218 lines
27 KiB
XML
218 lines
27 KiB
XML
<document id="6099569A81D14044680B22738C73858D" ID-DOI="10.5281/zenodo.186920" ID-GBIF-Dataset="695e995e-3a9a-41b8-bafa-ed8366b37ef4" ID-ISSN="1175-5326" ID-Zenodo-Dep="186920" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461059560947" checkinUser="plazi" docAuthor="Breure, Abraham S. H." docDate="2009" docId="CF1A2655FFC9BD7AE9A6F94F1D81FE5C" docLanguage="en" docName="zt02065p050.pdf" docOrigin="Zootaxa 2065" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Drymaeus (Drymaeus) rex Breure, 2009, spec.nov." docType="treatment" docVersion="8" lastPageNumber="43" masterDocId="33235E2DFFD9BD68E931FFE61949FF9C" masterDocTitle="New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World" masterLastPageNumber="50" masterPageNumber="25" pageNumber="41" updateTime="1698581906488" updateUser="plazi">
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<mods:titleInfo id="ED57255BDDD735BF86EB9A8E80DD6FDE">
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<mods:title id="B50433516C58DC34393703EE5B8D6FE2">New Orthalicidae (Mollusca, Gastropoda) from Venezuelan Guayana: unravelling secrets from the Lost World</mods:title>
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</mods:titleInfo>
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<mods:name id="1155F2016A371A9CDE6DA171BB17C357" type="personal">
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<mods:roleTerm id="E63F2AC428A50954E0AF2A4714CBC1C2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CDCAB949FA891A3DDDA31D490F37A705">Breure, Abraham S. H.</mods:namePart>
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<mods:typeOfResource id="3A039B3100A58F4E6B95329F00D93512">text</mods:typeOfResource>
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<mods:title id="87EA5B7817A244A957D3651E26C4CBF6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="4D9AD3ADAA40AD699670B5D163DC0B71">
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<mods:date id="28D6C822F2D433253BE7DA1F421CA2A8">2009</mods:date>
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<mods:detail id="5216B7468554557A295AFC70D734D52D" type="volume">
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<mods:number id="46812839B848CF2CC0EEB44693709BAD">2065</mods:number>
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<mods:extent id="31FEBF09A3DF4273EF5B4807C7F3B03E" unit="page">
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<mods:start id="ECCCE69CF1E14F8927B1422094C46941">25</mods:start>
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<mods:classification id="DDCEEA9C189A2734B5BED4031C9CCDCF">journal article</mods:classification>
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<mods:identifier id="C3E98990410B46DAA8B2322AE33647ED" type="DOI">10.5281/zenodo.186920</mods:identifier>
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<mods:identifier id="3D2857B86E1E361FB927DCD86AF8A027" type="GBIF-Dataset">695e995e-3a9a-41b8-bafa-ed8366b37ef4</mods:identifier>
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<mods:identifier id="CA52325B63530C0056F6EA1B88FE31A7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="08DDFFC7A2E90D0F3CF80779B82F8EFE" type="Zenodo-Dep">186920</mods:identifier>
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<treatment id="CF1A2655FFC9BD7AE9A6F94F1D81FE5C" ID-DOI="http://doi.org/10.5281/zenodo.5618382" ID-GBIF-Taxon="119613776" ID-Zenodo-Dep="5618382" LSID="urn:lsid:plazi:treatment:CF1A2655FFC9BD7AE9A6F94F1D81FE5C" httpUri="http://treatment.plazi.org/id/CF1A2655FFC9BD7AE9A6F94F1D81FE5C" lastPageId="18" lastPageNumber="43" pageId="16" pageNumber="41">
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<subSubSection id="0FA9C4C8FFC9BD78E9A6F94F1877F97A" pageId="16" pageNumber="41" type="nomenclature">
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<paragraph id="470C9743FFC9BD78E9A6F94F1B06F95F" blockId="16.[151,591,1705,1766]" box="[151,591,1705,1731]" pageId="16" pageNumber="41">
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<heading id="1C44202FFFC9BD78E9A6F94F1B06F95F" bold="true" box="[151,591,1705,1731]" fontSize="11" level="1" pageId="16" pageNumber="41" reason="1">
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<emphasis id="75C74B51FFC9BD78E9A6F94F1B06F95F" bold="true" box="[151,591,1705,1731]" pageId="16" pageNumber="41">
