142 lines
17 KiB
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142 lines
17 KiB
XML
<document id="B285162D99B64AF5E5662925995F8F2B" ID-CLB-Dataset="33935" ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="E7908052B7F06F7C572CA8683A03BA49" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha glandulosa Attems 1937" docType="treatment" docVersion="8" lastPageNumber="62" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="60" updateTime="1732923565725" updateUser="ExternalLinkService">
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<mods:title id="B7D633483AEB37DF2449C8510E686AA5">Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="F58A2584A93DBF17D2BF4EB960D8ED95">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="E09BED6B5BD61A6393729E16FF080702">Golovatch, Sergei I.</mods:namePart>
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<mods:namePart id="6BCCDA8FF5799F69EBAD2BBCEB79704E">Panha, Somsak</mods:namePart>
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<mods:typeOfResource id="96F153C6CDF1A9039D1942F80647B3B6">text</mods:typeOfResource>
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<mods:title id="91D70990AC60F311F8F2DECE34F108FB">ZooKeys</mods:title>
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<mods:date id="D06BCD3804525BD198E513D0E35B1F08">2011</mods:date>
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<mods:detail id="AF03A5A77C0911193312F2601F3A6D1A" type="volume">
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<mods:number id="D20166E6C2D7BF0FEC203D1A1EB11364">131</mods:number>
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<mods:url id="15169F2500A4F9D935703F7B1B747B5E">http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
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<mods:classification id="D0C6EA83EDB8FFCF222A5F74FC7DF1D4">journal article</mods:classification>
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<mods:identifier id="F1E63BC3396A41F7A19329680E3B01DE" type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
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<mods:identifier id="FA87BD12FA89976916996857DAE2281A" type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
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<treatment id="E7908052B7F06F7C572CA8683A03BA49" ID-GBIF-Taxon="152031103" LSID="urn:lsid:plazi:treatment:E7908052B7F06F7C572CA8683A03BA49" httpUri="http://treatment.plazi.org/id/E7908052B7F06F7C572CA8683A03BA49" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60" scope_class="Diplopoda" scope_family="Paradoxosomatidae" scope_order="Polydesmida">
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<subSubSection id="BEC33C42D282D6E2275B1EF4A2060859" pageId="59" pageNumber="60" type="nomenclature">
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<paragraph id="16F1F4EB452C27C002418DBFFDEBCADD" pageId="59" pageNumber="60">
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<taxonomicName id="16A658FA801ADFCD11B263CFCF395128" ID-CoL="6T8N9" LSID="http://species-id.net/wiki/Orthomorpha_glandulosa" authority="Attems, 1937" authorityName="Attems" authorityYear="1937" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha glandulosa" order="Polydesmida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="glandulosa">Orthomorpha glandulosa (Attems, 1937)</taxonomicName>
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Figs 5152
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</paragraph>
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</subSubSection>
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<subSubSection id="D564385D3FCA43D451A1A6B459A325D4" pageId="59" pageNumber="60" type="reference_group">
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<paragraph id="B58F31A2A6ACFDFD6737013AE6FA330E" pageId="59" pageNumber="60">
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<taxonomicName id="4260C8439B6BCDEA724A7C4563067F85" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus glandulosus" order="Polydesmida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="glandulosus">Pratinus glandulosus</taxonomicName>
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<bibRefCitation id="6A24C03F9AA27050CF23A81B1657A4CF" pageId="59" pageNumber="60">Attems 1937</bibRefCitation>
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: 119 (D).
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</paragraph>
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<paragraph id="DA5C7FF47381DC2393DBAFD0D16ACEAC" pageId="59" pageNumber="60">
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<taxonomicName id="4B93A963B5E19C20E0110AF3823B099F" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus glandulosus" order="Polydesmida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="glandulosus">Pratinus glandulosus</taxonomicName>
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-
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<bibRefCitation id="B62FC784249FE9B5C00F31CCA094A957" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">Attems 1938</bibRefCitation>
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: 220 (D).
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</paragraph>
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<paragraph id="95313CB0A121863468662132860F63F8" pageId="59" pageNumber="60">
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<taxonomicName id="9BEADF21450D8F5ECEFDBAF106C7FF25" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha glandulosa" order="Polydesmida" pageId="59" pageNumber="60" phylum="Arthropoda" rank="species" species="glandulosa">Orthomorpha glandulosa</taxonomicName>
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-
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<bibRefCitation id="83A6F0819F38CC44E6964FC6DFC0ABF5" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
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: 265 (M);
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<bibRefCitation id="D6465D9248CA2F7801C3DFF2D0096EA4" author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="355 - 364" title="A new species of Orthomorpha Bollman from Thailand observed in migration, with taxonomic notes on the genus (Diplopoda)." volume="107" year="1964">1964</bibRefCitation>
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: 361 (M, D);
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<bibRefCitation id="DAD372E547968B952253DF5771842F2C" author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">1968</bibRefCitation>
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: 56 (M);
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<bibRefCitation id="090673ABF29FB7193A318A3926734FA7" author="Hoffman, RL" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="699 - 719" title="Diplopoda from Malayan caves, collected by M. Pierre Strinati." volume="84" year="1977">Hoffman 1977</bibRefCitation>
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: 700 (M);
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<bibRefCitation id="A8BCBF954DDB9C93A2B61780B0A8E588" pageId="59" pageNumber="60">Golovatch 1998</bibRefCitation>
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: 42 (D, M);
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<bibRefCitation id="826974CB1223F4284F7255A14778F2A7" author="Enghoff, H" journalOrPublisher="Arthropoda Selecta" pageId="114" pageNumber="115" pagination="29 - 43" title="A review of the millipede fauna of Vietnam (Diplopoda)." volume="13" year="2004">Enghoff et al. 2004</bibRefCitation>
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: 38 (M, R).
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</paragraph>
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</subSubSection>
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<subSubSection id="C574BAF5D65AA36781667F0A41583B22" pageId="59" pageNumber="60" type="lectotype">
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<paragraph id="899024560EC70079FFB5532FA6309514" pageId="59" pageNumber="60">Lectotype.</paragraph>
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<paragraph id="9004D205D7A475981AC6C1243F69CF1B" pageId="59" pageNumber="60">♂ (NHMW-3506), Vietnam, Hon Ba Island, Nhatrang, 06.1930, leg. C. Dawydoff.</paragraph>
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</subSubSection>
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<subSubSection id="29E9482CBFE041E8FEDBB0CCFAA18AB6" pageId="59" pageNumber="60" type="paralectotypes">
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<paragraph id="68591F650915549D1B1CB7A0B424A777" pageId="59" pageNumber="60">Paralectotypes.</paragraph>
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<paragraph id="FB97002611B27F73A21BC600BE081C05" pageId="59" pageNumber="60">1 ♀ (NHMW-3506), same locality, together with lectotype. 1 ♀ (NHMW-3505), Vietnam, Darlac, frontier du Cambodge, 07.1930, leg. C. Dawydoff.</paragraph>
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<paragraph id="66289546DE5B7D7F68BE1A29B90E59F7" pageId="59" pageNumber="60">Lectotype designation proposed herewith is necessary to ensure the species is based on a complete male.</paragraph>
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</subSubSection>
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<subSubSection id="68DF628EF7788B51A8B105AB94279981" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60" type="redescription">
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<paragraph id="CCE0AC6910008D6C6B385282C7B7A9F3" pageId="59" pageNumber="60">Redescription.</paragraph>
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<paragraph id="F092DD9A2A6F020FA18654BE218B240F" pageId="59" pageNumber="60">
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Length ca 38 mm (lectotype), 31-34 mm (♀), width of midbody pro- and metazona 3.4 and 5.0 mm (lectotype), 2.9-3.4 and 4.2-4.4 mm (♀), respectively (vs 3.0 and 5.0 in width, as given in the available descriptions (
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<bibRefCitation id="BC8D181730F73B552F7AF5BDB67D1B32" pageId="59" pageNumber="60">Attems 1937</bibRefCitation>
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,
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<bibRefCitation id="E573DDE735E09EA96DDA45F9CED7AA50" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
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)). Coloration of alcohol material upon long-term preservation dark grey-brown (Fig. 51) with contrasting pallid paraterga and epiproct, and light brown venter and legs (vs dark castaneous brown with paraterga and epiproct yellow, and venter and legs light red-brown, as given in the descriptions (
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<bibRefCitation id="9FC5FF1FD122E689E8FE6272E25BF9C6" pageId="59" pageNumber="60">Attems 1937</bibRefCitation>
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,
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<bibRefCitation id="2AA539F0502BA3A560EEBAA0F90F9AFE" author="Attems, C" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Nouvelle Serie," pageId="114" pageNumber="115" pagination="187 - 321" title="Die von Dr. C. Dawydoff in Franzoesisch Indochina gesammelten Myriopoden." volume="6" year="1938">1938</bibRefCitation>
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)).
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</paragraph>
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<paragraph id="D69339C8ACDF7A4A4FF7040105B056E7" lastPageId="61" lastPageNumber="62" pageId="59" pageNumber="60">
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Head usual, clypeolabral region densely setose, surface of vertex smooth, with a few setae flanking a distinct epicranial suture. Antennae long and slender (Fig. 51A, B & J), extending behind segment 4 (♂) or surpassing segment 3 (♀) dorsally. Head in width <collum <segment 2 <3 = 4 <5-16 (♂), or head <collum <segments 3 and 4 <2 <5-16 (♀), gently and gradually tapering thereafter. Collum smooth, with three
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<pageBreakToken id="6D15E28A80CDD92A7CAF68390EE2CC44" pageId="60" pageNumber="61" start="start">transverse</pageBreakToken>
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rows of setae traceable only as insertion points, 4+4 anterior, 2+2 intermediate, and 3+3 posterior setae; caudal corner of paraterga acutangular (ca 75°), nearly pointed (Fig. 51A, B & J). Tegument poorly shining; metaterga coriaceous, rugulose, each postcollum one with two rows of fully abraded setae borne on minute tubercles growing increasingly strongly reduced towards epiproct: 2+2 in front row and 3+3 in caudal one; prozona very finely shagreened, surface below paraterga finely microgranulate. Axial line rather evident, starting from collum. Paraterga very strongly developed (Fig. 51A-G & J-L), set high (at ca 1/4 metazonital height), in ♂ evidently upturned, lying above dorsum on postcollum segments, in ♀ mostly below dorsum, rather thin in lateral view, a little thicker on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, better so in ♂, nearly pointed to pointed, caudal tip on paraterga 16-19 evidently curved mesad. Calluses on paraterga 2 delimited by a sulcus only dorsally, on following paraterga both dorsally and ventrally, rather broad. Paraterga 2 broad, anterior edge angulate, lateral edge with two minute incisions in anterior 1/3; posterior edge evidently concave (Fig. 51A, B & J). Paraterga 3 and 4 subequal, like subsequent paraterga, anterior edge broadly rounded, bordered and fused to callus, lateral edge with one minute incision in front 1/3. Ozopores evident,
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<pageBreakToken id="1E5E14D46991266C5C75BAC53C0D7ACB" pageId="61" pageNumber="62" start="start">lateral</pageBreakToken>
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, lying inside an ovoid groove, placed at about 1/3 metazonital length. Transverse sulcus complete on metaterga 4-18, incomplete on metaterga 2 and 3 (♂), or incomplete on metatergum 4 and complete on metaterga 5-18 (♀), shallow, not reaching bases of paraterga, ribbed at bottom, slightly sinuate anteromedially (Fig. 51A, C, F & J-L). Stricture between pro- and metazona narrow, shallow, beaded at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segments 2-4 (♂, ♀) (Fig. 51B, D & E), each with an evident sharp denticle caudally, thereafter increasingly strongly reduced until segment 10 (♂, ♀). Epiproct (Fig. 51E-G & L) conical, flattened dorsoventrally, apical papillae small, dentiform, directed caudoventrally; tip subtruncate; pre-apical papillae small, lying close to tip. Hypoproct (Fig. 51G) roundly subtrapeziform, setiferous knobs at caudal margin small and well-separated.
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</paragraph>
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<paragraph id="59E058609371A01E4B5EC3431AF3181A" pageId="61" pageNumber="62">Sterna sparsely setose, without modifications, but with a large, central, setose cone between ♂ coxae 4 (Fig. 51H & I). A paramedian pair of tubercles in front of gonopod aperture absent. Legs long and slender, midbody ones ca 1.3-1.4 (♂) or 1.2-1.3 (♀) as long as body height, prefemora without modifications, tarsal brushes present until ♂ legs 5.</paragraph>
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<paragraph id="36D619745F2A6CEDDE97E94FB2FA4B7F" pageId="61" pageNumber="62">
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Gonopods (Fig. 52) simple. Coxa long and slender, with several setae distodorsally. Prefemur rather large, densely setose, more than 2 times shorter than femorite +
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<normalizedToken id="619A38BD876F8C8B7D402348D8D75D76" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part. Femorite very slender, evidently curved, not enlarged distad,
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<normalizedToken id="B5D89E82931C8AB2AA88F2AC91B313FB" originalValue="“postfemoral”">"postfemoral"</normalizedToken>
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part demarcated by an oblique lateral sulcus; tip of solenophore small, trifid, with two subequal denticles (terminal and middle) and a larger subterminal lobule.
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</paragraph>
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<caption id="51C2666857F39A3DD63DCEF13B242D9A" pageId="61" pageNumber="62">
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<paragraph id="43DC5A43EEB4048E73AAA6A387C428C5" pageId="61" pageNumber="62">
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Figure 51.
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<taxonomicName id="197CA163AF6E6736393451E784DE1760" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha glandulosa" order="Polydesmida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="glandulosa">Orthomorpha glandulosa</taxonomicName>
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(Attems, 1937), ♂ lectotype (
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<normalizedToken id="0FB592EC299C21106E54B23EF37D909D" originalValue="A–I">A-I</normalizedToken>
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), ♀ paralectotype (
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<normalizedToken id="62358550FC32AEECF225DF507B7667FD" originalValue="J–L">J-L</normalizedToken>
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). A, B, J anterior part of body, dorsal, lateral and dorsal views, respectively C, D, K segments 10 and 11, dorsal, lateral and dorsal views, respectively
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<normalizedToken id="5023DCDDD75461D22B01BD9C0A8A6D12" originalValue="E–G">E-G</normalizedToken>
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, L posterior part of body, lateral, dorsal, ventral and dorsal views, respectively H, I sternal cones between coxae 4, subcaudal and sublateral views, respectively.
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</paragraph>
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</caption>
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<caption id="6C6DC543E5DC106F585F95715244BCE9" pageId="61" pageNumber="62">
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<paragraph id="07412C50E644F6BF5383EFEED65EF6E3" pageId="61" pageNumber="62">
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Figure 52.
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<taxonomicName id="368CFAD36F7E2CA997EBE04EDCAD429F" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha glandulosa" order="Polydesmida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="glandulosa">Orthomorpha glandulosa</taxonomicName>
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(Attems, 1937), ♂ lectotype. A, B left gonopod, mesal and lateral views, respectively.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |