treatments-xml/data/EB/13/87/EB138782C05A85504D992D82FAE5B2DF.xml

<document id="710C2B4055A3852C01609E696633B257" ID-CLB-Dataset="26112" ID-DOI="10.11646/zootaxa.4651.1.3" ID-GBIF-Dataset="0534b00e-640e-42d1-8706-80911b943bba" ID-ISSN="1175-5326" ID-Zenodo-Dep="3359026" ID-ZooBank="24EBA328-0834-48D1-84CA-B2E819EE56DD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564763567067" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2019" docId="EB138782C05A85504D992D82FAE5B2DF" docLanguage="en" docName="zootaxa.4651.1.3.pdf" docOrigin="Zootaxa 4651 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Salmoneus venustus Anker 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="58" masterDocId="172AFFFAC05B85574D0E2F2CFFA8B740" masterDocTitle="On three symbiotic species of the alpheid shrimp genus Salmoneus Holthuis, 1955 from the Indo-West Pacific, including one new to science (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="63" masterPageNumber="51" pageNumber="52" updateTime="1698748554678" updateUser="ExternalLinkService">
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<mods:title id="8E9D52CAA48F1BDED9D7E5A4453D3A1C">On three symbiotic species of the alpheid shrimp genus Salmoneus Holthuis, 1955 from the Indo-West Pacific, including one new to science (Malacostraca: Decapoda: Caridea)</mods:title>
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<subSubSection id="2BA0651FC05A85564D992D82FE5BB589" box="[151,499,686,713]" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="63053694C05A85564D992D82FE5BB589" blockId="1.[151,499,686,748]" box="[151,499,686,713]" pageId="1" pageNumber="52">
<heading id="384D81F8C05A85564D992D82FE5BB589" bold="true" box="[151,499,686,713]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
<emphasis id="51CEEA86C05A85564D992D82FE5BB589" bold="true" box="[151,499,686,713]" pageId="1" pageNumber="52">
<taxonomicName id="A4BA4D17C05A85564D992D82FE26B589" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[151,398,686,713]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05A85564D992D82FE26B589" bold="true" box="[151,398,686,713]" italics="true" pageId="1" pageNumber="52">Salmoneus venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05A85564C9B2D83FE5BB589" box="[405,499,687,713]" pageId="1" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="2BA0651FC05A85564D992DFEFEA8B5AC" box="[151,256,722,748]" pageId="1" pageNumber="52" type="description">
<paragraph id="63053694C05A85564D992DFEFEA8B5AC" blockId="1.[151,499,686,748]" box="[151,256,722,748]" pageId="1" pageNumber="52">
<figureCitation id="FB812A11C05A85564D992DFEFEA8B5AC" box="[151,256,722,748]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1807,1832]" captionStartId-1="4.[151,250,1689,1714]" captionStartId-2="5.[151,250,1940,1965]" captionStartId-3="6.[151,250,1894,1919]" captionTargetBox-0="[169,1414,285,1778]" captionTargetBox-1="[196,1395,200,1657]" captionTargetBox-2="[189,1372,641,1888]" captionTargetBox-3="[166,1421,166,1875]" captionTargetId-0="figure@3.[151,1436,273,1788]" captionTargetId-1="figure@4.[179,1409,181,1670]" captionTargetId-2="figure@5.[183,1404,628,1916]" captionTargetId-3="figure@6.[166,1421,166,1875]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." captionText-1="FIGURE 2. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—third maxilliped, lateral view; b—same, tip of ultimate article, lateral view; c—second pereiopod, lateral view; d—third pereiopod, lateral view; e—same, propodus and dactylus, mesial view; f—fourth pereiopod, lateral view; g—fifth pereiopod, lateral view." captionText-2="FIGURE 3. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—major cheliped, lateral view; b—same, mesial view; c—same, carpus and chela, ventrolateral view; d—same, chela fingers opened, ventrolateral view; e—minor cheliped, lateral view; f—same, carpus and chela, mesial view." captionText-3="FIGURE 4. Salmoneus venustus sp. nov.: non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH. 2015-08-0031 [a, b]; holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [c]; a—living individual, dorsal view; b—same, lateral view (both photographs by the author); c—fresh specimen post-mortem, lateral view (photograph courtesy of Dr. Ivan N. Marin)." httpUri-0="https://zenodo.org/record/3359028/files/figure.png" httpUri-1="https://zenodo.org/record/3359030/files/figure.png" httpUri-2="https://zenodo.org/record/3359032/files/figure.png" httpUri-3="https://zenodo.org/record/3359034/files/figure.png" pageId="1" pageNumber="52">Figs. 1–4</figureCitation>
</paragraph>
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<paragraph id="63053694C05A85564D992C36FEB5B43C" blockId="1.[151,1437,793,2045]" pageId="1" pageNumber="52">
<emphasis id="51CEEA86C05A85564D992C36FEEDB474" bold="true" box="[151,325,793,820]" pageId="1" pageNumber="52">Type material.</emphasis>
<materialsCitation id="D3D23CC9C05A85564C432C36FEB5B43C" ID-GBIF-Occurrence="2302320215" collectingDate="2012-10-24" collectorName="I. N. Marin" country="Vietnam" elevation="7" location="Nha Trang Bay" municipality="Che" pageId="1" pageNumber="52" specimenCount="1" typeStatus="holotype">
<typeStatus id="BC018836C05A85564C432C36FE12B474" box="[333,442,794,820]" pageId="1" pageNumber="52" type="holotype">Holotype</typeStatus>
: non-ovigerous specimen (cl 
<quantity id="A4429B71C05A85564E0D2C36FCF3B475" box="[771,859,794,821]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="52" unit="mm" value="6.4">6.4 mm</quantity>
, dissected), MNHN-IU-2014-20290, 
<collectingCountry id="1BAD7604C05A8556480E2C36FACAB474" box="[1280,1378,794,820]" name="Vietnam" pageId="1" pageNumber="52">Vietnam</collectingCountry>
, 
<location id="6665604FC05A855648602C36FEB9B419" LSID="urn:lsid:plazi:treatment:EB138782C05A85504D992D82FAE5B2DF:6665604FC05A855648602C36FEB9B419" country="Vietnam" municipality="Che" name="Nha Trang Bay" pageId="1" pageNumber="52">Nha Trang Bay</location>
, north side of 
<collectingMunicipality id="8361ACEEC05A85564CB32C12FE42B418" box="[445,490,830,856]" pageId="1" pageNumber="52">Che</collectingMunicipality>
Island, depth: 
<elevation id="E897D1A7C05A85564F942C12FD4EB419" box="[666,742,830,857]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="1" pageNumber="52" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">
<quantity id="A4429B71C05A85564F942C12FD4EB419" box="[666,742,830,857]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="1" pageNumber="52" unit="m" value="7.0" valueMax="8.0" valueMin="6.0">6–8 m</quantity>
</elevation>
, in burrow of 
<taxonomicName id="A4BA4D17C05A85564E9F2C12FBA9B418" authorityName="Stimpson" authorityYear="1866" box="[913,1025,830,856]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="51CEEA86C05A85564E9F2C12FBA9B418" box="[913,1025,830,856]" italics="true" pageId="1" pageNumber="52">Glypturus</emphasis>
</taxonomicName>
sp., suction pump, leg. 
<collectorName id="CE4F5342C05A855648152C13FA30B418" box="[1307,1432,830,857]" pageId="1" pageNumber="52">I.N. Marin</collectorName>
, 
<collectingDate id="0740E9BCC05A85564D992C4EFEB1B43C" box="[151,281,866,892]" pageId="1" pageNumber="52" value="2012-10-24">
<date id="17041054C05A85564D992C4EFEB1B43C" box="[151,281,866,892]" pageId="1" pageNumber="52" value="2012-10-24">24.10.2012</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
<paragraph id="63053694C05A85564DC92CA9FAF4B4A8" blockId="1.[151,1437,793,2045]" pageId="1" pageNumber="52">
<emphasis id="51CEEA86C05A85564DC92CA9FE18B4E0" bold="true" box="[199,432,901,928]" pageId="1" pageNumber="52">Additional material</emphasis>
. 1non-ovigerous specimen (cl 
<quantity id="A4429B71C05A85564E0F2CA9FCF0B4E1" box="[769,856,901,929]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="52" unit="mm" value="5.5">5.5 mm</quantity>
), 
<collectionCode id="05ABAE51C05A85564E662CAAFC70B4E0" box="[872,984,902,928]" pageId="1" pageNumber="52">OUMNH</collectionCode>
. 
<collectionCode id="05ABAE51C05A85564ED52CAAFBA8B4E0" box="[987,1024,902,928]" country="Austria" name="Zoological Collection, University of Vienna" pageId="1" pageNumber="52" type="University or college">ZC</collectionCode>
.2015-08-0031, 
<collectingCountry id="1BAD7604C05A855649BF2CAAFB50B4E0" box="[1201,1272,902,928]" name="Oman" pageId="1" pageNumber="52">Oman</collectingCountry>
, northern coast east of 
<collectingCountry id="1BAD7604C05A85564DE62C87FE95B485" box="[232,317,939,965]" name="Oman" pageId="1" pageNumber="52">Muscat</collectingCountry>
, shallow bay between Yiti and Sifah, shallow bay with sand-mud, fringed by mangroves, near shore and mangroves, depth at low tide: 
<quantity id="A4429B71C05A85564F142CE2FDF4B4A9" box="[538,604,974,1001]" metricMagnitude="-1" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="52" unit="m" value="0.7">0.7 m</quantity>
, in burrow in muddy sand, suction pump, leg. A. Anker, 
<date id="17041054C05A855649D92CE2FAF1B4A8" box="[1239,1369,974,1000]" pageId="1" pageNumber="52" value="2010-08-23">23.08.2010</date>
.
</paragraph>
<paragraph id="63053694C05A85564DC92CDDFDFBB3A5" blockId="1.[151,1437,793,2045]" pageId="1" pageNumber="52">
<emphasis id="51CEEA86C05A85564DC92CDDFEF9B34C" bold="true" box="[199,337,1009,1036]" pageId="1" pageNumber="52">Description</emphasis>
. Small-sized (cl 
<quantity id="A4429B71C05A85564F1C2CDDFD33B34D" box="[530,667,1009,1037]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.4" metricValueMin="5.5" pageId="1" pageNumber="52" unit="mm" value="5.95" valueMax="6.4" valueMin="5.5">5.5–6.4 mm</quantity>
) alpheid shrimp with stout, non-compressed body. Carapace mostly glabrous, with very few setae (
<typeStatus id="BC018836C05A85564CF72B3AFDF6B371" box="[505,606,1046,1073]" pageId="1" pageNumber="52" type="holotype">holotype</typeStatus>
) or more densely setose dorsally (
<typeStatus id="BC018836C05A85564EE42B3AFBF9B370" box="[1002,1105,1046,1072]" pageId="1" pageNumber="52" type="paratype">paratype</typeStatus>
). Rostrum moderately developed, broadly subtriangular, about as long as wide at base, acute distally, reaching distal margin of first article of antennular peduncle, lateral margins feebly concave; rostral or mid-dorsal carina absent (
<figureCitation id="FB812A11C05A8556497B2B72FB54B339" box="[1141,1276,1118,1145]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="1" pageNumber="52">Fig. 1a, b, h</figureCitation>
). Orbital teeth relatively small, sharp distally, pointing slightly mesially in dorsal view, anteriorly in lateral view (
<figureCitation id="FB812A11C05A8556480E2BAEFA27B3DD" box="[1280,1423,1154,1181]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="1" pageNumber="52">Fig. 1a, b, h</figureCitation>
). Pterygostomial region broadly rounded; anterolateral suture present (
<figureCitation id="FB812A11C05A85564EAE2B8AFC5AB381" box="[928,1010,1190,1217]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="1" pageNumber="52">Fig. 1b</figureCitation>
); cardiac notch well developed. Epistomial sclerite with short acute process.
</paragraph>
<paragraph id="63053694C05A85564DC92BC2FAAEB2A1" blockId="1.[151,1437,793,2045]" pageId="1" pageNumber="52">
Pleon with first to fourth pleura broadly rounded to slightly angular; fifth pleuron with posteroventral margin forming subacute angle; sixth pleonite with short suture but without articulated flap, posterior margin produced into long sharp tooth flanking telson (
<figureCitation id="FB812A11C05A85564F332A1AFD25B210" box="[573,653,1334,1360]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="1" pageNumber="52">Fig. 1c</figureCitation>
); preanal plate rounded. Telson fairly slender, subrectangular, slightly tapering distally, almost three times as long as maximal (proximal) width; dorsal surface with two pairs of small cuspidate setae both inserted at some distance from lateral margin, first pair slightly posterior to telson mid-length, second pair between 0.7 and 0.8 of telson length; posterior margin with shallow notch; posterolateral angles each with one pair of spiniform setae, mesial stouter and about 1.3–1.4 times as long as lateral (
<figureCitation id="FB812A11C05A855649852AEAFB51B2A1" box="[1163,1273,1478,1505]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="1" pageNumber="52">Fig. 1d, e</figureCitation>
).
</paragraph>
<paragraph id="63053694C05A85564DC92AC7FA88B169" blockId="1.[151,1437,793,2045]" pageId="1" pageNumber="52">
Eyes completely concealed in dorsal view, partly exposed in lateral view; cornea reduced, restricted to dorsolateral surface of eyestalk; anteromesial margin of eyestalk with dorsally produced tubercle (
<figureCitation id="FB812A11C05A855649862922FABAB169" box="[1160,1298,1550,1577]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="1" pageNumber="52">Fig. 1a, b, h</figureCitation>
).
</paragraph>
<paragraph id="63053694C05A85564DC9291EFBE5B009" blockId="1.[151,1437,793,2045]" pageId="1" pageNumber="52">
Antennular peduncle stout; stylocerite moderately long, slender, with sharp tip reaching or almost reaching midlength of second article; ventromesial carina with small, anteriorly directed tooth; second article about 1.3 times as long as wide; lateral antennular flagellum with very short fused portion, consisting of at most three discernable subdivisions, and well-developed secondary ramus, latter with five or so groups of aesthetascs; mesial antennular flagellum much stouter than lateral (
<figureCitation id="FB812A11C05A85564F2529EEFD3FB19D" box="[555,663,1730,1757]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="1" pageNumber="52">Fig. 1a, b</figureCitation>
). Antenna with stout basicerite, its distoventral margin bearing robust subacute tooth; scaphocerite ovoid in general shape, not reaching end of antennular peduncle, with straight lateral margin and very broad blade, latter highly convex anteriorly and significantly overreaching small sharp distolateral tooth; carpocerite short, cylindrical, reaching mid-length of scaphocerite (
<figureCitation id="FB812A11C05A85564EDC2802FB97B009" box="[978,1087,1838,1865]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="1" pageNumber="52">Fig. 1a, b</figureCitation>
).
</paragraph>
<paragraph id="63053694C05A85564DC9287EFEB3B0BD" blockId="1.[151,1437,793,2045]" pageId="1" pageNumber="52">
Mouthparts not dissected, typical for genus in external observation. Third maxilliped slender, pediform; coxa with somewhat elongated, oval-shaped lateral plate; antepenultimate article slightly flattened ventrolaterally; penultimate article about four times as long as wide; ultimate article with numerous rows of short serrulate setae and longer simple setae, tip with blunt corneous point, apparently without spiniform setae; arthrobranch well developed (
<figureCitation id="FB812A11C05A85564DAE28CEFEA5B0BD" box="[160,269,2018,2045]" captionStart="FIGURE 2" captionStartId="4.[151,250,1689,1714]" captionTargetBox="[196,1395,200,1657]" captionTargetId="figure@4.[179,1409,181,1670]" captionTargetPageId="4" captionText="FIGURE 2. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—third maxilliped, lateral view; b—same, tip of ultimate article, lateral view; c—second pereiopod, lateral view; d—third pereiopod, lateral view; e—same, propodus and dactylus, mesial view; f—fourth pereiopod, lateral view; g—fifth pereiopod, lateral view." httpUri="https://zenodo.org/record/3359030/files/figure.png" pageId="1" pageNumber="52">Fig. 2a, b</figureCitation>
).
</paragraph>
<paragraph id="63053694C05985554DC92FB5FCF8B524" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="53">
Chelipeds very asymmetrical in shape and dissimilar in size, carried flexed when not in use. Major cheliped relatively slender; ischium unarmed, flattened ventrolaterally; merus slender, more than six times as long as proximal width, widening distally, smooth, distodorsal and distomesial margins with rounded lobes, ventrolateral surface depressed distally; carpus short, cup-shaped, with strong subacute distoventral process; chela subcylindrical, more or less rounded in cross-section, with palm about as long as fingers, smooth, except for some small bumps on ventrolateral surface; fingers not gaping when closed, subequal, strongly crossing distally, not noticeably twisted, with evenly serrated cutting edges; cutting edge of both dactylus and pollex with about 14–15 teeth gradually increasing in size from most proximal tooth to mid-length teeth, then decreasing again towards most-distal tooth (
<figureCitation id="FB812A11C0598555482E2EB9FACAB6F0" box="[1312,1378,405,432]" captionStart="FIGURE 3" captionStartId="5.[151,250,1940,1965]" captionTargetBox="[189,1372,641,1888]" captionTargetId="figure@5.[183,1404,628,1916]" captionTargetPageId="5" captionText="FIGURE 3. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—major cheliped, lateral view; b—same, mesial view; c—same, carpus and chela, ventrolateral view; d—same, chela fingers opened, ventrolateral view; e—minor cheliped, lateral view; f—same, carpus and chela, mesial view." httpUri="https://zenodo.org/record/3359032/files/figure.png" pageId="2" pageNumber="53">Fig. 3</figureCitation>
a–d). Minor cheliped significantly smaller than major cheliped, slender; ischium unarmed on ventrolateral surface; merus longer than ischium, not swollen or widening distally, smooth, unarmed, slightly depressed ventrally; carpus about 1.1 times length of merus, cylindrical, slightly widening distally; chela 0.6 times length of carpus, simple, with palm subequal to fingers in length, smooth; fingers noticeably gaping when closed, subequal in length, crossing distally, cutting edges armed with four or five minute teeth (
<figureCitation id="FB812A11C05985554FD42D65FCEBB524" box="[730,835,585,612]" captionStart="FIGURE 3" captionStartId="5.[151,250,1940,1965]" captionTargetBox="[189,1372,641,1888]" captionTargetId="figure@5.[183,1404,628,1916]" captionTargetPageId="5" captionText="FIGURE 3. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—major cheliped, lateral view; b—same, mesial view; c—same, carpus and chela, ventrolateral view; d—same, chela fingers opened, ventrolateral view; e—minor cheliped, lateral view; f—same, carpus and chela, mesial view." httpUri="https://zenodo.org/record/3359032/files/figure.png" pageId="2" pageNumber="53">Fig. 3e, f</figureCitation>
).
</paragraph>
<paragraph id="63053694C05985554DC92D41FA91B3E3" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="53">
Second pereiopod slender; ischium unarmed on ventrolateral surface; merus noticeably longer than ischium; carpus with five subdivisions, first surpassing four others combined, with ratio approximately equal to 5.0/1.0/0.8/0.8/1.2; chela longer than distal-most carpal subdivision, simple (
<figureCitation id="FB812A11C05985554EF52D99FBE6B58F" box="[1019,1102,693,719]" captionStart="FIGURE 2" captionStartId="4.[151,250,1689,1714]" captionTargetBox="[196,1395,200,1657]" captionTargetId="figure@4.[179,1409,181,1670]" captionTargetPageId="4" captionText="FIGURE 2. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—third maxilliped, lateral view; b—same, tip of ultimate article, lateral view; c—second pereiopod, lateral view; d—third pereiopod, lateral view; e—same, propodus and dactylus, mesial view; f—fourth pereiopod, lateral view; g—fifth pereiopod, lateral view." httpUri="https://zenodo.org/record/3359030/files/figure.png" pageId="2" pageNumber="53">Fig. 2c</figureCitation>
). Third pereiopod slender; ischium with three cuspidate setae on ventrolateral surface; merus about seven times as long as wide, unarmed; carpus almost 0.7 times length of merus, noticeably more slender, with small spiniform seta distoventrally; propodus not noticeably longer than carpus, with two widely spaced spiniform setae on ventral margin, in addition to three longer spiniform setae near dactylar base; dactylus about half-length of propodus, slender, conical, simple, smoothly curving distally (
<figureCitation id="FB812A11C05985554C2D2C45FE27B4C3" box="[291,399,873,899]" captionStart="FIGURE 2" captionStartId="4.[151,250,1689,1714]" captionTargetBox="[196,1395,200,1657]" captionTargetId="figure@4.[179,1409,181,1670]" captionTargetPageId="4" captionText="FIGURE 2. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—third maxilliped, lateral view; b—same, tip of ultimate article, lateral view; c—second pereiopod, lateral view; d—third pereiopod, lateral view; e—same, propodus and dactylus, mesial view; f—fourth pereiopod, lateral view; g—fifth pereiopod, lateral view." httpUri="https://zenodo.org/record/3359030/files/figure.png" pageId="2" pageNumber="53">Fig. 2d, e</figureCitation>
). Fourth pereiopod more slender than third pereiopod; ischium unarmed on ventrolateral surface; merus almost eight times as long as wide; carpus 0.7 times length of merus, more slender than merus, unarmed distoventrally; propodus distinctly longer than carpus, without spiniform setae on ventral surface, distoventral margin adjacent to dactylus with one long spiniform seta; dactylus about half-length of propodus, similar to that of third pereiopod (
<figureCitation id="FB812A11C05985554C192CD5FECFB354" box="[279,359,1017,1044]" captionStart="FIGURE 2" captionStartId="4.[151,250,1689,1714]" captionTargetBox="[196,1395,200,1657]" captionTargetId="figure@4.[179,1409,181,1670]" captionTargetPageId="4" captionText="FIGURE 2. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—third maxilliped, lateral view; b—same, tip of ultimate article, lateral view; c—second pereiopod, lateral view; d—third pereiopod, lateral view; e—same, propodus and dactylus, mesial view; f—fourth pereiopod, lateral view; g—fifth pereiopod, lateral view." httpUri="https://zenodo.org/record/3359030/files/figure.png" pageId="2" pageNumber="53">Fig. 2f</figureCitation>
). Fifth pereiopod as slender as fourth pereiopod; ischium unarmed; merus almost eight times as long as wide, unarmed; carpus noticeably more slender than merus, as long as merus, unarmed distoventrally; propodus very long, slender, 1.5 times as long as carpus, with numerous rows of progressively longer serrulate setae forming dense cleaning brush on distal ventrolateral surface, ventral and ventromesial margin without spiniform setae; dactylus about 0.4 times length of propodus, similar to that of third and fourth pereiopods (
<figureCitation id="FB812A11C059855549D72BA5FA85B3E3" box="[1241,1325,1161,1187]" captionStart="FIGURE 2" captionStartId="4.[151,250,1689,1714]" captionTargetBox="[196,1395,200,1657]" captionTargetId="figure@4.[179,1409,181,1670]" captionTargetPageId="4" captionText="FIGURE 2. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—third maxilliped, lateral view; b—same, tip of ultimate article, lateral view; c—second pereiopod, lateral view; d—third pereiopod, lateral view; e—same, propodus and dactylus, mesial view; f—fourth pereiopod, lateral view; g—fifth pereiopod, lateral view." httpUri="https://zenodo.org/record/3359030/files/figure.png" pageId="2" pageNumber="53">
Fig. 
<quantity id="A4429B71C059855548032BA5FA85B3E3" box="[1293,1325,1161,1187]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="2" pageNumber="53" unit="g" value="2.0">2g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="63053694C05985554DC92B81FAFBB273" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="53">
Second pleopod with appendix masculina as long as appendix interna, furnished with numerous short stiff setae, as illustrated (
<figureCitation id="FB812A11C05985554C382BFDFE2CB3AC" box="[310,388,1233,1260]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="2" pageNumber="53">Fig. 1f</figureCitation>
). Uropod with lateral lobe of protopod ending in subacute tooth; exopod broadly ovoid, with small distolateral tooth and moderately developed distolateral spiniform seta; diaeresis sinuous, with broad subtriangular lobe mesial to spiniform seta; endopod as long as exopod, ovoid, without specific features (
<figureCitation id="FB812A11C059855549FC2A35FAECB273" box="[1266,1348,1305,1331]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1832]" captionTargetBox="[169,1414,285,1778]" captionTargetId="figure@3.[151,1436,273,1788]" captionTargetPageId="3" captionText="FIGURE 1. Salmoneus venustus sp. nov.: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view." httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="2" pageNumber="53">
Fig. 
<quantity id="A4429B71C059855548292A35FAECB273" box="[1319,1348,1305,1331]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="2" pageNumber="53" unit="g" value="1.0">1g</quantity>
</figureCitation>
).
</paragraph>
<paragraph id="63053694C05985554DC92A11FDB5B217" blockId="2.[151,1437,153,2015]" box="[199,541,1341,1367]" pageId="2" pageNumber="53">Gill formula typical for genus.</paragraph>
<paragraph id="63053694C05985554DC92A4CFE39B14B" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="53">
<emphasis id="51CEEA86C05985554DC92A4CFEF0B23B" bold="true" box="[199,344,1376,1403]" pageId="2" pageNumber="53">Colouration</emphasis>
. Semitransparent-whitish, with red chromatophores forming diffuse transverse bands on pleon, some red chromatophores scattered over selected areas of carapace; antennules, antennae and tail fan also with patches of red chromatophores; major chela hyaline-white; walking legs, antennular and antennal flagella colourless and semi-translucent; yellowish and brown-orange inner organs (including ovaries) visible through partly translucent carapace (
<figureCitation id="FB812A11C05985554C302ADDFE2BB14B" box="[318,387,1521,1547]" captionStart="FIGURE 4" captionStartId="6.[151,250,1894,1919]" captionTargetBox="[166,1421,166,1875]" captionTargetId="figure@6.[166,1421,166,1875]" captionTargetPageId="6" captionText="FIGURE 4. Salmoneus venustus sp. nov.: non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH. 2015-08-0031 [a, b]; holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [c]; a—living individual, dorsal view; b—same, lateral view (both photographs by the author); c—fresh specimen post-mortem, lateral view (photograph courtesy of Dr. Ivan N. Marin)." httpUri="https://zenodo.org/record/3359034/files/figure.png" pageId="2" pageNumber="53">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="63053694C05985554DC92939FD23B16F" blockId="2.[151,1437,153,2015]" box="[199,651,1556,1583]" pageId="2" pageNumber="53">
<materialsCitation id="D3D23CC9C05985554DC92939FD20B16F" ID-GBIF-Occurrence="2302320210" box="[199,648,1556,1583]" country="Vietnam" location="Nha Trang Bay" pageId="2" pageNumber="53" specimenCount="1" typeStatus="holotype">
<emphasis id="51CEEA86C05985554DC92939FEC8B16F" bold="true" box="[199,352,1556,1583]" pageId="2" pageNumber="53">
<typeStatus id="BC018836C05985554DC92939FEAAB16F" box="[199,258,1557,1583]" pageId="2" pageNumber="53">Type</typeStatus>
locality
</emphasis>
. 
<location id="6665604FC05985554C602939FDB2B16F" LSID="urn:lsid:plazi:treatment:EB138782C05A85504D992D82FAE5B2DF:6665604FC05985554C602939FDB2B16F" box="[366,538,1557,1583]" country="Vietnam" name="Nha Trang Bay" pageId="2" pageNumber="53">Nha Trang Bay</location>
, 
<collectingCountry id="1BAD7604C05985554F2B2939FD20B16F" box="[549,648,1557,1583]" name="Vietnam" pageId="2" pageNumber="53">Vietnam</collectingCountry>
</materialsCitation>
.
</paragraph>
<paragraph id="63053694C05985554DC92914FE17B137" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="53">
<emphasis id="51CEEA86C05985554DC92914FEF1B113" bold="true" box="[199,345,1592,1619]" pageId="2" pageNumber="53">Distribution</emphasis>
. Indo-West Pacific: currently only known from Nha Trang Bay in southern 
<collectingCountry id="1BAD7604C059855549A02915FAB8B113" box="[1198,1296,1593,1619]" name="Vietnam" pageId="2" pageNumber="53">Vietnam</collectingCountry>
, and west of 
<collectingCountry id="1BAD7604C05985554D992971FF41B137" box="[151,233,1629,1655]" name="Oman" pageId="2" pageNumber="53">Muscat</collectingCountry>
in northern 
<collectingCountry id="1BAD7604C05985554C7D2971FE13B137" box="[371,443,1629,1655]" name="Oman" pageId="2" pageNumber="53">Oman</collectingCountry>
.
</paragraph>
<paragraph id="63053694C05985554DC929ADFCA8B180" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="53">
<emphasis id="51CEEA86C05985554DC929ADFE8EB1DB" bold="true" box="[199,294,1665,1691]" pageId="2" pageNumber="53">Ecology</emphasis>
. Shallow mud-sand flats, from about 0.7 down to 
<quantity id="A4429B71C05985554E5529ADFC21B1DB" box="[859,905,1665,1691]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="53" unit="m" value="8.0">8 m</quantity>
; associated with burrows, presumably made by large ghost shrimps, such as 
<taxonomicName id="A4BA4D17C05985554CD72989FDE1B1FF" authorityName="Stimpson" authorityYear="1866" box="[473,585,1701,1727]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="51CEEA86C05985554CD72989FDE1B1FF" box="[473,585,1701,1727]" italics="true" pageId="2" pageNumber="53">Glypturus</emphasis>
</taxonomicName>
sp. (see below).
</paragraph>
<paragraph id="63053694C05985554DC129E5FE07B06B" blockId="2.[151,1437,153,2015]" pageId="2" pageNumber="53">
<emphasis id="51CEEA86C05985554DC129E5FEF8B1A3" bold="true" box="[207,336,1737,1763]" pageId="2" pageNumber="53">Etymology</emphasis>
. The new species’ name is the Latin adjective 
<taxonomicName id="A4BA4D17C05985554E6B29E6FC6DB1A3" authorityName="Anker" authorityYear="2019" box="[869,965,1738,1763]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="venustus">
<emphasis id="51CEEA86C05985554E6B29E6FC6DB1A3" box="[869,965,1738,1763]" italics="true" pageId="2" pageNumber="53">venustus</emphasis>
</taxonomicName>
, for graceful or attractive, alluding to the pretty reddish colouration of the new species (contrasting to many bland, rather whitish or pale-yellowish species in the genus 
<taxonomicName id="A4BA4D17C05985554C2B283DFE36B06B" authorityName="Holthuis" authorityYear="1955" box="[293,414,1809,1835]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="51CEEA86C05985554C2B283DFE36B06B" box="[293,414,1809,1835]" italics="true" pageId="2" pageNumber="53">Salmoneus</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="63053694C05985544DC92819FC08B7BB" blockId="2.[151,1437,153,2015]" lastBlockId="3.[151,1436,153,251]" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
<emphasis id="51CEEA86C05985554DC92819FE9CB00F" bold="true" box="[199,308,1845,1871]" pageId="2" pageNumber="53">Remarks</emphasis>
. 
<taxonomicName id="A4BA4D17C05985554C4C2819FD8AB00F" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[322,546,1845,1871]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05985554C4C2819FD8AB00F" box="[322,546,1845,1871]" italics="true" pageId="2" pageNumber="53">Salmoneus venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05985554F272819FD2AB00F" box="[553,642,1845,1871]" pageId="2" pageNumber="53" rank="species">
<emphasis id="51CEEA86C05985554F272819FD2AB00F" bold="true" box="[553,642,1845,1871]" pageId="2" pageNumber="53">sp. nov.</emphasis>
</taxonomicNameLabel>
is still distinguishable from all other species of 
<taxonomicName id="A4BA4D17C059855549992819FAB8B00F" authorityName="Holthuis" authorityYear="1955" box="[1175,1296,1845,1871]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="51CEEA86C059855549992819FAB8B00F" box="[1175,1296,1845,1871]" italics="true" pageId="2" pageNumber="53">Salmoneus</emphasis>
</taxonomicName>
by a combination of morphological characters and colouration. In the general shape of the frontal and dorsal areas of the carapace, including the very broad, laterally convex rostrum, the absence of a strong mid-dorsal carina, the absence of a dorsal depression, and the eyes being concealed in dorsal view, as well as in the general shape, degree of asymmetry and armature of the major and minor chelipeds, 
<taxonomicName id="A4BA4D17C05985554FB828E9FC9BB09F" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[694,819,1989,2015]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05985554FB828E9FC9BB09F" box="[694,819,1989,2015]" italics="true" pageId="2" pageNumber="53">S. venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05985554E3528E9FC3DB09F" box="[827,917,1989,2015]" pageId="2" pageNumber="53" rank="species">
<emphasis id="51CEEA86C05985554E3528E9FC3DB09F" bold="true" box="[827,917,1989,2015]" pageId="2" pageNumber="53">sp. nov.</emphasis>
</taxonomicNameLabel>
appears to be most similar to 
<taxonomicName id="A4BA4D17C059855449E728E9FEF9B7F3" authority="(Coutiere, 1897)" authorityName="Coutiere" authorityYear="1897" baseAuthorityName="Coutiere" baseAuthorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" lastPageId="3" lastPageNumber="54" order="Decapoda" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="51CEEA86C059855549E728E9FA34B09F" box="[1257,1436,1989,2015]" italics="true" pageId="2" pageNumber="53">S. serratidigitus</emphasis>
(
<bibRefCitation id="072B4B65C05885544D912FB5FEE2B7F3" author="Coutiere, H." box="[159,330,153,179]" pageId="3" pageNumber="54" pagination="380 - 386" refId="ref6646" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05885544C522FB5FD20B7F3" authority="(De Man, 1910)" authorityName="De Man" authorityYear="1910" baseAuthorityName="De Man" baseAuthorityYear="1910" box="[348,648,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="51CEEA86C05885544C522FB5FE66B7F3" box="[348,462,153,179]" italics="true" pageId="3" pageNumber="54">S. sibogae</emphasis>
(
<bibRefCitation id="072B4B65C05885544CD22FB5FD28B7F3" author="De Man, J. G." box="[476,640,153,179]" pageId="3" pageNumber="54" pagination="287 - 319" refId="ref6857" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05885544F9C2FB5FC72B7F3" authority="(Coutiere, 1897)" authorityName="Coutiere" authorityYear="1897" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[658,986,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05885544F9C2FB5FCB2B7F3" box="[658,794,153,179]" italics="true" pageId="3" pageNumber="54">S. latirostris</emphasis>
(
<bibRefCitation id="072B4B65C05885544E262FB5FC7BB7F3" author="Coutiere, H." box="[808,979,153,179]" pageId="3" pageNumber="54" pagination="380 - 386" refId="ref6646" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05885544EEB2FB5FA30B7F3" authority="Anker &amp; Marin, 2006" authorityName="Anker &amp; Marin" authorityYear="2006" box="[997,1432,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="nhatrangensis">
<emphasis id="51CEEA86C05885544EEB2FB5FB37B7F3" box="[997,1183,153,179]" italics="true" pageId="3" pageNumber="54">S. nhatrangensis</emphasis>
<bibRefCitation id="072B4B65C058855449AA2FB5FA30B7F3" author="Anker, A. &amp; Marin, I. N." box="[1188,1432,153,179]" pageId="3" pageNumber="54" pagination="295 - 319" refId="ref6175" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. The Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">Anker &amp; Marin, 2006</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05885544D992F91FDA6B797" authority="(Edmondson, 1930)" authorityName="Edmondson" authorityYear="1930" baseAuthorityName="Edmondson" baseAuthorityYear="1930" box="[151,526,189,215]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="mauiensis">
<emphasis id="51CEEA86C05885544D992F91FE8CB797" box="[151,292,189,215]" italics="true" pageId="3" pageNumber="54">S. mauiensis</emphasis>
(
<bibRefCitation id="072B4B65C05885544C3A2F91FDAFB797" author="Edmondson, C. H." box="[308,519,189,215]" pageId="3" pageNumber="54" pagination="1 - 18" refId="ref7138" refString="Edmondson, C. H. (1930) New Hawaiian Crustacea. Occasional Papers of the Bernice P. Bishop Museum, 9, 1 - 18, pl. 1." type="journal article" year="1930">Edmondson, 1930</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05885544F152F91FCEBB798" authority="(Schmitt, 1936)" authorityName="Schmitt" authorityYear="1936" baseAuthorityName="Schmitt" baseAuthorityYear="1936" box="[539,835,189,216]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="arubae">
<emphasis id="51CEEA86C05885544F152F91FD2FB797" box="[539,647,189,215]" italics="true" pageId="3" pageNumber="54">S. arubae</emphasis>
(
<bibRefCitation id="072B4B65C05885544F962F91FC94B798" author="Schmitt, W. L." box="[664,828,189,216]" pageId="3" pageNumber="54" pagination="363 - 378" refId="ref7612" refString="Schmitt, W. L. (1936) Zoologische Ergebnisse einer Reise nach Bonaire, Curacao und Aruba im Jahre 1930. No. 16. Macruran and anomuran Crustacea from Bonaire, Curacao und Aruba. Zoologische Jahrbu ¨ cher, 67, 363 - 378, pls. 11 - 13." type="journal article" year="1936">Schmitt, 1936</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05885544E5E2F91FB4AB797" authority="Manning &amp; Chace, 1990" authorityName="Manning &amp; Chace" authorityYear="1990" box="[848,1250,189,215]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="51CEEA86C05885544E5E2F91FC16B797" box="[848,958,189,215]" italics="true" pageId="3" pageNumber="54">S. setosus</emphasis>
<bibRefCitation id="072B4B65C05885544EC82F91FB4AB797" author="Manning, R. B. &amp; Chace, F. A. Jr." box="[966,1250,189,215]" pageId="3" pageNumber="54" pagination="1 - 91" refId="ref7384" refString="Manning, R. B. &amp; Chace, F. A. Jr. (1990) Decapod and stomatopod Crustacea from Ascension Island, South Atlantic Ocean. Smithsonian Contributions to Zoology, 503, 1 - 91. https: // doi. org / 10.5479 / si. 00810282.503" type="journal article" year="1990">Manning &amp; Chace, 1990</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C058855449E02F91FF7AB7BB" authority="Anker, 2007" authorityName="Anker" authorityYear="2007" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="rocas">
<emphasis id="51CEEA86C058855449E02F91FAE0B797" box="[1262,1352,189,215]" italics="true" pageId="3" pageNumber="54">S. rocas</emphasis>
Anker, 2007
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05885544DD32FCDFDE5B7BB" authority="Manning &amp; Chace, 1990" authorityName="Manning &amp; Chace" authorityYear="1990" box="[221,589,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="51CEEA86C05885544DD32FCDFE86B7BB" box="[221,302,225,251]" italics="true" pageId="3" pageNumber="54">S. teres</emphasis>
<bibRefCitation id="072B4B65C05885544C382FCDFDE5B7BB" author="Manning, R. B. &amp; Chace, F. A. Jr." box="[310,589,225,251]" pageId="3" pageNumber="54" pagination="1 - 91" refId="ref7384" refString="Manning, R. B. &amp; Chace, F. A. Jr. (1990) Decapod and stomatopod Crustacea from Ascension Island, South Atlantic Ocean. Smithsonian Contributions to Zoology, 503, 1 - 91. https: // doi. org / 10.5479 / si. 00810282.503" type="journal article" year="1990">Manning &amp; Chace, 1990</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="A4BA4D17C05885544F862FCDFC35B7BB" authority="Grippa, 2004" authorityName="Grippa" authorityYear="2004" box="[648,925,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="kekovae">
<emphasis id="51CEEA86C05885544F862FCDFD56B7BB" box="[648,766,225,251]" italics="true" pageId="3" pageNumber="54">S. kekovae</emphasis>
<bibRefCitation id="072B4B65C05885544E0B2FCDFC35B7BB" author="Grippa, G. B." box="[773,925,225,251]" pageId="3" pageNumber="54" pagination="45 - 52" refId="ref7227" refString="Grippa, G. B. (2004) Salmoneus kekovae, a new species of alpheid shrimp (Crustacea: Decapoda: Caridea) from the south-western coast of Turkey. Mediterranean Marine Science, 5, 45 - 52. https: // doi. org / 10.12681 / mms. 202" type="journal article" year="2004">Grippa, 2004</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="37C5661CC05885544D992823FCBCB0F8" httpUri="https://zenodo.org/record/3359028/files/figure.png" pageId="3" pageNumber="54" startId="3.[151,250,1807,1832]" targetBox="[169,1414,285,1778]" targetPageId="3">
<paragraph id="63053694C05885544D992823FCBCB0F8" blockId="3.[151,1437,1807,1976]" pageId="3" pageNumber="54">
<emphasis id="51CEEA86C05885544D992823FEBFB068" bold="true" box="[151,279,1807,1832]" pageId="3" pageNumber="54">FIGURE 1.</emphasis>
<taxonomicName id="A4BA4D17C05885544C2E283CFE47B067" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[288,495,1808,1832]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05885544C2E283CFE47B067" box="[288,495,1808,1832]" italics="true" pageId="3" pageNumber="54">Salmoneus venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05885544CF7283CFDE5B068" box="[505,589,1808,1832]" pageId="3" pageNumber="54" rank="species">
<emphasis id="51CEEA86C05885544CF7283CFDE5B068" bold="true" box="[505,589,1808,1832]" pageId="3" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [a–g]; non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH.ZC. 2015-08-0031 [h]; a—frontal region, dorsal view; b—same, lateral view; c—posterior pleonites, lateral view; d—telson, dorsal view; e—same, posterior margin, dorsal view; f—second pleopod, appendix interna and appendix masculina, lateral view; g—uropod, dorsal view; h—rostro-orbital area of carapace and eyes, lateral view.
</paragraph>
</caption>
<caption id="37C5661CC05F85534D9929B5FE76B05E" httpUri="https://zenodo.org/record/3359030/files/figure.png" pageId="4" pageNumber="55" startId="4.[151,250,1689,1714]" targetBox="[196,1395,200,1657]" targetPageId="4">
<paragraph id="63053694C05F85534D9929B5FE76B05E" blockId="4.[151,1437,1689,1822]" pageId="4" pageNumber="55">
<emphasis id="51CEEA86C05F85534D9929B5FEBEB1F2" bold="true" box="[151,278,1689,1714]" pageId="4" pageNumber="55">FIGURE 2.</emphasis>
<taxonomicName id="A4BA4D17C05F85534C2E29B6FE47B1F2" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[288,495,1690,1714]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05F85534C2E29B6FE47B1F2" box="[288,495,1690,1714]" italics="true" pageId="4" pageNumber="55">Salmoneus venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05F85534CF729B6FDE5B1F2" box="[505,589,1690,1714]" pageId="4" pageNumber="55" rank="species">
<emphasis id="51CEEA86C05F85534CF729B6FDE5B1F2" bold="true" box="[505,589,1690,1714]" pageId="4" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—third maxilliped, lateral view; b—same, tip of ultimate article, lateral view; c—second pereiopod, lateral view; d—third pereiopod, lateral view; e—same, propodus and dactylus, mesial view; f—fourth pereiopod, lateral view; g—fifth pereiopod, lateral view.
</paragraph>
</caption>
<subSubSection id="2BA0651FC05F85504DC92860FAE5B2DF" lastPageId="7" lastPageNumber="58" pageId="4" pageNumber="55" type="reference_group">
<paragraph id="63053694C05F85524DC92860FE39B523" blockId="4.[151,1437,1868,2039]" lastBlockId="5.[151,1437,153,611]" lastPageId="5" lastPageNumber="56" pageId="4" pageNumber="55">
<taxonomicName id="A4BA4D17C05F85534DC92860FE00B026" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[199,424,1868,1894]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05F85534DC92860FE00B026" box="[199,424,1868,1894]" italics="true" pageId="4" pageNumber="55">Salmoneus venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05F85534CBE2861FDA1B027" box="[432,521,1869,1895]" pageId="4" pageNumber="55" rank="species">
<emphasis id="51CEEA86C05F85534CBE2861FDA1B027" bold="true" box="[432,521,1869,1895]" pageId="4" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
may be separated from 
<taxonomicName id="A4BA4D17C05F85534E162861FC64B026" ID-CoL="85Y2S" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[792,972,1868,1895]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="51CEEA86C05F85534E162861FC64B026" box="[792,972,1868,1895]" italics="true" pageId="4" pageNumber="55">S. serratidigitus</emphasis>
</taxonomicName>
, 
<emphasis id="51CEEA86C05F85534ED42861FB76B026" box="[986,1246,1868,1895]" italics="true" pageId="4" pageNumber="55">
<taxonomicName id="A4BA4D17C05F85534ED42861FBE5B026" ID-CoL="85Y2W" baseAuthorityName="De Man" baseAuthorityYear="1910" box="[986,1101,1868,1895]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="sibogae">S. sibogae</taxonomicName>
<taxonomicName id="A4BA4D17C05F8553495B2861FB76B026" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[1109,1246,1868,1895]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="latirostris">S. latirostris</taxonomicName>
</emphasis>
, 
<taxonomicName id="A4BA4D17C05F855349E32861FAD1B026" ID-CoL="854PQ" baseAuthorityName="Edmondson" baseAuthorityYear="1930" box="[1261,1401,1869,1895]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="mauiensis">
<emphasis id="51CEEA86C05F855349E32861FAD1B026" box="[1261,1401,1869,1895]" italics="true" pageId="4" pageNumber="55">S. mauiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F855348892861FE9EB0CA" ID-CoL="85Y2J" authorityName="Anker &amp; Marin" authorityYear="2006" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="nhatrangensis">
<emphasis id="51CEEA86C05F855348892861FE9EB0CA" italics="true" pageId="4" pageNumber="55">S. nhatrangensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534C4D285DFE06B0CA" ID-CoL="85XZQ" baseAuthorityName="Schmitt" baseAuthorityYear="1936" box="[323,430,1904,1931]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="arubae">
<emphasis id="51CEEA86C05F85534C4D285DFE06B0CA" box="[323,430,1904,1931]" italics="true" pageId="4" pageNumber="55">S. arubae</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534CB2285DFD80B0CA" ID-CoL="85Y2V" authorityName="Manning &amp; Chace" authorityYear="1990" box="[444,552,1905,1931]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="51CEEA86C05F85534CB2285DFD80B0CA" box="[444,552,1905,1931]" italics="true" pageId="4" pageNumber="55">S. setosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534F38285DFD26B0CA" ID-CoL="85Y2Q" authorityName="Anker" authorityYear="2007" box="[566,654,1905,1931]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="rocas">
<emphasis id="51CEEA86C05F85534F38285DFD26B0CA" box="[566,654,1905,1931]" italics="true" pageId="4" pageNumber="55">S. rocas</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534F92285DFD44B0CA" ID-CoL="85Y32" authorityName="Manning &amp; Chace" authorityYear="1990" box="[668,748,1905,1931]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="51CEEA86C05F85534F92285DFD44B0CA" box="[668,748,1905,1931]" italics="true" pageId="4" pageNumber="55">S. teres</emphasis>
</taxonomicName>
, and 
<taxonomicName id="A4BA4D17C05F85534E27285DFC37B0CA" ID-CoL="85Y2D" authorityName="Grippa" authorityYear="2004" box="[809,927,1904,1931]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="kekovae">
<emphasis id="51CEEA86C05F85534E27285DFC37B0CA" box="[809,927,1904,1931]" italics="true" pageId="4" pageNumber="55">S. kekovae</emphasis>
</taxonomicName>
by the longer second article of the antennular peduncle, being 1.3 times as long as wide (vs. as long as wide or wider than long in the other species); from 
<taxonomicName id="A4BA4D17C05F8553485B28B9FEA4B092" ID-CoL="85Y2S" baseAuthorityName="Coutiere" baseAuthorityYear="1897" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="51CEEA86C05F8553485B28B9FEA4B092" italics="true" pageId="4" pageNumber="55">S. serratidigitus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534C142895FE25B092" ID-CoL="85Y2W" baseAuthorityName="De Man" baseAuthorityYear="1910" box="[282,397,1976,2003]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="51CEEA86C05F85534C142895FE25B092" box="[282,397,1976,2003]" italics="true" pageId="4" pageNumber="55">S. sibogae</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534C922895FD8DB092" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[412,549,1976,2003]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05F85534C922895FD8DB092" box="[412,549,1976,2003]" italics="true" pageId="4" pageNumber="55">S. latirostris</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534F3A2895FD68B092" ID-CoL="854PQ" baseAuthorityName="Edmondson" baseAuthorityYear="1930" box="[564,704,1977,2003]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="mauiensis">
<emphasis id="51CEEA86C05F85534F3A2895FD68B092" box="[564,704,1977,2003]" italics="true" pageId="4" pageNumber="55">S. mauiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534FC02895FC80B092" ID-CoL="85Y2Q" authorityName="Anker" authorityYear="2007" box="[718,808,1977,2003]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="rocas">
<emphasis id="51CEEA86C05F85534FC02895FC80B092" box="[718,808,1977,2003]" italics="true" pageId="4" pageNumber="55">S. rocas</emphasis>
</taxonomicName>
, and 
<taxonomicName id="A4BA4D17C05F85534E682895FC74B092" ID-CoL="85Y2D" authorityName="Grippa" authorityYear="2004" box="[870,988,1976,2003]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="kekovae">
<emphasis id="51CEEA86C05F85534E682895FC74B092" box="[870,988,1976,2003]" italics="true" pageId="4" pageNumber="55">S. kekovae</emphasis>
</taxonomicName>
by the much shallower median incision on the posterior margin of the telson; from 
<taxonomicName id="A4BA4D17C05F85534F7F28F1FC8BB0B6" ID-CoL="85Y2S" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[625,803,2012,2039]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="51CEEA86C05F85534F7F28F1FC8BB0B6" box="[625,803,2012,2039]" italics="true" pageId="4" pageNumber="55">S. serratidigitus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534E3E28F1FC0AB0B6" ID-CoL="85Y2W" baseAuthorityName="De Man" baseAuthorityYear="1910" box="[816,930,2012,2039]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="51CEEA86C05F85534E3E28F1FC0AB0B6" box="[816,930,2012,2039]" italics="true" pageId="4" pageNumber="55">S. sibogae</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F85534EA128F1FB9FB0B6" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[943,1079,2012,2039]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05F85534EA128F1FB9FB0B6" box="[943,1079,2012,2039]" italics="true" pageId="4" pageNumber="55">S. latirostris</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F8553494A28F1FB56B0B6" ID-CoL="85Y2J" authorityName="Anker &amp; Marin" authorityYear="2006" box="[1092,1278,2012,2039]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="nhatrangensis">
<emphasis id="51CEEA86C05F8553494A28F1FB56B0B6" box="[1092,1278,2012,2039]" italics="true" pageId="4" pageNumber="55">S. nhatrangensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05F8553480428F1FA3DB0B6" ID-CoL="854PQ" baseAuthorityName="Edmondson" baseAuthorityYear="1930" box="[1290,1429,2013,2039]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="mauiensis">
<emphasis id="51CEEA86C05F8553480428F1FA3DB0B6" box="[1290,1429,2013,2039]" italics="true" pageId="4" pageNumber="55">S. mauiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05E85524D992FB5FEA9B7F3" ID-CoL="85XZQ" baseAuthorityName="Schmitt" baseAuthorityYear="1936" box="[151,257,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="arubae">
<emphasis id="51CEEA86C05E85524D992FB5FEA9B7F3" box="[151,257,153,179]" italics="true" pageId="5" pageNumber="56">S. arubae</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05E85524C002FB5FED2B7F3" ID-CoL="85Y2V" authorityName="Manning &amp; Chace" authorityYear="1990" box="[270,378,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="51CEEA86C05E85524C002FB5FED2B7F3" box="[270,378,153,179]" italics="true" pageId="5" pageNumber="56">S. setosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05E85524C892FB5FE77B7F3" ID-CoL="85Y2Q" authorityName="Anker" authorityYear="2007" box="[391,479,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="rocas">
<emphasis id="51CEEA86C05E85524C892FB5FE77B7F3" box="[391,479,153,179]" italics="true" pageId="5" pageNumber="56">S. rocas</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05E85524CE22FB5FD93B7F3" ID-CoL="85Y32" authorityName="Manning &amp; Chace" authorityYear="1990" box="[492,571,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="51CEEA86C05E85524CE22FB5FD93B7F3" box="[492,571,153,179]" italics="true" pageId="5" pageNumber="56">S. teres</emphasis>
</taxonomicName>
and 
<taxonomicName id="A4BA4D17C05E85524F7E2FB5FD4DB7F3" ID-CoL="85Y2D" authorityName="Grippa" authorityYear="2004" box="[624,741,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="kekovae">
<emphasis id="51CEEA86C05E85524F7E2FB5FD4DB7F3" box="[624,741,153,179]" italics="true" pageId="5" pageNumber="56">S. kekovae</emphasis>
</taxonomicName>
by the distinctly shorter stylocerite, reaching to the mid-length of the second article of the antennular peduncle (vs. at least to the distal margin of the second and often to the midlength of the third article in the other species); from 
<taxonomicName id="A4BA4D17C05E85524FE92FCDFC33B7BB" ID-CoL="85Y2S" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[743,923,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="51CEEA86C05E85524FE92FCDFC33B7BB" box="[743,923,225,251]" italics="true" pageId="5" pageNumber="56">S. serratidigitus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05E85524EA72FCDFBB5B7BB" ID-CoL="85Y2W" baseAuthorityName="De Man" baseAuthorityYear="1910" box="[937,1053,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="51CEEA86C05E85524EA72FCDFBB5B7BB" box="[937,1053,225,251]" italics="true" pageId="5" pageNumber="56">S. sibogae</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05E855249252FCDFB10B7BB" ID-CoL="854PQ" baseAuthorityName="Edmondson" baseAuthorityYear="1930" box="[1067,1208,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="mauiensis">
<emphasis id="51CEEA86C05E855249252FCDFB10B7BB" box="[1067,1208,225,251]" italics="true" pageId="5" pageNumber="56">S. mauiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05E855249C82FCDFA9CB7BB" ID-CoL="85Y2V" authorityName="Manning &amp; Chace" authorityYear="1990" box="[1222,1332,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="51CEEA86C05E855249C82FCDFA9CB7BB" box="[1222,1332,225,251]" italics="true" pageId="5" pageNumber="56">S. setosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05E8552484C2FCDFA34B7BB" ID-CoL="85Y2Q" authorityName="Anker" authorityYear="2007" box="[1346,1436,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="rocas">
<emphasis id="51CEEA86C05E8552484C2FCDFA34B7BB" box="[1346,1436,225,251]" italics="true" pageId="5" pageNumber="56">S. rocas</emphasis>
</taxonomicName>
and 
<taxonomicName id="A4BA4D17C05E85524DC92E29FE95B65F" ID-CoL="85Y2D" authorityName="Grippa" authorityYear="2004" box="[199,317,261,287]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="kekovae">
<emphasis id="51CEEA86C05E85524DC92E29FE95B65F" box="[199,317,261,287]" italics="true" pageId="5" pageNumber="56">S. kekovae</emphasis>
</taxonomicName>
by the shape of the rostrum and size and direction of the extra-corneal teeth; from 
<taxonomicName id="A4BA4D17C05E855249EF2E29FA35B65F" ID-CoL="85Y2J" authorityName="Anker &amp; Marin" authorityYear="2006" box="[1249,1437,261,287]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="nhatrangensis">
<emphasis id="51CEEA86C05E855249EF2E29FA35B65F" box="[1249,1437,261,287]" italics="true" pageId="5" pageNumber="56">S. nhatrangensis</emphasis>
</taxonomicName>
by the much shallower rostro-orbital notches; from 
<taxonomicName id="A4BA4D17C05E85524FD02E05FCE2B603" ID-CoL="85XZQ" baseAuthorityName="Schmitt" baseAuthorityYear="1936" box="[734,842,297,323]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="arubae">
<emphasis id="51CEEA86C05E85524FD02E05FCE2B603" box="[734,842,297,323]" italics="true" pageId="5" pageNumber="56">S. arubae</emphasis>
</taxonomicName>
by the much longer minor cheliped carpus and third pereiopod dactylus; from 
<taxonomicName id="A4BA4D17C05E85524CB92E61FD8BB627" ID-CoL="85Y2V" authorityName="Manning &amp; Chace" authorityYear="1990" box="[439,547,333,359]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="51CEEA86C05E85524CB92E61FD8BB627" box="[439,547,333,359]" italics="true" pageId="5" pageNumber="56">S. setosus</emphasis>
</taxonomicName>
by the absence of numerous thickened setae on the carapace and pleon; from 
<taxonomicName id="A4BA4D17C05E855248892E61FF64B6CB" ID-CoL="85Y32" authorityName="Manning &amp; Chace" authorityYear="1990" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="teres">
<emphasis id="51CEEA86C05E855248892E61FF64B6CB" italics="true" pageId="5" pageNumber="56">S. teres</emphasis>
</taxonomicName>
in the absence of a spiniform seta on the ischium of the second pereiopod; and from 
<taxonomicName id="A4BA4D17C05E8552499C2E5DFAA0B6CB" ID-CoL="85Y2D" authorityName="Grippa" authorityYear="2004" box="[1170,1288,369,395]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="kekovae">
<emphasis id="51CEEA86C05E8552499C2E5DFAA0B6CB" box="[1170,1288,369,395]" italics="true" pageId="5" pageNumber="56">S. kekovae</emphasis>
</taxonomicName>
by the much less swollen major chela (cf. 
<bibRefCitation id="072B4B65C05E85524CD72EB9FDD2B6EF" author="Coutiere, H." box="[473,634,405,431]" pageId="5" pageNumber="56" pagination="380 - 386" refId="ref6646" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle 2, 380 - 386." type="journal article" year="1897">Coutière 1897</bibRefCitation>
, 
<bibRefCitation id="072B4B65C05E85524F8A2EB9FD68B6EF" author="Coutiere, H." box="[644,704,405,431]" pageId="5" pageNumber="56" pagination="1 - 559" refId="ref6680" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, 6 pls. 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">1899</bibRefCitation>
: fig. 21; 
<bibRefCitation id="072B4B65C05E85524E2E2EB9FC13B6EF" author="De Man, J. G." box="[800,955,405,431]" pageId="5" pageNumber="56" pagination="133 - 465" refId="ref6896" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga Expeditie, 39 a 1, 133 - 465; (1915) Supplement-Explanations of plates of Alpheidae. E. J. Brill, Leiden, 23 pls. https: // doi. org / 10.5962 / bhl. title. 10512" type="book chapter" year="1911">De Man 1911</bibRefCitation>
: fig. 10; 
<bibRefCitation id="072B4B65C05E855249152EB9FB1CB6EF" author="Schmitt, W. L." box="[1051,1204,405,431]" pageId="5" pageNumber="56" pagination="363 - 378" refId="ref7612" refString="Schmitt, W. L. (1936) Zoologische Ergebnisse einer Reise nach Bonaire, Curacao und Aruba im Jahre 1930. No. 16. Macruran and anomuran Crustacea from Bonaire, Curacao und Aruba. Zoologische Jahrbu ¨ cher, 67, 363 - 378, pls. 11 - 13." type="journal article" year="1936">Schmitt 1936</bibRefCitation>
: fig. 2; Banner 1953: fig. 2; 
<bibRefCitation id="072B4B65C05E85524DEC2E95FE44B693" author="Banner, D. M. &amp; Banner, A. H." box="[226,492,441,467]" pageId="5" pageNumber="56" pagination="1 - 99" refId="ref6502" refString="Banner, D. M. &amp; Banner, A. H. (1981). Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
: fig. 8; 
<bibRefCitation id="072B4B65C05E85524F4C2E95FCFDB693" author="Manning, R. B. &amp; Chace, F. A. Jr." box="[578,853,441,467]" pageId="5" pageNumber="56" pagination="1 - 91" refId="ref7384" refString="Manning, R. B. &amp; Chace, F. A. Jr. (1990) Decapod and stomatopod Crustacea from Ascension Island, South Atlantic Ocean. Smithsonian Contributions to Zoology, 503, 1 - 91. https: // doi. org / 10.5479 / si. 00810282.503" type="journal article" year="1990">Manning &amp; Chace 1990</bibRefCitation>
: figs. 9, 10; 
<bibRefCitation id="072B4B65C05E85524EEE2E95FBD9B693" author="Grippa, G. B." box="[992,1137,441,467]" pageId="5" pageNumber="56" pagination="45 - 52" refId="ref7227" refString="Grippa, G. B. (2004) Salmoneus kekovae, a new species of alpheid shrimp (Crustacea: Decapoda: Caridea) from the south-western coast of Turkey. Mediterranean Marine Science, 5, 45 - 52. https: // doi. org / 10.12681 / mms. 202" type="journal article" year="2004">Grippa 2004</bibRefCitation>
: figs. 1, 2; 
<bibRefCitation id="072B4B65C05E855249E32E95FF7BB6B7" author="Anker, A. &amp; Marin, I. N." pageId="5" pageNumber="56" pagination="295 - 319" refId="ref6175" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. The Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">Anker &amp; Marin 2006</bibRefCitation>
: fig. 2; Anker 2007: fig. 4). In addition, the presence of a strong distoventral process on the major cheliped carpus, the absence of spiniform setae on the fourth pereiopod, and the eyestalks with reduced cornea and pointed anteromesial tubercle, combined with several other characters, separate 
<taxonomicName id="A4BA4D17C05E85524ECC2D09FB97B57F" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[962,1087,549,575]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05E85524ECC2D09FB97B57F" box="[962,1087,549,575]" italics="true" pageId="5" pageNumber="56">S. venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05E855249492D09FB09B57F" box="[1095,1185,549,575]" pageId="5" pageNumber="56" rank="species">
<emphasis id="51CEEA86C05E855249492D09FB09B57F" bold="true" box="[1095,1185,549,575]" pageId="5" pageNumber="56">sp. nov.</emphasis>
</taxonomicNameLabel>
from most of the species mentioned above.
</paragraph>
<caption id="37C5661CC05E85524D9928B8FA2FB0B5" httpUri="https://zenodo.org/record/3359032/files/figure.png" pageId="5" pageNumber="56" startId="5.[151,250,1940,1965]" targetBox="[189,1372,641,1888]" targetPageId="5">
<paragraph id="63053694C05E85524D9928B8FA2FB0B5" blockId="5.[151,1437,1940,2037]" pageId="5" pageNumber="56">
<emphasis id="51CEEA86C05E85524D9928B8FEBEB0ED" bold="true" box="[151,278,1940,1965]" pageId="5" pageNumber="56">FIGURE 3.</emphasis>
<taxonomicName id="A4BA4D17C05E85524C2E28B9FE47B0ED" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[288,495,1941,1965]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05E85524C2E28B9FE47B0ED" box="[288,495,1941,1965]" italics="true" pageId="5" pageNumber="56">Salmoneus venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05E85524CF728B9FDE5B0ED" box="[505,589,1941,1965]" pageId="5" pageNumber="56" rank="species">
<emphasis id="51CEEA86C05E85524CF728B9FDE5B0ED" bold="true" box="[505,589,1941,1965]" pageId="5" pageNumber="56">sp. nov.</emphasis>
</taxonomicNameLabel>
: holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290; a—major cheliped, lateral view; b—same, mesial view; c—same, carpus and chela, ventrolateral view; d—same, chela fingers opened, ventrolateral view; e—minor cheliped, lateral view; f—same, carpus and chela, mesial view.
</paragraph>
</caption>
<caption id="37C5661CC05D85514D99284AFDFFB0AB" httpUri="https://zenodo.org/record/3359034/files/figure.png" pageId="6" pageNumber="57" startId="6.[151,250,1894,1919]" targetBox="[166,1421,166,1875]" targetPageId="6">
<paragraph id="63053694C05D85514D99284AFDFFB0AB" blockId="6.[151,1437,1894,2027]" pageId="6" pageNumber="57">
<emphasis id="51CEEA86C05D85514D99284AFEBBB03F" bold="true" box="[151,275,1894,1919]" pageId="6" pageNumber="57">FIGURE 4.</emphasis>
<taxonomicName id="A4BA4D17C05D85514C17284BFE4CB03F" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[281,484,1895,1919]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05D85514C17284BFE4CB03F" box="[281,484,1895,1919]" italics="true" pageId="6" pageNumber="57">Salmoneus venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05D85514CE4284BFD92B03F" box="[490,570,1895,1919]" pageId="6" pageNumber="57" rank="species">
<emphasis id="51CEEA86C05D85514CE4284BFD92B03F" bold="true" box="[490,570,1895,1919]" pageId="6" pageNumber="57">sp. nov.</emphasis>
</taxonomicNameLabel>
: non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH. 2015-08-0031 [a, b]; holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [c]; a—living individual, dorsal view; b—same, lateral view (both photographs by the author); c—fresh specimen 
<emphasis id="51CEEA86C05D85514999289CFABFB087" box="[1175,1303,1968,1991]" italics="true" pageId="6" pageNumber="57">post-mortem</emphasis>
, lateral view (photograph courtesy of Dr. Ivan N. Marin).
</paragraph>
</caption>
<paragraph id="63053694C05C85504DC92FB5FC13B6EF" blockId="7.[151,1437,153,1439]" pageId="7" pageNumber="58">
The characteristic colour pattern of 
<taxonomicName id="A4BA4D17C05C85504F512FB5FD74B7F3" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[607,732,153,179]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05C85504F512FB5FD74B7F3" box="[607,732,153,179]" italics="true" pageId="7" pageNumber="58">S. venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05C85504FEB2FB5FCE8B7F3" box="[741,832,153,179]" pageId="7" pageNumber="58" rank="species">
<emphasis id="51CEEA86C05C85504FEB2FB5FCE8B7F3" bold="true" box="[741,832,153,179]" pageId="7" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
, which basically consists of diffuse red bands on the pleon (
<figureCitation id="FB812A11C05C85504DEB2F91FE82B797" box="[229,298,189,215]" captionStart="FIGURE 4" captionStartId="6.[151,250,1894,1919]" captionTargetBox="[166,1421,166,1875]" captionTargetId="figure@6.[166,1421,166,1875]" captionTargetPageId="6" captionText="FIGURE 4. Salmoneus venustus sp. nov.: non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH. 2015-08-0031 [a, b]; holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [c]; a—living individual, dorsal view; b—same, lateral view (both photographs by the author); c—fresh specimen post-mortem, lateral view (photograph courtesy of Dr. Ivan N. Marin)." httpUri="https://zenodo.org/record/3359034/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
), allows an easy separation of living individuals of the new species from those of 
<taxonomicName id="A4BA4D17C05C855049C82F91FAD1B797" ID-CoL="85Y2S" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[1222,1401,189,215]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="serratidigitus">
<emphasis id="51CEEA86C05C855049C82F91FAD1B797" box="[1222,1401,189,215]" italics="true" pageId="7" pageNumber="58">S. serratidigitus</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05C855048892F91FEAFB7BB" ID-CoL="854PQ" baseAuthorityName="Edmondson" baseAuthorityYear="1930" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="mauiensis">
<emphasis id="51CEEA86C05C855048892F91FEAFB7BB" italics="true" pageId="7" pageNumber="58">S. mauiensis</emphasis>
</taxonomicName>
(both uniformly bright yellow-orange), 
<taxonomicName id="A4BA4D17C05C85504FC32FCDFCE8B7BB" ID-CoL="85Y2W" baseAuthorityName="De Man" baseAuthorityYear="1910" box="[717,832,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="sibogae">
<emphasis id="51CEEA86C05C85504FC32FCDFCE8B7BB" box="[717,832,225,251]" italics="true" pageId="7" pageNumber="58">S. sibogae</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4BA4D17C05C85504E412FCDFBA2B7BB" ID-CoL="85Y2J" authorityName="Anker &amp; Marin" authorityYear="2006" box="[847,1034,225,251]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="nhatrangensis">
<emphasis id="51CEEA86C05C85504E412FCDFBA2B7BB" box="[847,1034,225,251]" italics="true" pageId="7" pageNumber="58">S. nhatrangensis</emphasis>
</taxonomicName>
(both whitish or pale yellowish), 
<taxonomicName id="A4BA4D17C05C855048892FCDFF40B65F" ID-CoL="85Y2V" authorityName="Manning &amp; Chace" authorityYear="1990" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="setosus">
<emphasis id="51CEEA86C05C855048892FCDFF40B65F" italics="true" pageId="7" pageNumber="58">S. setosus</emphasis>
</taxonomicName>
(bright yellow) and 
<taxonomicName id="A4BA4D17C05C85504CC52E29FDE9B65F" ID-CoL="85Y2D" authorityName="Grippa" authorityYear="2004" box="[459,577,261,287]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="kekovae">
<emphasis id="51CEEA86C05C85504CC52E29FDE9B65F" box="[459,577,261,287]" italics="true" pageId="7" pageNumber="58">S. kekovae</emphasis>
</taxonomicName>
(pink-orange) (cf. 
<bibRefCitation id="072B4B65C05C85504E1D2E29FC1DB65F" author="Coutiere, H." box="[787,949,261,287]" pageId="7" pageNumber="58" pagination="1 - 559" refId="ref6680" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, 6 pls. 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">Coutière 1899</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C85504ECE2E29FB2FB65F" author="Edmondson, C. H." box="[960,1159,261,287]" pageId="7" pageNumber="58" pagination="1 - 18" refId="ref7138" refString="Edmondson, C. H. (1930) New Hawaiian Crustacea. Occasional Papers of the Bernice P. Bishop Museum, 9, 1 - 18, pl. 1." type="journal article" year="1930">Edmondson 1930</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C8550499C2E29FA3FB65F" author="Banner, D. M. &amp; Banner, A. H." box="[1170,1431,261,287]" pageId="7" pageNumber="58" pagination="1 - 99" refId="ref6502" refString="Banner, D. M. &amp; Banner, A. H. (1981). Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C85504D992E05FE8FB603" author="Grippa, G. B." box="[151,295,297,323]" pageId="7" pageNumber="58" pagination="45 - 52" refId="ref7227" refString="Grippa, G. B. (2004) Salmoneus kekovae, a new species of alpheid shrimp (Crustacea: Decapoda: Caridea) from the south-western coast of Turkey. Mediterranean Marine Science, 5, 45 - 52. https: // doi. org / 10.12681 / mms. 202" type="journal article" year="2004">Grippa 2004</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C85504C3E2E05FDB7B603" author="Anker, A. &amp; Marin, I. N." box="[304,543,297,323]" pageId="7" pageNumber="58" pagination="295 - 319" refId="ref6175" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. The Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">Anker &amp; Marin 2006</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C85504F262E05FD47B604" author="Anker, A. &amp; Vera Caripe, J." box="[552,751,297,324]" pageId="7" pageNumber="58" pagination="185 - 191" refId="ref6345" refString="Anker, A. &amp; Vera Caripe, J. (2016) Leptalpheus pereirai sp. nov., a new alphied shrimp from Panama and Venezuela (Decapoda: Caridea). Zootaxa, 4127 (1), 185 - 191. https: // doi. org / 10.11646 / zootaxa. 4127.1.11" type="journal article" year="2016">
Anker 
<emphasis id="51CEEA86C05C85504F7A2E06FD04B603" box="[628,684,297,323]" italics="true" pageId="7" pageNumber="58">et al.</emphasis>
2016
</bibRefCitation>
). The only other known brightly transversely red-banded species of 
<taxonomicName id="A4BA4D17C05C85504DE92E61FEC8B627" ID-CoL="84TDH" authorityName="Holthuis" authorityYear="1955" box="[231,352,333,359]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="51CEEA86C05C85504DE92E61FEC8B627" box="[231,352,333,359]" italics="true" pageId="7" pageNumber="58">Salmoneus</emphasis>
</taxonomicName>
are 
<taxonomicName id="A4BA4D17C05C85504C9E2E61FDB0B627" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[400,536,333,359]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05C85504C9E2E61FDB0B627" box="[400,536,333,359]" italics="true" pageId="7" pageNumber="58">S. latirostris</emphasis>
</taxonomicName>
and 
<taxonomicName id="A4BA4D17C05C85504F402E61FC24B627" ID-CoL="85Y22" authority="(Coutiere, 1897)" authorityName="Coutiere" authorityYear="1897" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[590,908,333,359]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="51CEEA86C05C85504F402E61FD62B627" box="[590,714,333,359]" italics="true" pageId="7" pageNumber="58">S. cristatus</emphasis>
(
<bibRefCitation id="072B4B65C05C85504FD72E61FC2CB627" author="Coutiere, H." box="[729,900,333,359]" pageId="7" pageNumber="58" pagination="380 - 386" refId="ref6646" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
)
</taxonomicName>
, although some other species of the genus have darker brown-purple transverse bands (see below) or display red chromatophores on the carapace only (
<bibRefCitation id="072B4B65C05C855048192E5DFF79B6EF" author="Anker, A. &amp; Firdaus, M. &amp; Pratama, I. S." pageId="7" pageNumber="58" pagination="489 - 495" refId="ref6047" refString="Anker, A., Firdaus, M. &amp; Pratama, I. S. (2014) Salmoneus yoyo nov. sp., a peculiar new infaunal shrimp from Lombok, Indonesia (Decapoda, Caridea, Alpheidae). Zootaxa, 3852 (4), 489 - 495. https: // doi. org / 10.11646 / zootaxa. 3852.4.6" type="journal article" year="2014">
Anker 
<emphasis id="51CEEA86C05C8550486B2E5EFA35B6CB" box="[1381,1437,369,395]" italics="true" pageId="7" pageNumber="58">et al.</emphasis>
2014
</bibRefCitation>
) or on the carapace and the first two pleonites (Anker 
<emphasis id="51CEEA86C05C85504E3D2EBAFCC3B6EF" box="[819,875,405,431]" italics="true" pageId="7" pageNumber="58">et al.</emphasis>
2015).
</paragraph>
<paragraph id="63053694C05C85504DC92E95FD86B378" blockId="7.[151,1437,153,1439]" pageId="7" pageNumber="58">
<taxonomicName id="A4BA4D17C05C85504DC92E95FE1BB693" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[199,435,441,467]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05C85504DC92E95FE1BB693" box="[199,435,441,467]" italics="true" pageId="7" pageNumber="58">Salmoneus latirostris</emphasis>
</taxonomicName>
was described by 
<bibRefCitation id="072B4B65C05C85504F732E95FC80B693" author="Coutiere, H." box="[637,808,441,467]" pageId="7" pageNumber="58" pagination="380 - 386" refId="ref6646" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle 2, 380 - 386." type="journal article" year="1897">Coutière (1897</bibRefCitation>
, 
<bibRefCitation id="072B4B65C05C85504E3F2E95FCC2B693" author="Coutiere, H." box="[817,874,441,467]" pageId="7" pageNumber="58" pagination="1 - 559" refId="ref6680" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, 6 pls. 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">1899</bibRefCitation>
) as “incolore, corps régulièrement annelé de rouge vif” (= colourless, with regular intense transverse red banding), and by L.B. Holthuis in 
<bibRefCitation id="072B4B65C05C855049672EF1FAD6B6B7" author="Banner, D. M. &amp; Banner, A. H." box="[1129,1406,477,503]" pageId="7" pageNumber="58" pagination="1 - 99" refId="ref6502" refString="Banner, D. M. &amp; Banner, A. H. (1981). Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner (1981)</bibRefCitation>
as red banded on the pleon, with one red band at the middle or the end of the tail fan and three transverse rows of red chromatophores or red lines on the carapace, and with the antennular and antennal peduncles red or with red spots (with one specimen having a distinct red spot on the scaphocerite). According to the brief notes in 
<bibRefCitation id="072B4B65C05C855049E22D65FA30B523" author="Coutiere, H." box="[1260,1432,585,611]" pageId="7" pageNumber="58" pagination="380 - 386" refId="ref6646" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle 2, 380 - 386." type="journal article" year="1897">Coutière (1897</bibRefCitation>
, 
<bibRefCitation id="072B4B65C05C85504D992D41FF7BB5C7" author="Coutiere, H." box="[151,211,621,647]" pageId="7" pageNumber="58" pagination="1 - 559" refId="ref6680" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, 6 pls. 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">1899</bibRefCitation>
), 
<taxonomicName id="A4BA4D17C05C85504DEB2D41FEC8B5C7" ID-CoL="85Y22" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[229,352,621,647]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="51CEEA86C05C85504DEB2D41FEC8B5C7" box="[229,352,621,647]" italics="true" pageId="7" pageNumber="58">S. cristatus</emphasis>
</taxonomicName>
is also red-banded, similar to 
<taxonomicName id="A4BA4D17C05C85504FA22D41FC9CB5C7" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[684,820,621,647]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05C85504FA22D41FC9CB5C7" box="[684,820,621,647]" italics="true" pageId="7" pageNumber="58">S. latirostris</emphasis>
</taxonomicName>
(“coloration comme chez 
<taxonomicName id="A4BA4D17C05C855049572D41FB49B5C7" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[1113,1249,621,647]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05C855049572D41FB49B5C7" box="[1113,1249,621,647]" italics="true" pageId="7" pageNumber="58">S. latirostris</emphasis>
</taxonomicName>
”). L.B. Holthuis in 
<bibRefCitation id="072B4B65C05C85504DBD2DBDFE6DB5EB" author="Banner, D. M. &amp; Banner, A. H." box="[179,453,657,683]" pageId="7" pageNumber="58" pagination="1 - 99" refId="ref6502" refString="Banner, D. M. &amp; Banner, A. H. (1981). Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner (1981)</bibRefCitation>
supplied more detailed colour notes for 
<taxonomicName id="A4BA4D17C05C85504E8F2DBDFC54B5EB" ID-CoL="85Y22" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[897,1020,657,683]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="51CEEA86C05C85504E8F2DBDFC54B5EB" box="[897,1020,657,683]" italics="true" pageId="7" pageNumber="58">S. cristatus</emphasis>
</taxonomicName>
, which can be summarised as following: body transparent, whitish; carapace with short median red line extending from the rostral tip to slightly behind the rostral base; posterior portion of the carapace with transverse red line, its lateral ends curving slightly forward; third paired red lines extending ventrally from near the eyes; second to sixth pleonites each with distinct red line running from bases of pleura along posterior margin of tergum; uropodal endopod with one red spot basally; antennular peduncle with some red spots; eggs orange red. Based on the above colour descriptions, both 
<taxonomicName id="A4BA4D17C05C855049EB2C69FAC5B41F" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[1253,1389,837,863]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05C855049EB2C69FAC5B41F" box="[1253,1389,837,863]" italics="true" pageId="7" pageNumber="58">S. latirostris</emphasis>
</taxonomicName>
and 
<taxonomicName id="A4BA4D17C05C85504D992C45FEBCB4C3" ID-CoL="85Y22" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[151,276,873,899]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="51CEEA86C05C85504D992C45FEBCB4C3" box="[151,276,873,899]" italics="true" pageId="7" pageNumber="58">S. cristatus</emphasis>
</taxonomicName>
have red lines or bands on the carapace, which are absent in 
<taxonomicName id="A4BA4D17C05C85504EC32C45FBE2B4C3" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[973,1098,873,899]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05C85504EC32C45FBE2B4C3" box="[973,1098,873,899]" italics="true" pageId="7" pageNumber="58">S. venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05C8550495D2C45FB05B4C3" box="[1107,1197,873,899]" pageId="7" pageNumber="58" rank="species">
<emphasis id="51CEEA86C05C8550495D2C45FB05B4C3" bold="true" box="[1107,1197,873,899]" pageId="7" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
In addition, in 
<taxonomicName id="A4BA4D17C05C855048532C45FF77B4E7" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05C855048532C45FF77B4E7" italics="true" pageId="7" pageNumber="58">S. venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05C85504DE82CA1FE97B4E7" box="[230,319,909,935]" pageId="7" pageNumber="58" rank="species">
<emphasis id="51CEEA86C05C85504DE82CA1FE97B4E7" bold="true" box="[230,319,909,935]" pageId="7" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
the red bands on the pleon appear to be narrower and somewhat diffuse (
<figureCitation id="FB812A11C05C8550497F2CA1FB1DB4E7" box="[1137,1205,909,935]" captionStart="FIGURE 4" captionStartId="6.[151,250,1894,1919]" captionTargetBox="[166,1421,166,1875]" captionTargetId="figure@6.[166,1421,166,1875]" captionTargetPageId="6" captionText="FIGURE 4. Salmoneus venustus sp. nov.: non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH. 2015-08-0031 [a, b]; holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [c]; a—living individual, dorsal view; b—same, lateral view (both photographs by the author); c—fresh specimen post-mortem, lateral view (photograph courtesy of Dr. Ivan N. Marin)." httpUri="https://zenodo.org/record/3359034/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
), rather than sharply marked, as in 
<taxonomicName id="A4BA4D17C05C85504C3C2C9DFE12B48B" ID-CoL="85Y2G" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[306,442,945,971]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="51CEEA86C05C85504C3C2C9DFE12B48B" box="[306,442,945,971]" italics="true" pageId="7" pageNumber="58">S. latirostris</emphasis>
</taxonomicName>
and 
<taxonomicName id="A4BA4D17C05C85504CFE2C9DFDC3B48B" ID-CoL="85Y22" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[496,619,945,971]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="51CEEA86C05C85504CFE2C9DFDC3B48B" box="[496,619,945,971]" italics="true" pageId="7" pageNumber="58">S. cristatus</emphasis>
</taxonomicName>
(A. Anker, pers. obs.). These differences in the colouration add to several morphological differences between these three species, especially between the new species and 
<taxonomicName id="A4BA4D17C05C855049C32CF9FAE1B4AF" ID-CoL="85Y22" baseAuthorityName="Coutiere" baseAuthorityYear="1897" box="[1229,1353,981,1007]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="51CEEA86C05C855049C32CF9FAE1B4AF" box="[1229,1353,981,1007]" italics="true" pageId="7" pageNumber="58">S. cristatus</emphasis>
</taxonomicName>
, which possesses a well-developed rostral and mid-dorsal carina not seen in 
<taxonomicName id="A4BA4D17C05C85504E982CD5FBBAB353" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[918,1042,1017,1043]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05C85504E982CD5FBBAB353" box="[918,1042,1017,1043]" italics="true" pageId="7" pageNumber="58">S. venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05C855049172CD5FBDAB353" box="[1049,1138,1017,1043]" pageId="7" pageNumber="58" rank="species">
<emphasis id="51CEEA86C05C855049172CD5FBDAB353" bold="true" box="[1049,1138,1017,1043]" pageId="7" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<bibRefCitation id="072B4B65C05C8550498E2CD5FA8CB353" author="Coutiere, H." box="[1152,1316,1017,1043]" pageId="7" pageNumber="58" pagination="1 - 559" refId="ref6680" refString="Coutiere, H. (1899) Les Alpheidae. Morphologie externe et interne, formes larvaires, bionomie. Annales des Sciences Naturelles, Zoologie et Paleontologie, 8 (9), 1 - 559, 6 pls. 6. https: // doi. org / 10.5962 / bhl. title. 13143" type="book chapter" year="1899">Coutière 1899</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C855048212CD5FE82B377" author="Banner, D. M. &amp; Banner, A. H." pageId="7" pageNumber="58" pagination="1 - 99" refId="ref6502" refString="Banner, D. M. &amp; Banner, A. H. (1981). Annotated checklist of the alpheid shrimp of the Red Sea and Gulf of Aden. Zoologische Verhandelingen, Leiden, 190, 1 - 99." type="journal article" year="1981">Banner &amp; Banner 1981</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C85504C382B30FD8AB378" author="Duris, Z. &amp; Horka, I." box="[310,546,1052,1080]" pageId="7" pageNumber="58" pagination="773 - 793" refId="ref6964" refString="Duris, Z. &amp; Horka, I. (2016) Salmoneus chadwickae, a new alpheid shrimp (Crustacea: Decapoda: Alpheidae) from the Red Sea, with remarks on related or regional congeners. Marine Biodiversity, 46, 773 - 793. https: // doi. org / 10.1007 / s 12526 - 015 - 0427 - 4" type="journal article" year="2016">Ďuriš &amp; Horká 2016</bibRefCitation>
).
</paragraph>
<paragraph id="63053694C05C85504DC92B6DFF48B3AC" blockId="7.[151,1437,153,1439]" pageId="7" pageNumber="58">
The two species presently known from incomplete specimens, 
<taxonomicName id="A4BA4D17C05C85504E9D2B6DFB7DB31B" ID-CoL="85Y29" authority="(De Man, 1910)" authorityName="De Man" authorityYear="1910" baseAuthorityName="De Man" baseAuthorityYear="1910" box="[915,1237,1089,1115]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="hilarulus">
<emphasis id="51CEEA86C05C85504E9D2B6DFBBEB31B" box="[915,1046,1089,1115]" italics="true" pageId="7" pageNumber="58">S. hilarulus</emphasis>
(
<bibRefCitation id="072B4B65C05C855049282B6DFB65B31B" author="De Man, J. G." box="[1062,1229,1089,1115]" pageId="7" pageNumber="58" pagination="287 - 319" refId="ref6857" refString="De Man, J. G. (1910) Diagnoses of new species of macrurous decapod Crustacea from the &quot; Siboga-Expedition &quot;. Tijdschrift der Nederlandsche Dierkundige Vereeniging, 2 (11), 287 - 319." type="journal article" year="1910">De Man, 1910</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="A4BA4D17C05C855048012B6DFE09B33F" ID-CoL="85Y2Z" authority="Banner &amp; Banner, 1966" authorityName="Banner &amp; Banner" authorityYear="1966" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="tafaongae">
<emphasis id="51CEEA86C05C855048012B6DFA34B31B" box="[1295,1436,1089,1115]" italics="true" pageId="7" pageNumber="58">S. tafaongae</emphasis>
<bibRefCitation id="072B4B65C05C85504D992B49FE09B33F" author="Banner, A. H. &amp; Banner, D. M." box="[151,417,1125,1151]" pageId="7" pageNumber="58" pagination="145 - 188" refId="ref6453" refString="Banner, A. H. &amp; Banner, D. M. (1966) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part X. Collections from Fiji, Tonga and Samoa. Pacific Science, 20, 145 - 188." type="journal article" year="1966">Banner &amp; Banner, 1966</bibRefCitation>
</taxonomicName>
(
<typeStatus id="BC018836C05C85504CA12B49FDBBB33F" box="[431,531,1125,1151]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
lost in the fire of the Hawaii Marine Laboratory) differ from 
<taxonomicName id="A4BA4D17C05C855049CE2B49FA94B33F" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[1216,1340,1125,1151]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05C855049CE2B49FA94B33F" box="[1216,1340,1125,1151]" italics="true" pageId="7" pageNumber="58">S. venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05C8550484D2B49FA34B33F" box="[1347,1436,1125,1151]" pageId="7" pageNumber="58" rank="species">
<emphasis id="51CEEA86C05C8550484D2B49FA34B33F" bold="true" box="[1347,1436,1125,1151]" pageId="7" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
in several important features, such as the much stouter dactylus of the third pereiopod in the former species (cf. 
<bibRefCitation id="072B4B65C05C855048752BA5FEA4B387" author="De Man, J. G." pageId="7" pageNumber="58" pagination="133 - 465" refId="ref6896" refString="De Man, J. G. (1911) The Decapoda of the Siboga Expedition. Part II. Family Alpheidae. Siboga Expeditie, 39 a 1, 133 - 465; (1915) Supplement-Explanations of plates of Alpheidae. E. J. Brill, Leiden, 23 pls. https: // doi. org / 10.5962 / bhl. title. 10512" type="book chapter" year="1911">De Man 1911</bibRefCitation>
: fig. 
<quantity id="A4429B71C05C85504C4C2B81FEC6B387" box="[322,366,1197,1223]" metricMagnitude="-2" metricUnit="kg" metricValue="1.0" pageId="7" pageNumber="58" unit="g" value="10.0">10g</quantity>
), and the much longer rostrum and up-turned orbital teeth in the latter species (
<bibRefCitation id="072B4B65C05C855049D92B81FF7BB3AC" author="Banner, A. H. &amp; Banner, D. M." pageId="7" pageNumber="58" pagination="145 - 188" refId="ref6453" refString="Banner, A. H. &amp; Banner, D. M. (1966) Contributions to the knowledge of the alpheid shrimp of the Pacific Ocean. Part X. Collections from Fiji, Tonga and Samoa. Pacific Science, 20, 145 - 188." type="journal article" year="1966">Banner &amp; Banner 1966</bibRefCitation>
).
</paragraph>
<paragraph id="63053694C05C85504DC92BD9FAE5B2DF" blockId="7.[151,1437,153,1439]" pageId="7" pageNumber="58">
<taxonomicName id="A4BA4D17C05C85504DC92BD9FE01B24F" ID-CoL="85Y34" authority="Anker, 2019" authorityName="Anker" authorityYear="2019" box="[199,425,1269,1295]" class="Malacostraca" family="Alpheidae" genus="Salmoneus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="venustus" status="sp. nov.">
<emphasis id="51CEEA86C05C85504DC92BD9FE01B24F" box="[199,425,1269,1295]" italics="true" pageId="7" pageNumber="58">Salmoneus venustus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4AFD57FDC05C85504CBC2BD9FDA5B24F" box="[434,525,1269,1295]" pageId="7" pageNumber="58" rank="species">
<emphasis id="51CEEA86C05C85504CBC2BD9FDA5B24F" bold="true" box="[434,525,1269,1295]" pageId="7" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
appears to be an obligate burrow symbiont, as indicated by the significantly reduced, poorly pigmented corneas (
<figureCitation id="FB812A11C05C85504F162A35FDF3B273" box="[536,603,1305,1331]" captionStart="FIGURE 4" captionStartId="6.[151,250,1894,1919]" captionTargetBox="[166,1421,166,1875]" captionTargetId="figure@6.[166,1421,166,1875]" captionTargetPageId="6" captionText="FIGURE 4. Salmoneus venustus sp. nov.: non-ovigerous specimen (cl 5.5 mm) from northern Oman, OUMNH. 2015-08-0031 [a, b]; holotype, non-ovigerous specimen (cl 6.4 mm) from Nha Trang Bay, Vietnam, MNHN-IU-2014-20290 [c]; a—living individual, dorsal view; b—same, lateral view (both photographs by the author); c—fresh specimen post-mortem, lateral view (photograph courtesy of Dr. Ivan N. Marin)." httpUri="https://zenodo.org/record/3359034/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
) and the microhabitat / collection data. Brief field notes accompanying the Vietnamese specimen (
<typeStatus id="BC018836C05C85504C962A11FE55B218" box="[408,509,1341,1368]" pageId="7" pageNumber="58" type="holotype">holotype</typeStatus>
) suggest that its host may have been a large ghost shrimp from the genus 
<taxonomicName id="A4BA4D17C05C855048222A11FA34B217" authorityName="Stimpson" authorityYear="1866" box="[1324,1436,1341,1367]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="51CEEA86C05C855048222A11FA34B217" box="[1324,1436,1341,1367]" italics="true" pageId="7" pageNumber="58">Glypturus</emphasis>
</taxonomicName>
sp. (
<taxonomicName id="A4BA4D17C05C85504DC92A4DFED8B23C" ID-CoL="5b184d39-f8ea-420d-acad-9cb703e2a9c3" authorityName="Dana" authorityYear="1852" box="[199,368,1377,1404]" class="Malacostraca" family="Callianassidae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="family">Callianassidae</taxonomicName>
). Spacious galleries of 
<taxonomicName id="A4BA4D17C05C85504F772A4DFD41B23B" authorityName="Stimpson" authorityYear="1866" box="[633,745,1377,1403]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="51CEEA86C05C85504F772A4DFD41B23B" box="[633,745,1377,1403]" italics="true" pageId="7" pageNumber="58">Glypturus</emphasis>
</taxonomicName>
spp. are known to harbour a number of other alpheid “commensals” (e.g. 
<bibRefCitation id="072B4B65C05C85504C322AA9FDACB2DF" author="Anker, A. &amp; Marin, I. N." box="[316,516,1413,1439]" pageId="7" pageNumber="58" pagination="295 - 319" refId="ref6175" refString="Anker, A. &amp; Marin, I. N. (2006) New records and species of Alpheidae (Crustacea, Decapoda) from Vietnam. Part I. Genus Salmoneus Holthuis, 1955. The Raffles Bulletin of Zoology, 54, 295 - 319." type="journal article" year="2006">
Anker 
<emphasis id="51CEEA86C05C85504C872AAAFE69B2DF" box="[393,449,1413,1439]" italics="true" pageId="7" pageNumber="58">et al.</emphasis>
2006
</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C85504F012AA9FCB8B2DF" author="Dworschak, P. C. &amp; Anker A." box="[527,784,1413,1439]" pageId="7" pageNumber="58" pagination="1 - 16" refId="ref7078" refString="Dworschak, P. C., Marin. I. N. &amp; Anker A. (2006) A new species of Naushonia Kingsley, 1897 (Decapoda: Thalassinidea: Laomediidae) from Vietnam and the Philippines with notes on the genus Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997. Zootaxa, 1372, 1 - 16." type="journal article" year="2006">
Dworschak 
<emphasis id="51CEEA86C05C85504F9B2AAAFD66B2DF" box="[661,718,1413,1439]" italics="true" pageId="7" pageNumber="58">et al.</emphasis>
2006
</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C85504E142AA9FBEDB2DF" author="Anker, A. &amp; Dworschak, P. C." box="[794,1093,1413,1439]" pageId="7" pageNumber="58" pagination="290 - 302" refId="ref5997" refString="Anker, A. &amp; Dworschak, P. C. (2007) Jengalpheops rufus, gen. nov., sp. nov., a new commensal alpheid shrimp from the Philippines (Crustacea: Decapoda). Zoological Studies, 46, 290 - 302." type="journal article" year="2007">Anker &amp; Dworschak 2007</bibRefCitation>
; 
<bibRefCitation id="072B4B65C05C855049412AA9FAE9B2DF" author="Anker, A. &amp; Marin, I. N." box="[1103,1345,1413,1439]" pageId="7" pageNumber="58" pagination="91 - 107" refId="ref6224" refString="Anker, A. &amp; Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker &amp; Marin 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>