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<taxonomicName id="80B3ECC0FFC9BD78E9A6F94F189EF95F" box="[151,471,1705,1731]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="41" phylum="Mollusca" rank="species" species="rex" status="sp. nov." subGenus="Drymaeus">
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<emphasis id="75C74B51FFC9BD78E9A6F94F189EF95F" bold="true" box="[151,471,1705,1731]" italics="true" pageId="16" pageNumber="41">Drymaeus (Drymaeus) rex</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="6EF4F62AFFC9BD78E8EEF94F1B06F95F" box="[479,591,1705,1731]" pageId="16" pageNumber="41" rank="species">spec.nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="470C9743FFC9BD78E9A6F92A1877F97A" blockId="16.[151,591,1705,1766]" box="[151,318,1740,1766]" pageId="16" pageNumber="41">
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(
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<figureCitation id="DF888BC6FFC9BD78E9AEF92A19A2F97A" box="[159,235,1740,1766]" captionStart="FIGURE 7" captionStartId="10.[151,262,950,974]" captionTargetBox="[165,1434,203,909]" captionTargetId="figure@10.[151,1436,194,926]" captionTargetPageId="10" captionText="FIGURE 7. Drymaeus (Drymaeus) rex spec. nov. A. Genitalia UF 284722; B. Longitudinal section penis; C. Spermatophore; D. Pallial organs. Scale bar— 5 mm." httpUri="https://zenodo.org/record/186927/files/figure.png" pageId="16" pageNumber="41">Figs 7</figureCitation>
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,
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<date id="330DB183FFC9BD78E9C9F92A1858F97A" box="[248,273,1740,1766]" pageId="16" pageNumber="41">
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<figureCitation id="DF888BC6FFC9BD78E9C9F92A184FF97A" box="[248,262,1740,1766]" captionStart="FIGURE 8" captionStartId="16.[151,255,1462,1486]" captionTargetBox="[151,1434,274,1435]" captionTargetId="figure@16.[151,1435,274,1437]" captionTargetPageId="16" captionText="FIGURE 8. Shells in Drymaeus (Drymaeus). A ─ C. D. (D.) extraneus (Haas, 1955). A ─ B. Holotype FMNH 49736; C. FMNH 49734. D ─ F. D. (D.) steyermarki (Haas, 1955), holotype FMNH 49735. G ─ H. D. (D.) yapacanensis Breure & Eskens, 1981, holotype RMNH 55331. I ─ S. Drymaeus (Drymaeus) rex spec. nov. I ─ M, Q ─ S. Paratypes UF 284745, 284730, 284759, 284748, 284714, 284719, 284719 and 284715 respectively; N ─ P. Holotype UF 284726. Scale bar— 1 cm." httpUri="https://zenodo.org/record/186928/files/figure.png" pageId="16" pageNumber="41">8</figureCitation>
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I
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</date>
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─S)
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</paragraph>
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</subSubSection>
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<subSubSection id="0FA9C4C8FFC9BD78E9A6F8FF1AEAF81D" pageId="16" pageNumber="41" type="diagnosis">
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<paragraph id="470C9743FFC9BD78E9A6F8FF1AEAF81D" blockId="16.[151,1437,1817,2036]" pageId="16" pageNumber="41">
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<emphasis id="75C74B51FFC9BD78E9A6F8FF185EF8AF" bold="true" box="[151,279,1817,1843]" pageId="16" pageNumber="41">Diagnosis.</emphasis>
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A relatively large species of
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<taxonomicName id="80B3ECC0FFC9BD78EB5DF8FF1BADF8AE" box="[620,740,1817,1842]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="41" phylum="Mollusca" rank="genus">
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<emphasis id="75C74B51FFC9BD78EB5DF8FF1BADF8AE" box="[620,740,1817,1842]" italics="true" pageId="16" pageNumber="41">Drymaeus</emphasis>
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</taxonomicName>
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, with a white line below the suture and typically with three interrupted spiral colour bands on the last whorl, crossing weaker axial streaks of a less intense colour. Aperture with a white, slightly expanded lip, bordered pink inside.
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</paragraph>
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</subSubSection>
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<subSubSection id="0FA9C4C8FFC9BD79E9F4F86B180AFBCE" lastPageId="17" lastPageNumber="42" pageId="16" pageNumber="41" type="description">
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<paragraph id="470C9743FFC9BD79E9F4F86B1D66FD92" blockId="16.[151,1437,1817,2036]" lastBlockId="17.[151,1437,152,2034]" lastPageId="17" lastPageNumber="42" pageId="16" pageNumber="41">
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<emphasis id="75C74B51FFC9BD78E9F4F86B182CF83B" bold="true" box="[197,357,1933,1959]" pageId="16" pageNumber="41">Description.</emphasis>
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Shell up to
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<quantity id="804B3AA6FFC9BD78E8CFF86B1B25F83B" box="[510,620,1933,1959]" metricMagnitude="-2" metricUnit="m" metricValue="4.07" pageId="16" pageNumber="41" unit="mm" value="40.7">40.7 mm</quantity>
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(further dimension given in
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<tableCitation id="0A31A2F8FFC9BD78EAE3F86B1D64F83B" box="[978,1069,1933,1959]" captionStart="TABLE 1" captionStartId="8.[151,240,267,291]" captionTargetBox="[151,1436,422,659]" captionTargetPageId="8" captionText="TABLE 1. Dimensions in selected species of Plekocheilus and Drymaeus in mm. Abbreviations: D, shell diameter; H, shell height; HA, height of aperture; LW, height of last whorl; M, mean; max, maximum; min, minimum; n, number of specimens measured; s, standard deviation; W, number of whorls; WA, width of aperture. For depositories of material, see section on material and methods." httpUri="http://table.plazi.org/id/13CCC7CBFFD1BD60E9A6FEED1B69FE17" pageId="16" pageNumber="41" tableUuid="13CCC7CBFFD1BD60E9A6FEED1B69FE17">Table 1</tableCitation>
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), 2.2 times longer than wide, narrowly rimate with rather convex sides, rather elongated, solid. Colour very variable, whitish-yellowish to yellowish-brown, with the following patterns: a) chestnut-brown axial streaks and three, darker interrupted spiral bands; b) with axial chestnut-brown streaks of varying width, sometimes coalescing; c) axial streaks chestnut-brown and zig-zagging. The spiral bands, if present, are evenly spaced on the last whorl, one at the periphery. Upper whorls in the same ground colour as the rest of the shell or with a pinkish hue; in the latter case the apex is dark reddish-black. Surface somewhat shining, in most cases dull; the upper whorls smooth, on the remaining whorls the growth striae are slightly thickened, most noticeable on the third and fourth whorl. Protoconch with a grating sculpture of axial riblets and spiral striae, which are of equal strength. Whorls 5.2, rather convex; suture slightly impressed and bordered by a whitish to yellowish line below. Aperture elongate-ovate, 1.8 times longer than wide, 0.52 times the total length. Peristome thin, expanded, especially below, whitish with a pink or orange border inside. Columellar margin curved, reflexed, dilated above and receding. Parietal wall with a thin callus, with a pink or orange hue.
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</paragraph>
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<paragraph id="470C9743FFC8BD79E9F4FDFC199DFCF6" blockId="17.[151,1437,152,2034]" pageId="17" pageNumber="42">
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<emphasis id="75C74B51FFC8BD79E9F4FDFC180AFDA8" bold="true" box="[197,323,538,564]" pageId="17" pageNumber="42">Genitalia.</emphasis>
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Proximal part of the penis with a sheath (ca. 1/5 the length of the phallus), distally slightly swollen and passing without external differentiation into the epihallus. The flagellum is slightly tapering, relatively thick, ca. 1/8 the length of the phallus; the distally attached retractor muscle relatively short and thin. Internally three different zones may be distinguished in the penis: proximal part with longitudinal folds of various lengths, the median part with fewer folds bearing cross-folds, the distal part with a bulgy fold and smaller folds at right angles; epiphallus with longitudinal folds, flagellum with short longitudinal ridges. Spermathecal duct somewhat swollen proximally, subcylindrical in its lower part, thereafter constricted towards the elongate-globose spermatheca. The spermatophore is curved, cylindrical and tapering at both ends.
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</paragraph>
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<paragraph id="470C9743FFC8BD79E9F4FC90180AFBCE" blockId="17.[151,1437,152,2034]" pageId="17" pageNumber="42">
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<emphasis id="75C74B51FFC8BD79E9F4FC901830FC0C" bold="true" box="[197,377,886,912]" pageId="17" pageNumber="42">Pallial organs.</emphasis>
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Nephridium elongate-truncated, passing into a very short adrenal (primary) ureter, which tapers into the relatively long and wide adrectal (secondary) ureter. Urinary opening at inner side of the pneumostome. Pericardium situated alongside the nephridium, triangularly shaped; ventriculum about twice the size of the aorta. Pulmonary vein ramified towards the mantle edge; a secundary vein runs parallel under the adrectal ureter, with side veins towards the main pulmonary vein, becoming stronger and ramified near the pneumostome.
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</paragraph>
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</subSubSection>
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<subSubSection id="0FA9C4C8FFC8BD79E9F4FBB81B1FFA67" pageId="17" pageNumber="42" type="materials_examined">
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<paragraph id="470C9743FFC8BD79E9F4FBB81B1FFA67" blockId="17.[151,1437,152,2034]" pageId="17" pageNumber="42">
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<emphasis id="75C74B51FFC8BD79E9F4FBB81830FBE4" bold="true" box="[197,377,1118,1144]" pageId="17" pageNumber="42">
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<typeStatus id="980829E1FFC8BD79E9F4FBB81849FBE4" box="[197,256,1118,1144]" pageId="17" pageNumber="42">Type</typeStatus>
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material.
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</emphasis>
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<collectingCountry id="3FA4D7D3FFC8BD79E8B5FBB81B49FBE4" box="[388,512,1118,1144]" name="Venezuela" pageId="17" pageNumber="42">Venezuela</collectingCountry>
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, Estado Amazonas, Cerro de la Neblina massif,
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<quantity id="804B3AA6FFC8BD79ED0FFBB81DD8FBE4" box="[1086,1169,1118,1144]" metricMagnitude="3" metricUnit="m" metricValue="5.3" pageId="17" pageNumber="42" unit="km" value="5.3">5.3 km</quantity>
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NE Pico Phelps at the base of the escarpment
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<quantity id="804B3AA6FFC8BD79E89CFB6318A1FB03" box="[429,488,1157,1183]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="42" unit="km" value="1.0">1 km</quantity>
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NE Pico Maguire, Camp VII [
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<geoCoordinate id="2287F184FFC8BD79EA64FB631AB6FB03" box="[853,1023,1157,1183]" direction="north" orientation="latitude" pageId="17" pageNumber="42" precision="15" value="0.84444445">00° 50’ 40” N</geoCoordinate>
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<geoCoordinate id="2287F184FFC8BD79ED36FB631DF3FB03" box="[1031,1210,1157,1183]" direction="west" orientation="longitude" pageId="17" pageNumber="42" precision="15" value="-65.969444">65° 58’ 10” W</geoCoordinate>
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], ca.
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<quantity id="804B3AA6FFC8BD79EDC9FB631C1DFB03" box="[1272,1364,1157,1183]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="17" pageNumber="42" unit="m" value="1800.0">1800 m</quantity>
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, R.W. McDiarmid leg.,
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<date id="330DB183FFC8BD79E851FB4A18C4FB5A" box="[352,397,1196,1222]" pageId="17" pageNumber="42">31.i</date>
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─
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<date id="330DB183FFC8BD79E893FB4A1B55FB5A" box="[418,540,1196,1222]" pageId="17" pageNumber="42" value="1985-02-10">10.ii.1985</date>
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||
.
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<typeStatus id="980829E1FFC8BD79EB16FB4A1BDDFB5A" box="[551,660,1196,1222]" pageId="17" pageNumber="42" type="holotype">Holotype</typeStatus>
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UF 284726,
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<typeStatus id="980829E1FFC8BD79EA1EFB4A1AEDFB5A" box="[815,932,1196,1222]" pageId="17" pageNumber="42" type="paratype">paratypes</typeStatus>
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: Ibidem, UF 284725/1, 284730/1); Ibidem, L. Cocroft leg., 10─
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<date id="330DB183FFC8BD79E8A6FB341B5EFB70" box="[407,535,1234,1260]" pageId="17" pageNumber="42" value="1985-02-13">13.ii.1985</date>
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(UF 284736/1); Ibidem, A. Paolillo leg., 10─
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<date id="330DB183FFC8BD79ED6DFB341D95FB70" box="[1116,1244,1234,1260]" pageId="17" pageNumber="42" value="1985-02-13">13.ii.1985</date>
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(UF 284744/1, 284745/1); Ibidem,
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<quantity id="804B3AA6FFC8BD79E8BAFB1F1880FA8F" box="[395,457,1273,1299]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="42" unit="km" value="3.0">3 km</quantity>
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NE Pico Phelps, Camp II [
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<geoCoordinate id="2287F184FFC8BD79EA29FB1F1A80FA8F" box="[792,969,1273,1299]" direction="north" orientation="latitude" pageId="17" pageNumber="42" precision="15" value="0.8277778">00° 49’ 40” N</geoCoordinate>
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||
<geoCoordinate id="2287F184FFC8BD79EAE0FB1F1DC5FA8F" box="[977,1164,1273,1299]" direction="west" orientation="longitude" pageId="17" pageNumber="42" precision="15" value="-65.98333">65° 59’ 00” W</geoCoordinate>
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], 2085─
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<quantity id="804B3AA6FFC8BD79EDC2FB1F1C1BFA8F" box="[1267,1362,1273,1299]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="17" pageNumber="42" unit="m" value="2100.0">2100 m</quantity>
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, R.W. McDiarmid leg.,
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<date id="330DB183FFC8BD79E85AFAC618A2FAA6" box="[363,491,1312,1338]" pageId="17" pageNumber="42" value="1985-02-28">28.ii.1985</date>
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, in
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<taxonomicName id="80B3ECC0FFC8BD79EB28FAC61BE9FAA5" box="[537,672,1312,1337]" class="Gastropoda" family="Cancellariidae" genus="Brocchinia" kingdom="Animalia" order="Neogastropoda" pageId="17" pageNumber="42" phylum="Mollusca" rank="genus">
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<emphasis id="75C74B51FFC8BD79EB28FAC61BE9FAA5" box="[537,672,1312,1337]" italics="true" pageId="17" pageNumber="42">Brocchinia</emphasis>
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</taxonomicName>
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plants (
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<collectionCode id="21A20F86FFC8BD79EA32FAC61A29FAA6" LSID="urn:lsid:biocol.org:col:34992" box="[771,864,1312,1338]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="17" pageNumber="42">RMNH</collectionCode>
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||
<date id="330DB183FFC8BD79EA59FAC61A8CFAA6" box="[872,965,1312,1338]" pageId="17" pageNumber="42" value="26-11-20">112026</date>
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||
/2, UF 284714/2, 284715/1, 284719/2, 284720/1, 284721/1, 284722/2 +
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<specimenCount id="51B55CCAFFC8BD79EB11FAA01B2FFAFC" box="[544,614,1350,1376]" pageId="17" pageNumber="42" type="generic">1 juv.</specimenCount>
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||
); Ibidem,
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||
<quantity id="804B3AA6FFC8BD79EBECFAA01A50FAFC" box="[733,793,1350,1376]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="42" unit="km" value="2.0">2 km</quantity>
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NNW Pico Phelps, in leaf litter, Camp V [
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<geoCoordinate id="2287F184FFC8BD79EC21FAA019E5FA1B" direction="north" orientation="latitude" pageId="17" pageNumber="42" precision="15" value="0.8194444">00° 49’ 10” N</geoCoordinate>
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||
<geoCoordinate id="2287F184FFC8BD79E984FA8B1822FA1B" box="[181,363,1389,1415]" direction="west" orientation="longitude" pageId="17" pageNumber="42" precision="15" value="-66.00056">66° 00’ 02” W</geoCoordinate>
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], ca.
