, dorsal. Male B habitus, dorsal C antennule (setae omitted) D leg 5, dorsal E leg 6. Scale bars: 0.02 mm (A, C-E); 0.1 mm (B)." figureDoi="10.3897/zookeys.1115.83266.figure27" httpUri="https://binary.pensoft.net/fig/720747" pageId="0" pageNumber="1">27B</figureCitation>
) as in female. Body length 890
<normalizedToken id="F5876700C92BDBBAF2259FF76CA963B7" originalValue="μm">μm</normalizedToken>
in dissected and figured paratype (length range 821-890
<normalizedToken id="CA2C3FC9E923CB153C4A8EF228EBBEA1" originalValue="μm">μm</normalizedToken>
). Prosome 600
<normalizedToken id="4572F60EE4606854AECDF7389DC2D4D8" originalValue="μm">μm</normalizedToken>
long. Cephalothorax slightly wider than long (374
<normalizedToken id="3C09C79ADE43496C4F9A7B2465FD392D" originalValue="×">x</normalizedToken>
407
<normalizedToken id="997E748415931990DB3542F958A7C611" originalValue="μm">μm</normalizedToken>
). Urosome five-segmented, Genital somite much wider than long. First two free abdominal somites broadened distally, with pointed posterolateral corners. Caudal ramus 1.44
<normalizedToken id="028423167898B185434A69073761C3EC" originalValue="×">x</normalizedToken>
longer than wide (46
<normalizedToken id="80AB76C25EC45CB235F1E0EB113E9ECB" originalValue="×">x</normalizedToken>
32
<normalizedToken id="41A5B68FE3E17CF7F7EEC3FC632B3406" originalValue="μm">μm</normalizedToken>
), armed as in female.
</paragraph>
<paragraph id="374D4C9678EC333A1B24F4DAAF4024E4" pageId="0" pageNumber="1">
Antennule (Fig.
<figureCitation id="9969C09C3AFBCFCB995DFA91FF04C175" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Acontiophorus estivalis sp. nov., female A leg 5, dorsal. Male B habitus, dorsal C antennule (setae omitted) D leg 5, dorsal E leg 6. Scale bars: 0.02 mm (A, C-E); 0.1 mm (B)." figureDoi="10.3897/zookeys.1115.83266.figure27" httpUri="https://binary.pensoft.net/fig/720747" pageId="0" pageNumber="1">27C</figureCitation>
) 168
<normalizedToken id="D67E34CB2CF43682226656564F476BD5" originalValue="μm">μm</normalizedToken>
long, 11-segmented; setae entangled, difficult to distinguish from one another; aesthetascs five on second segment, two on third, one on each 7th and 10th segment; aesthetasc on 10th segment large. Antenna as in female.
</paragraph>
<paragraph id="2F1951321FF3F43CA13AE37CE872DA80" pageId="0" pageNumber="1">
Oral siphon, mandible, maxillule, maxilla, maxilliped, and legs 1-4 same as those of female. Leg 5 (Fig.
<figureCitation id="1537511469D325BBAA52CC2DDD87E934" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Acontiophorus estivalis sp. nov., female A leg 5, dorsal. Male B habitus, dorsal C antennule (setae omitted) D leg 5, dorsal E leg 6. Scale bars: 0.02 mm (A, C-E); 0.1 mm (B)." figureDoi="10.3897/zookeys.1115.83266.figure27" httpUri="https://binary.pensoft.net/fig/720747" pageId="0" pageNumber="1">27D</figureCitation>
) also shaped as in female; inner distal seta on protopod pinnate; exopodal segment 1.33
<normalizedToken id="C964E6E6F343EC45A2DA69015CA7A5EE" originalValue="×">x</normalizedToken>
longer than wide (32
<normalizedToken id="D2EC41C20E341CC3B256F720116B7BA9" originalValue="×">x</normalizedToken>
24
<normalizedToken id="71E4FBEB58F79E485725FD40612C7D42" originalValue="μm">μm</normalizedToken>
), armed as in female. Leg 6 (Fig.
<figureCitation id="CC64A1468D6E20062411E95039CB19B3" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Acontiophorus estivalis sp. nov., female A leg 5, dorsal. Male B habitus, dorsal C antennule (setae omitted) D leg 5, dorsal E leg 6. Scale bars: 0.02 mm (A, C-E); 0.1 mm (B)." figureDoi="10.3897/zookeys.1115.83266.figure27" httpUri="https://binary.pensoft.net/fig/720747" pageId="0" pageNumber="1">27E</figureCitation>
) represented by three setae (two larger, weakly pinnate and one smaller naked) on genital operculum.
</paragraph>
</subSubSection>
<subSubSection id="0B1F43BC88015E3180A33880F5F16DF8" pageId="0" pageNumber="1" type="etymology">
<paragraph id="92DDC368567B82F04B1E2381BD117FED" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="B43E48107C495C72553A9A03FE7E153F" pageId="0" pageNumber="1">
The specific name
<emphasis id="D0C3C1B193EFF7188F13CF442799A2F8" italics="true" pageId="0" pageNumber="1">estivalis</emphasis>
is derived from Latin
<emphasis id="D2AF946437D3919229B75A288923F1EE" italics="true" pageId="0" pageNumber="1">estival</emphasis>
(summer), indicating the discovery of the new species in the summer.
</paragraph>
</subSubSection>
<subSubSection id="45DF93042DF04C253B7FEFAA8BEC508A" pageId="0" pageNumber="1" type="remarks">
<paragraph id="76DAE40DA5671C343B8150B6C0F0217E" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="BF61A1678115E4404F8A4DC9092A2258" pageId="0" pageNumber="1">
The segmentation of the antennule appears to be a reliable character for the differentiation of
<taxonomicName id="A9A9DA71C605BB374ED9103161396B8E" authorityName="Brady" authorityYear="1880" class="Copepoda" family="Asterocheridae" genus="Acontiophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acontiophorus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="F229A1281EAE8FFB9676F3F989BD7827" italics="true" pageId="0" pageNumber="1">Acontiophorus</emphasis>
</taxonomicName>
species.
<taxonomicName id="2EA549B7B6EEF4FA755E315E6F6976D8" genus="Aconiophorus" lsidName="Aconiophorus estivalis" pageId="0" pageNumber="1" rank="species" species="estivalis">
<emphasis id="D8E8F6CDE49DCCE2539B66C2AEB2C4C7" italics="true" pageId="0" pageNumber="1">Aconiophorus estivalis</emphasis>
</taxonomicName>
sp. nov. has an 11-segmented antennule in the female; this feature is shared with three congeners,
<taxonomicName id="3920D316B9B8C164B69212988A2191FE" class="Hexanauplia" kingdom="Animalia" lsidName="A. antennatus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="DA578A1525CB7881C2FD7B7879B069B0" italics="true" pageId="0" pageNumber="1">A. antennatus</emphasis>
</taxonomicName>
Hansen, 1923,
<taxonomicName id="F05E00D7F8B3F81DC9DD9F9B850BFACD" class="Hexanauplia" kingdom="Animalia" lsidName="A. scutatus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="ACA7893CE0E0C645C6B0A636DF7A6721" italics="true" pageId="0" pageNumber="1">A. scutatus</emphasis>
</taxonomicName>
(Brady &amp; Robertson, 1873) and
<taxonomicName id="3BCF5358F8B567E805AF8FE2BA43CC16" class="Hexanauplia" kingdom="Animalia" lsidName="A. zealandicus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zealandicus">
<emphasis id="8D696102E5866F8659C5E8E59CD7AAF4" italics="true" pageId="0" pageNumber="1">A. zealandicus</emphasis>
</taxonomicName>
Sewell, 1944.
<taxonomicName id="FF6CEAFACD78222AB725E750D7B1E3B3" genus="Acontiphorus" lsidName="Acontiphorus antennatus" pageId="0" pageNumber="1" rank="species" species="antennatus">
<emphasis id="30689AE35647D75AA321B0A31A8AB8F8" italics="true" pageId="0" pageNumber="1">Acontiphorus antennatus</emphasis>
</taxonomicName>
was redescribed by
<bibRefCitation id="5A51401168FD6719583A2E5E82E6061F" author="Eiselt, VJ" journalOrPublisher="Mathematisch-Naturwissenschaftliche Klasse" pageId="0" pageNumber="1" pagination="177 - 185" refId="B13" refString="Eiselt, VJ, 1969. Revision von Acontiophorus antennatus Hansen 1923 und materialien zur variabilitaet von Acontiophorus armatus Brady 1880 (Cyclop. Siph., Copepoda, Crust.). Sitzungsberichten der Oesterreichischen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 177 (8-10): 177 - 185" title="Revision von Acontiophorus antennatus Hansen 1923 und materialien zur variabilitaet von Acontiophorus armatus Brady 1880 (Cyclop. Siph., Copepoda, Crust.). Sitzungsberichten der Oesterreichischen Akademie der Wissenschaften." volume="177" year="1969">Eiselt (1969)</bibRefCitation>
and according to his illustration and description, the caudal ramus of
<taxonomicName id="F4D8F61DC32EEAB9EE04067A588854D4" class="Hexanauplia" kingdom="Animalia" lsidName="A. antennatus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="A8A19059B6481EB95B08080554D9B8F2" italics="true" pageId="0" pageNumber="1">A. antennatus</emphasis>
</taxonomicName>
is ~ 4
<normalizedToken id="2B2A4C6E3D2D50D76A8C931C5A15DB2B" originalValue="×">x</normalizedToken>
longer than wide in the female (cf. 2.12
<normalizedToken id="41139A505FFA1B4F0A434171B33B5FA0" originalValue="×">x</normalizedToken>
longer than wide in
<taxonomicName id="F13B665A67DAEFE453A4B0C98B46A461" class="Hexanauplia" kingdom="Animalia" lsidName="A. estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="D425618BFBBB8CA67FCEB93939AFAFFB" italics="true" pageId="0" pageNumber="1">A. estivalis</emphasis>
</taxonomicName>
sp. nov.) and the third exopodal segment of leg 1 is armed with three spines and four setae (formula III, 4; against III, 5 in
<taxonomicName id="241790CBA2CEE2DFE541B507478AEF48" class="Hexanauplia" kingdom="Animalia" lsidName="A. estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="8CE4A4568E5E5003D09FA318B6686A3F" italics="true" pageId="0" pageNumber="1">A. estivalis</emphasis>
</taxonomicName>
sp. nov.). In
<taxonomicName id="CCCB2842406E4663C200AD671572CA05" class="Hexanauplia" kingdom="Animalia" lsidName="A. zealandicus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zealandicus">
<emphasis id="A3DF3D6CE6692CB0C955BF990C4B93DF" italics="true" pageId="0" pageNumber="1">A. zealandicus</emphasis>
</taxonomicName>
the male antennule is ten-segmented (cf. 11-segmented in
<taxonomicName id="D4CCD28F32CF9CBBEB643A817619DB4F" class="Hexanauplia" kingdom="Animalia" lsidName="A. estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="A10A4CA3A4711E25937E405B4786AE17" italics="true" pageId="0" pageNumber="1">A. estivalis</emphasis>
</taxonomicName>
sp. nov.) and the oral siphon is extremely long, extending beyond the caudal rami (
<bibRefCitation id="CD346E32731A5C3C8C5E697D49FD56A7" author="Nicholls, AG" journalOrPublisher="Records of the South Australian Museum" pageId="0" pageNumber="1" pagination="1 - 62" refId="B44" refString="Nicholls, AG, 1944. Littoral Copepoda from South Australia (II) Calanoida, Cyclopoida, Notodelphyoida, Monstrilloida and Caligoida. Records of the South Australian Museum 8: 1 - 62" title="Littoral Copepoda from South Australia (II) Calanoida, Cyclopoida, Notodelphyoida, Monstrilloida and Caligoida." volume="8" year="1944">Nicholls 1944</bibRefCitation>
) (cf. extending to middle of genital double-somite in
<taxonomicName id="73C696F637AD7BD0C98D9F6CB6AFE6C8" authority="estivalis" class="Hexanauplia" genus="A." kingdom="Animalia" lsidName="A." order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">A. estivalis</taxonomicName>
sp. nov.). Therefore,
<taxonomicName id="8CC211AE5F546E2DF87C72D8B48C74A1" class="Hexanauplia" kingdom="Animalia" lsidName="A. antennatus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antennatus">
<emphasis id="D4E420E9A9C88CF9F3F3E178A71B8C3F" italics="true" pageId="0" pageNumber="1">A. antennatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4445EC54F49A2110DE56E529123C9646" class="Hexanauplia" kingdom="Animalia" lsidName="A. zelandicus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="zelandicus">
<emphasis id="8440D16A3053A35744F4F5FBD0AA6FE6" italics="true" pageId="0" pageNumber="1">A. zelandicus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4F711B6C45CF5F708AA31B8F5D964809" class="Hexanauplia" kingdom="Animalia" lsidName="A. estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="1BE90A6523B7EB0FB5D555C2149D0366" italics="true" pageId="0" pageNumber="1">A. estivalis</emphasis>
</taxonomicName>
sp. nov. with confidence.
<taxonomicName id="7C2A60D6CEAD5A085E942022A6CCEFCB" authorityName="Lee &amp; Chang &amp; Kim" authorityYear="2022" class="Copepoda" family="Asterocheridae" genus="Acontiophorus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acontiophorus estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="BEE249D2BBC6C16462C7BF73EBEA64B8" italics="true" pageId="0" pageNumber="1">Acontiophorus estivalis</emphasis>
</taxonomicName>
sp. nov. closely resembles
<taxonomicName id="F2379349434CBC61FE41E6FB087F78A7" class="Hexanauplia" kingdom="Animalia" lsidName="A. scutatus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="796FFB2BF1DC7AEDA0BE25761A530668" italics="true" pageId="0" pageNumber="1">A. scutatus</emphasis>
</taxonomicName>
. As noticeable differences between them, the caudal ramus of
<taxonomicName id="AD632AD1FC110EDC0AEFE84CA0CE8B99" class="Hexanauplia" kingdom="Animalia" lsidName="A. scutatus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="60F916A3688FF65EF96EC6CE1F436506" italics="true" pageId="0" pageNumber="1">A. scutatus</emphasis>
</taxonomicName>
is 3
<normalizedToken id="445D55D96B39709BD3B766BFDF0BB7D3" originalValue="×">x</normalizedToken>
longer than wide, and the oral siphon of the latter species extends to the caudal rami (
<bibRefCitation id="4CF0E56414635C525050BCF87795B98C" author="Sars, GO" journalOrPublisher="Bergen Museum, Bergen" pageId="0" pageNumber="1" pagination="105 - 140" refId="B49" refString="Sars, GO, 1915. Copepoda Cyclopoida. Parts IX &amp; X. Ascomyzontidae (concluded), Acontiophoridae, Myzopontiidae, Dyspontiidae, Artotrogidae, Cancerillidae. An account of the Crustacea of Norway with short descriptions and figures of all the species. Bergen Museum, Bergen 6: 105 - 140" title="Copepoda Cyclopoida. Parts IX &amp; X. Ascomyzontidae (concluded), Acontiophoridae, Myzopontiidae, Dyspontiidae, Artotrogidae, Cancerillidae. An account of the Crustacea of Norway with short descriptions and figures of all the species." volume="6" year="1915">Sars 1915</bibRefCitation>
) (cf. the siphon extends to the middle of the genital double-somite in
<taxonomicName id="DB877E323326C04DC461CB2B48FEB8E3" class="Hexanauplia" kingdom="Animalia" lsidName="A. estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="B09F924442A6E5BC264ECF7F7B6A652D" italics="true" pageId="0" pageNumber="1">A. estivalis</emphasis>
</taxonomicName>
sp. nov.). Additionally, the first segment of the male antennule of
<taxonomicName id="F2068717EF5055802EE016E82F12C011" class="Hexanauplia" kingdom="Animalia" lsidName="A. scutatus" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="scutatus">
<emphasis id="7227BDBCFFE65D30BEBEBF4CECA7C824" italics="true" pageId="0" pageNumber="1">A. scutatus</emphasis>
</taxonomicName>
bears an aesthetasc, according to the illustration of
<bibRefCitation id="009E86164AD1625D248FAAD4DD918D20" author="Sars, GO" journalOrPublisher="Bergen Museum, Bergen" pageId="0" pageNumber="1" pagination="105 - 140" refId="B49" refString="Sars, GO, 1915. Copepoda Cyclopoida. Parts IX &amp; X. Ascomyzontidae (concluded), Acontiophoridae, Myzopontiidae, Dyspontiidae, Artotrogidae, Cancerillidae. An account of the Crustacea of Norway with short descriptions and figures of all the species. Bergen Museum, Bergen 6: 105 - 140" title="Copepoda Cyclopoida. Parts IX &amp; X. Ascomyzontidae (concluded), Acontiophoridae, Myzopontiidae, Dyspontiidae, Artotrogidae, Cancerillidae. An account of the Crustacea of Norway with short descriptions and figures of all the species." volume="6" year="1915">Sars (1915)</bibRefCitation>
, which is absent in
<taxonomicName id="E8A4F9ECF534E31F2620612F6D489665" class="Hexanauplia" kingdom="Animalia" lsidName="A. estivalis" order="Siphonostomatoida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="estivalis">
<emphasis id="B1C3ED775E65CC3B534FF71580E9C52E" italics="true" pageId="0" pageNumber="1">A. estivalis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
</treatment>
</document>