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<quantity id="804B3AA6FFC8BD79E89BFA8B1B41FA1B" box="[426,520,1389,1415]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="17" pageNumber="42" unit="m" value="1300.0">1300 m</quantity>
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, F.G. Thompson leg.,
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<date id="330DB183FFC8BD79EA21FA8B1AC7FA1B" box="[784,910,1389,1415]" pageId="17" pageNumber="42" value="1984-04-13">13.iv.1984</date>
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(UF 284759/1); Idem, Idem,
|
||
<quantity id="804B3AA6FFC8BD79EDC4FA8B1C1EFA1B" box="[1269,1367,1389,1415]" metricMagnitude="3" metricUnit="m" metricValue="6.25" pageId="17" pageNumber="42" unit="km" value="6.25">6.25 km</quantity>
|
||
NNE Pico Phelps, Camp XI [
|
||
<geoCoordinate id="2287F184FFC8BD79E898FA721B06FA32" box="[425,591,1428,1454]" direction="north" orientation="latitude" pageId="17" pageNumber="42" precision="15" value="0.8666667">00° 52’ 00” N</geoCoordinate>
|
||
<geoCoordinate id="2287F184FFC8BD79EB67FA721A4EFA32" box="[598,775,1428,1454]" direction="west" orientation="longitude" pageId="17" pageNumber="42" precision="15" value="-65.97833">65° 58’ 42” W</geoCoordinate>
|
||
], 1400─
|
||
<quantity id="804B3AA6FFC8BD79EA58FA721A8CFA32" box="[873,965,1428,1454]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="42" unit="m" value="1500.0">1500 m</quantity>
|
||
, R.W. McDiarmid leg.,
|
||
<date id="330DB183FFC8BD79EDEAFA721C79FA32" box="[1243,1328,1428,1454]" pageId="17" pageNumber="42" value="1985-02">ii.1985</date>
|
||
, in forest (UF 284748/
|
||
<specimenCount id="51B55CCAFFC8BD79E804FA5C18E6FA48" box="[309,431,1466,1492]" pageId="17" pageNumber="42" type="subadult">1 subadult</specimenCount>
|
||
); Ibidem, 2.0 km SW Neblina Base Camp,
|
||
<quantity id="804B3AA6FFC8BD79EA83FA5C1D48FA48" box="[946,1025,1466,1492]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="42" unit="m" value="200.0">200 m</quantity>
|
||
, R.W. McDiarmid leg.,
|
||
<date id="330DB183FFC8BD79EC2CFA5C1CDFFA48" box="[1309,1430,1466,1492]" pageId="17" pageNumber="42" value="1985-02-22">22.ii.1985</date>
|
||
, in tree
|
||
<quantity id="804B3AA6FFC8BD79E9D9FA07185CFA67" box="[232,277,1505,1531]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="42" unit="m" value="2.0">2 m</quantity>
|
||
off ground (UF 284747/1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FA9C4C8FFC8BD7AE9F4F9E11990FEBA" lastPageId="18" lastPageNumber="43" pageId="17" pageNumber="42" type="discussion">
|
||
<paragraph id="470C9743FFC8BD79E9F4F9E11C48F8AC" blockId="17.[151,1437,152,2034]" pageId="17" pageNumber="42">
|
||
<emphasis id="75C74B51FFC8BD79E9F4F9E11876F9BD" bold="true" box="[197,319,1543,1569]" pageId="17" pageNumber="42">Remarks.</emphasis>
|
||
<collectingCountry id="3FA4D7D3FFC8BD79E876F9EE1821F9BE" box="[327,360,1544,1570]" name="American Samoa" pageId="17" pageNumber="42">As</collectingCountry>
|
||
often seen in
|
||
<taxonomicName id="80B3ECC0FFC8BD79EB21F9EE1BC0F9BD" box="[528,649,1544,1569]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="42" phylum="Mollusca" rank="genus">
|
||
<emphasis id="75C74B51FFC8BD79EB21F9EE1BC0F9BD" box="[528,649,1544,1569]" italics="true" pageId="17" pageNumber="42">Drymaeus</emphasis>
|
||
</taxonomicName>
|
||
, this species shows a great deal of variation in colour and pattern. One specimen (UF 284730) is evenly coloured, the last whorl lighter than the upper whorls, the axial streaks hardly noticeable. This species is also rather variable in shape. Three specimens are somewhat aberrant and have hardly any transition between columellar and parietal margin, giving the aperture a skewed appearance. Of these, one specimen (UF 284759) has the columellar margin straight and the aperture is ear-shaped. In two other specimens (UF 284747, 284748) the aperture is also somewhat ear-shaped, but the columella shows a slight fold. The specimens have been collected at lower elevations (1450, 1300 and
|
||
<quantity id="804B3AA6FFC8BD79ED58F9161DFAF896" box="[1129,1203,1776,1802]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="42" unit="m" value="200.0">200 m</quantity>
|
||
respectively), but it is not clear from the material at hand if there is a correlation between elevation and morphology.
|
||
</paragraph>
|
||
<paragraph id="470C9743FFC8BD7AE9F4F8DB1990FEBA" blockId="17.[151,1437,152,2034]" lastBlockId="18.[151,1436,152,448]" lastPageId="18" lastPageNumber="43" pageId="17" pageNumber="42">
|
||
<emphasis id="75C74B51FFC8BD79E9F4F8DB1B71F8CB" bold="true" box="[197,568,1853,1879]" pageId="17" pageNumber="42">Comparison with other taxa.</emphasis>
|
||
This species is closely allied to
|
||
<taxonomicName id="80B3ECC0FFC8BD79EAF2F8DB187FF8E2" authority="Breure & Eskens, 1981" authorityName="Breure & Eskens" authorityYear="1981" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="42" phylum="Mollusca" rank="species" species="yapacanensis" subGenus="Drymaeus">
|
||
<emphasis id="75C74B51FFC8BD79EAF2F8DB1C6AF8CA" box="[963,1315,1853,1878]" italics="true" pageId="17" pageNumber="42">Drymaeus (D.) yapacanensis</emphasis>
|
||
<bibRefCitation id="2322EAB2FFC8BD79EC1AF8DB187FF8E2" author="Breure" pageId="17" pageNumber="49" refString="Breure, A. S. H. & Eskens, A. A. C. (1981) Notes on and descriptions of Bulimulidae (Mollusca, Gastropoda), II. Zoologische Verhandelingen, 186, 1 ─ 111." type="journal article" year="1981">Breure & Eskens, 1981</bibRefCitation>
|
||
</taxonomicName>
|
||
, from which it differs in (1) being stouter; (2) having a more swollen last whorl; (3) having less whorls. This new species is also very similar to
|
||
<taxonomicName id="80B3ECC0FFC8BD79EBF4F86D1DEAF838" authority="Haas, 1955" authorityName="Haas" authorityYear="1955" box="[709,1187,1930,1956]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="42" phylum="Mollusca" rank="species" species="steyermarki" subGenus="Drymaeus">
|
||
<emphasis id="75C74B51FFC8BD79EBF4F86D1D4CF838" box="[709,1029,1931,1956]" italics="true" pageId="17" pageNumber="42">Drymaeus (D.) steyermarki</emphasis>
|
||
(
|
||
<bibRefCitation id="2322EAB2FFC8BD79ED24F86C1DD2F838" author="Haas" box="[1045,1179,1930,1956]" pageId="17" pageNumber="49" refString="Haas, F. (1955) On some small collections of inland shells from South America. Fieldiana, Zoology, 34, 361 ─ 387." type="journal article" year="1955">Haas, 1955</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from which it differs in (1) being larger; (2) missing the thread-like columellar fold. It may also be compared to
|
||
<taxonomicName id="80B3ECC0FFC8BD79EDFCF8571870F86E" authority="Haas, 1955" authorityName="Haas" authorityYear="1955" class="Gastropoda" family="Bulimulidae" genus="Bulimulus" higherTaxonomySource="CoL" kingdom="Animalia" order="Stylommatophora" pageId="17" pageNumber="42" phylum="Mollusca" rank="species" species="extraneus" subGenus="Drymaeus">
|
||
<emphasis id="75C74B51FFC8BD79EDFCF8571CD5F856" box="[1229,1436,1969,1994]" italics="true" pageId="17" pageNumber="42">D. (D.) extraneus</emphasis>
|
||
(
|
||
<bibRefCitation id="2322EAB2FFC8BD79E991F83E1866F86E" author="Haas" box="[160,303,2008,2034]" pageId="17" pageNumber="49" refString="Haas, F. (1955) On some small collections of inland shells from South America. Fieldiana, Zoology, 34, 361 ─ 387." type="journal article" year="1955">Haas, 1955</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, from which it differs in (1) having the surface less shining; (2) the less dilated columellar margin; (3) the pink lip; (4) the different colour pattern.
|
||
<taxonomicName id="80B3ECC0FFCBBD7AEA07FF7E1C6FFF2E" authority="Albers, 1854" authorityName="Albers" authorityYear="1854" box="[822,1318,152,178]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="43" phylum="Mollusca" rank="species" species="edmuelleri" subGenus="Drymaeus">
|
||
<emphasis id="75C74B51FFCBBD7AEA07FF7E1D27FF2D" box="[822,1134,152,177]" italics="true" pageId="18" pageNumber="43">Drymaeus (D.) edmuelleri</emphasis>
|
||
(
|
||
<bibRefCitation id="2322EAB2FFCBBD7AED4EFF7E1C57FF2E" author="Albers" box="[1151,1310,152,178]" pageId="18" pageNumber="49" refString="Albers, J. C. (1854) Diagnosen neuer Bulimus - Arten. Malakologische Blatter, 1, 30 ─ 32." type="journal article" year="1854">Albers, 1854</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, of which the
|
||
<typeStatus id="980829E1FFCBBD7AE9F3FF5819BDFF44" box="[194,244,190,216]" pageId="18" pageNumber="43">type</typeStatus>
|
||
material (ZMB 111.929) has been studied, resembles this new species but differs in (1) being smaller and slenderer; (2) having the last whorl less swollen and somewhat protruded at base; (3) having straight sides.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0FA9C4C8FFCBBD7AE9F4FED41D81FE5C" pageId="18" pageNumber="43" type="etymology">
|
||
<paragraph id="470C9743FFCBBD7AE9F4FED41D81FE5C" blockId="18.[151,1436,152,448]" pageId="18" pageNumber="43">
|
||
<emphasis id="75C74B51FFCBBD7AE9F4FED4181FFED0" bold="true" box="[197,342,306,332]" pageId="18" pageNumber="43">Etymology.</emphasis>
|
||
Latin
|
||
<taxonomicName id="80B3ECC0FFCBBD7AE899FED41884FED7" box="[424,461,306,331]" class="Gastropoda" family="Orthalicidae" genus="Drymaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="18" pageNumber="43" phylum="Mollusca" rank="species" species="rex">
|
||
<emphasis id="75C74B51FFCBBD7AE899FED41884FED7" box="[424,461,306,331]" italics="true" pageId="18" pageNumber="43">rex</emphasis>
|
||
</taxonomicName>
|
||
, king. Named in honour of Roy W. McDiarmid who collected most of the
|
||
<typeStatus id="980829E1FFCBBD7AEC59FED41CD5FED0" box="[1384,1436,306,332]" pageId="18" pageNumber="43">type</typeStatus>
|
||
material. “Roy” is the Old French word for king, but also the Anglicized spelling of a nickname from Scottish Gaelic ruadh (red, referring to the reddish apex in some specimens). The name also refers to the majestical appearance of this species when collected alive. The epithet is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |