303 lines
36 KiB
XML
303 lines
36 KiB
XML
<document id="E8495FDCE5F84158ECB60EDCDADA26A5" ID-CLB-Dataset="21869" ID-DOI="10.5852/ejt.2016.236" ID-GBIF-Dataset="2dfee902-a7fc-472b-bf1d-27029b3504f3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3847840" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590436812908" checkinUser="valdenar" docAuthor="Herbert, David G." docDate="2016" docId="C146B323FFB8FFE1FDAF9D60FC65F3A2" docLanguage="en" docName="ejt-2016-236.pdf.imf" docOrigin="European Journal of Taxonomy 236" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Gulella hadroglossa Herbert 2016, sp. nov." docType="treatment" docUuid="B0429B6A-4C92-40BB-B2E7-A707A32ADE96" docUuidSource="ZooBank" docVersion="10" lastPageNumber="9" masterDocId="3D7FCB5BFFBEFFE9FFC0994BFF82F57F" masterDocTitle="New narrow-range endemic land snails from the sky islands of northern South Africa (Gastropoda: Streptaxidae and Urocyclidae)" masterLastPageNumber="29" masterPageNumber="1" pageNumber="7" updateTime="1698841246942" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:mods id="0D920E02F64B17868A644BAAF4D9FAAD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4438DA8328B11BE1ACEEF3596751910D">
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<mods:title id="0659BD6558BB06506D74D6BB8DE00BBA">New narrow-range endemic land snails from the sky islands of northern South Africa (Gastropoda: Streptaxidae and Urocyclidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="9B705596525831ADCC00C0C0B12E5FD9" type="personal">
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<mods:role id="A580071E1EF266F8D7482ECD0150A2CE">
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<mods:roleTerm id="DC5591A747ADE8B19772717A35E60BB6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="71664809B3530A0514EAE8970A3ADC0F">Herbert, David G.</mods:namePart>
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<mods:typeOfResource id="9E27A7EC3FE3BD880FD7C79398D84675">text</mods:typeOfResource>
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<mods:title id="58F09A459233ADB272BF12A41EB8C6E5">European Journal of Taxonomy</mods:title>
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</mods:titleInfo>
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<mods:part id="C52B613E75B5ACF3323D1264DB1E7985">
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<mods:date id="D4285A25C7BEE0AC842FD7B430844322">2016</mods:date>
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<mods:detail id="18F1D09D8A2C979DB08DFCE34BC36AAB" type="pubDate">
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<mods:number id="6FC13F2706E6815B878C41C7DDCFF62D">2016-10-12</mods:number>
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</mods:detail>
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<mods:detail id="163C8186A6AF1166D2488A2636308A73" type="volume">
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<mods:number id="0C672F6F7A6C376CDDFF200778889E76">236</mods:number>
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</mods:detail>
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<mods:start id="48BA2DF37719026CD9873378B170840C">1</mods:start>
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<mods:end id="CF30F999B89CE7DAE1C93F143BDC6E45">29</mods:end>
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<mods:classification id="6F84869E3A52D205B821FD6E3930CE15">journal article</mods:classification>
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<mods:identifier id="AE104E4C3EB33998C4FE2CFB8A04F374" type="CLB-Dataset">21869</mods:identifier>
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<mods:identifier id="9B9D9021B77EE683FBE959E9098060C6" type="DOI">10.5852/ejt.2016.236</mods:identifier>
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<mods:identifier id="B9F66F6EA0442BDDEFF9EF12954F6FC5" type="GBIF-Dataset">2dfee902-a7fc-472b-bf1d-27029b3504f3</mods:identifier>
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<mods:identifier id="E68DB458D14592924527A6D5835B6CE1" type="ISSN">2118-9773</mods:identifier>
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<mods:identifier id="878F57D8E8ED3B2CF55939806F58647E" type="Zenodo-Dep">3847840</mods:identifier>
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<mods:identifier id="B6D8B0649B9F374B13597929CF5519B7" type="ZooBank">B51BF718-79F5-47F5-8740-BA181CE88257</mods:identifier>
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<treatment id="C146B323FFB8FFE1FDAF9D60FC65F3A2" ID-DOI="http://doi.org/10.5281/zenodo.3854758" ID-GBIF-Taxon="164282651" ID-Zenodo-Dep="3854758" LSID="urn:lsid:zoobank.org:act:B0429B6A-4C92-40BB-B2E7-A707A32ADE96" httpUri="http://treatment.plazi.org/id/C146B323FFB8FFE1FDAF9D60FC65F3A2" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7">
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<subSubSection id="01F551BEFFB8FFEFFDAF9D60FB4FF117" pageId="6" pageNumber="7" type="nomenclature">
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<paragraph id="49500235FFB8FFEFFDAF9D60FC47F139" blockId="6.[359,1229,1067,1164]" box="[623,965,1067,1094]" pageId="6" pageNumber="7">
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<heading id="1218B559FFB8FFEFFDAF9D60FC47F139" box="[623,965,1067,1094]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="7" reason="2">
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<taxonomicName id="8EEF79B6FFB8FFEFFDAF9D60FCE1F139" ID-CoL="3HLRL" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[623,867,1067,1094]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="hadroglossa" status="sp. nov.">
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<emphasis id="7B9BDE27FFB8FFEFFDAF9D60FCE1F139" bold="true" box="[623,867,1067,1094]" italics="true" pageId="6" pageNumber="7">Gulella hadroglossa</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="60A8635CFFB8FFEFFCAA9D67FC47F139" box="[874,965,1068,1094]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="49500235FFB8FFEFFEA79D05FB4FF117" blockId="6.[359,1229,1067,1164]" box="[359,1229,1102,1128]" pageId="6" pageNumber="7">
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<uri id="3D7E0E37FFB8FFEFFEA79D05FB4FF117" box="[359,1229,1102,1128]" pageId="6" pageNumber="7">
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urn:lsid:zoobank.org:act:
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<uuid id="3D4938E0FFB8FFEFFD459D05FB4FF117" box="[645,1229,1102,1128]" pageId="6" pageNumber="7">B0429B6A-4C92-40BB-B2E7-A707A32ADE96</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="01F551BEFFB8FFEFFD389D3AFCBEF1F3" box="[760,828,1137,1164]" pageId="6" pageNumber="7" type="description">
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<paragraph id="49500235FFB8FFEFFD389D3AFCBEF1F3" blockId="6.[359,1229,1067,1164]" box="[760,828,1137,1164]" pageId="6" pageNumber="7">
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<figureCitation id="D1D41EB0FFB8FFEFFD389D3AFCBEF1F3" box="[760,828,1137,1164]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[193,1394,777,1884]" captionTargetId="figure@7.[189,1399,765,1889]" captionTargetPageId="7" captionText="Fig. 3. Gulella hadroglossa sp. nov. A–E. Holotype, height 3.0 mm, diameter 1.5 mm (NMSA P0230/ T4084). F–G. Juvenile specimen showing apertural dentition, diameter 1.5 mm (paratype, NMSA P0417/T4085)." figureDoi="http://doi.org/10.5281/zenodo.3847852" httpUri="https://zenodo.org/record/3847852/files/figure.png" pageId="6" pageNumber="7">Fig. 3</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="01F551BEFFB8FFEFFF7D9DF8FD85F060" pageId="6" pageNumber="7" type="etymology">
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<paragraph id="49500235FFB8FFEFFF7D9DF8FEC6F1B2" blockId="6.[189,324,1203,1229]" box="[189,324,1203,1229]" pageId="6" pageNumber="7">
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<heading id="1218B559FFB8FFEFFF7D9DF8FEC6F1B2" bold="true" box="[189,324,1203,1229]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
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<emphasis id="7B9BDE27FFB8FFEFFF7D9DF8FEC6F1B2" bold="true" box="[189,324,1203,1229]" pageId="6" pageNumber="7">Etymology</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="49500235FFB8FFEFFF7D9DA9FD85F060" blockId="6.[189,1398,1249,1311]" pageId="6" pageNumber="7">
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||
From
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||
<emphasis id="7B9BDE27FFB8FFEFFEC89DAAFEDAF184" box="[264,344,1249,1275]" italics="true" pageId="6" pageNumber="7">hadros</emphasis>
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||
(Gr.) ‘well-developed, great, strong’, and
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<emphasis id="7B9BDE27FFB8FFEFFCA19DAAFC2EF184" box="[865,940,1249,1275]" italics="true" pageId="6" pageNumber="7">glossa</emphasis>
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(Gr.) ‘tongue’; referring to the large, tongue-like parietal lamella.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="01F551BEFFB8FFEEFF7D9C0DFD57F72F" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="materials_examined">
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<paragraph id="49500235FFB8FFEFFF7D9C0DFE28F01F" blockId="6.[189,426,1350,1376]" box="[189,426,1350,1376]" pageId="6" pageNumber="7">
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<heading id="1218B559FFB8FFEFFF7D9C0DFE28F01F" bold="true" box="[189,426,1350,1376]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
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||
<emphasis id="7B9BDE27FFB8FFEFFF7D9C0DFE28F01F" bold="true" box="[189,426,1350,1376]" pageId="6" pageNumber="7">Material examined</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="49500235FFB8FFEFFF2A9C3EFEDEF0F0" blockId="6.[189,1399,1397,1529]" box="[234,348,1397,1423]" pageId="6" pageNumber="7">
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<emphasis id="7B9BDE27FFB8FFEFFF2A9C3EFEDEF0F0" bold="true" box="[234,348,1397,1423]" pageId="6" pageNumber="7">
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<typeStatus id="9654BC97FFB8FFEFFF2A9C3EFEDEF0F0" box="[234,348,1397,1423]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="49500235FFB8FFEFFF7D9CD3FB84F086" blockId="6.[189,1399,1397,1529]" pageId="6" pageNumber="7">
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<materialsCitation id="F9870868FFB8FFEFFF7D9CD3FB84F086" ID-GBIF-Occurrence="2625465305" collectingDate="2014-12-04" collectionCode="NMSA" collectorName="D. Herbert & L. Davis & M. Cole" country="South Africa" elevation="1545" latitude="-24.56847" location="Picnic Trail" longLatPrecision="1" longitude="30.8592" municipality="Mariepskop Forest Reserve" pageId="6" pageNumber="7" specimenCode="P0230" specimenCount="1" stateProvince="Mpumalanga" typeStatus="holotype">
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<collectingCountry id="31F842A5FFB8FFEFFF7D9CD3FE14F0CD" box="[189,406,1432,1458]" name="South Africa" pageId="6" pageNumber="7">SOUTH AFRICA</collectingCountry>
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:
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<collectingRegion id="8B2BCCD7FFB8FFEFFE629CD3FDC2F0CD" box="[418,576,1432,1458]" country="South Africa" name="Mpumalanga" pageId="6" pageNumber="7">Mpumalanga</collectingRegion>
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,
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<collectingMunicipality id="A934984FFFB8FFEFFD8C9CD3FC11F0CD" box="[588,915,1432,1458]" pageId="6" pageNumber="7">Mariepskop Forest Reserve</collectingMunicipality>
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,
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<location id="4C3054EEFFB8FFEFFC609CD3FBAEF0CD" LSID="urn:lsid:plazi:treatment:C146B323FFB8FFE1FDAF9D60FC65F3A2:4C3054EEFFB8FFEFFC609CD3FBAEF0CD" box="[928,1068,1432,1458]" country="South Africa" latitude="-24.56847" longLatPrecision="1" longitude="30.8592" municipality="Mariepskop Forest Reserve" name="Picnic Trail" pageId="6" pageNumber="7" stateProvince="Mpumalanga">Picnic Trail</location>
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,
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,
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<geoCoordinate id="2CDB64F2FFB8FFEFFB1B9CD3FAF3F0CD" box="[1243,1393,1432,1459]" degrees="30.85920" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="30.8592">30.85920° E</geoCoordinate>
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,
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<quantity id="8E17AFD0FFB8FFEFFF7D9CF0FE95F0A9" box="[189,279,1467,1494]" metricMagnitude="3" metricUnit="m" metricValue="1.545" pageId="6" pageNumber="7" unit="m" value="1545.0">
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<elevation id="C2C2E506FFB8FFEFFF7D9CF0FE95F0A9" box="[189,279,1467,1494]" metricMagnitude="3" metricUnit="m" metricValue="1.545" pageId="6" pageNumber="7" unit="m" value="1545.0">1545 m</elevation>
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</quantity>
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||
, northern mist-belt forest, in leaf-litter,
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||
<collectorName id="E41A67E3FFB8FFEFFD259CF7FCE7F0AA" box="[741,869,1467,1494]" pageId="6" pageNumber="7">D. Herbert</collectorName>
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,
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<collectorName id="E41A67E3FFB8FFEFFCB09CF7FC56F0AA" box="[880,980,1467,1494]" pageId="6" pageNumber="7">L. Davis</collectorName>
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&
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<collectorName id="E41A67E3FFB8FFEFFC399CF7FBD9F0AA" box="[1017,1115,1467,1494]" pageId="6" pageNumber="7">M. Cole</collectorName>
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, stn 14-27,
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<date id="3D5124F5FFB8FFEFFB229CF0FAF0F0AA" box="[1250,1394,1467,1494]" pageId="6" pageNumber="7" value="2014-12-04">
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<collectingDate id="2D15DD1DFFB8FFEFFB229CF0FAF0F0AA" box="[1250,1394,1467,1494]" pageId="6" pageNumber="7" value="2014-12-04">4 Dec. 2014</collectingDate>
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</date>
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||
, height 3.0 mm, diameter
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||
<quantity id="8E17AFD0FFB8FFEFFE249C95FDBCF086" box="[484,574,1502,1529]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="mm" value="1.5">1.5 mm</quantity>
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(
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<collectionCode id="2FFE9AF0FFB8FFEFFD8D9C95FD20F087" box="[589,674,1502,1528]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="6" pageNumber="7" type="Museum">NMSA</collectionCode>
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<specimenCode id="1949AA4EFFB8FFEFFD699C95FD77F087" box="[681,757,1502,1528]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="6" pageNumber="7" type="Museum">P0230</specimenCode>
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||
/T4084, dry specimen).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="49500235FFB8FFEFFF2A9F6BFEE5F345" blockId="6.[189,1399,1568,1911]" box="[234,359,1568,1594]" pageId="6" pageNumber="7">
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<emphasis id="7B9BDE27FFB8FFEFFF2A9F6BFEE5F345" bold="true" box="[234,359,1568,1594]" pageId="6" pageNumber="7">
|
||
<typeStatus id="9654BC97FFB8FFEFFF2A9F6BFEE5F345" box="[234,359,1568,1594]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
</paragraph>
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<paragraph id="49500235FFB8FFEFFF7D9F08FC7AF208" blockId="6.[189,1399,1568,1911]" pageId="6" pageNumber="7">
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<collectingCountry id="31F842A5FFB8FFEFFF7D9F08FE1BF322" box="[189,409,1603,1629]" name="South Africa" pageId="6" pageNumber="7">SOUTH AFRICA</collectingCountry>
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||
:
|
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<collectingRegion id="8B2BCCD7FFB8FFEFFE689F08FDC4F322" box="[424,582,1603,1629]" country="South Africa" name="Mpumalanga" pageId="6" pageNumber="7">Mpumalanga</collectingRegion>
|
||
, same data as
|
||
<typeStatus id="9654BC97FFB8FFEFFD3D9F08FCE6F322" box="[765,868,1603,1629]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFB8FFEFFCB79F08FC4EF322" box="[887,972,1603,1629]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="6" pageNumber="7" type="Museum">NMSA</collectionCode>
|
||
P0417/T4085, five dry specimens;
|
||
<collectionCode id="2FFE9AF0FFB8FFEFFF7D9F2CFEAEF3FE" box="[189,300,1639,1665]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="6" pageNumber="7" type="Museum">NHMUK</collectionCode>
|
||
20160038, one dry specimen;
|
||
<collectionCode id="2FFE9AF0FFB8FFEFFD799F2DFC96F3FE" box="[697,788,1638,1665]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
|
||
.5004142, one dry specimen); Mariepskop Forest Reserve,
|
||
<geoCoordinate id="2CDB64F2FFB8FFEFFEEE9FC2FE41F3DC" box="[302,451,1673,1700]" degrees="24.55117" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-24.55117">24.55117° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFB8FFEFFE149FC2FDEEF3DB" box="[468,620,1673,1700]" degrees="30.89395" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="30.89395">30.89395° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFB8FFEFFDBD9FC2FD59F3DB" box="[637,731,1673,1700]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="6" pageNumber="7" unit="m" value="1460.0">1460 m</quantity>
|
||
, indigenous Afromontane forest, in leaf-litter on forest floor, J.L. Horn,
|
||
<date id="3D5124F5FFB8FFEFFEBE9FE6FD97F3B8" box="[382,533,1708,1735]" pageId="6" pageNumber="7" value="2005-04-23">23 Apr. 2005</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFB8FFEFFDE39FE6FDFAF3B8" box="[547,632,1709,1735]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="6" pageNumber="7" type="Museum">NMSA</collectionCode>
|
||
W3500/T4081, four dry specimens); Mariepskop Forest Reserve, Bushpig Trail,
|
||
<geoCoordinate id="2CDB64F2FFB8FFEFFEB09F84FD86F395" box="[368,516,1743,1770]" degrees="24.56795" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-24.56795">24.56795° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFB8FFEFFDD39F9BFD28F395" box="[531,682,1744,1770]" degrees="30.86138" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="30.86138">30.86138° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFB8FFEFFD789F84FC97F395" box="[696,789,1743,1770]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="6" pageNumber="7" unit="m" value="1520.0">1520 m</quantity>
|
||
, northern mist-belt forest, in leaf-litter, D. Herbert, L. Davis & M. Cole, stn 14-22a,
|
||
<date id="3D5124F5FFB8FFEFFD8C9FB8FD5CF272" box="[588,734,1779,1805]" pageId="6" pageNumber="7" value="2014-12-03">3 Dec. 2014</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFB8FFEFFD2F9FB8FCC6F272" box="[751,836,1779,1805]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="6" pageNumber="7" type="Museum">NMSA</collectionCode>
|
||
P0263/T4082, one dry specimen); Mariepskop Forest Reserve, east facing slope,
|
||
<geoCoordinate id="2CDB64F2FFB8FFEFFD8B9E5DFD5FF24F" box="[587,733,1814,1841]" degrees="24.57108" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-24.57108">24.57108° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFB8FFEFFD2A9E5DFCFCF24E" box="[746,894,1814,1841]" degrees="30.86014" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="30.86014">30.86014° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFB8FFEFFC4A9E5DFC67F24E" box="[906,997,1814,1841]" metricMagnitude="3" metricUnit="m" metricValue="1.519" pageId="6" pageNumber="7" unit="m" value="1519.0">1519 m</quantity>
|
||
, leg. M. Cole,
|
||
<date id="3D5124F5FFB8FFEFFB519E5DFAAAF24F" box="[1169,1320,1814,1840]" pageId="6" pageNumber="7" value="2010-10-18">18 Oct. 2010</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFB8FFEFFAF79E5DFAF5F24F" box="[1335,1399,1814,1840]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="6" pageNumber="7">ELM</collectionCode>
|
||
D16635/T033, three dry specimens); Mariepskop Forest Reserve, in forest beside road at
|
||
<quantity id="8E17AFD0FFB8FFEFFB0E9E72FAF3F22B" box="[1230,1393,1849,1876]" metricMagnitude="3" metricUnit="m" metricValue="1.61" metricValueMax="1.7" metricValueMin="1.52" pageId="6" pageNumber="7" unit="m" value="1610.0" valueMax="1700.0" valueMin="1520.0">1520–1700 m</quantity>
|
||
, leg. M. Cole,
|
||
<date id="3D5124F5FFB8FFEFFE9E9E16FD89F208" box="[350,523,1885,1911]" pageId="6" pageNumber="7" value="2014-12-03" valueMax="2014-12-04" valueMin="2014-12-03">3–4 Dec. 2014</date>
|
||
(
|
||
<collectionCode id="2FFE9AF0FFB8FFEFFDD99E16FDDBF208" box="[537,601,1885,1911]" country="South Africa" httpUri="http://grbio.org/cool/9trn-613c" name="East London Museum" pageId="6" pageNumber="7">ELM</collectionCode>
|
||
D18019/T034, six dry specimens).
|
||
</paragraph>
|
||
<paragraph id="49500235FFB9FFEEFF7D9840FEDFF45A" blockId="7.[189,349,267,293]" box="[189,349,267,293]" pageId="7" pageNumber="8">
|
||
<heading id="1218B559FFB9FFEEFF7D9840FEDFF45A" bold="true" box="[189,349,267,293]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
|
||
<emphasis id="7B9BDE27FFB9FFEEFF7D9840FEDFF45A" bold="true" box="[189,349,267,293]" pageId="7" pageNumber="8">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFB9FFEEFF7D9871FE9FF408" blockId="7.[189,1399,313,375]" pageId="7" pageNumber="8">
|
||
<collectingCountry id="31F842A5FFB9FFEEFF7D9871FE17F42B" box="[189,405,314,340]" name="South Africa" pageId="7" pageNumber="8">SOUTH AFRICA</collectingCountry>
|
||
:
|
||
<collectingRegion id="8B2BCCD7FFB9FFEEFE629872FDC2F42C" box="[418,576,313,339]" country="South Africa" name="Mpumalanga" pageId="7" pageNumber="8">Mpumalanga</collectingRegion>
|
||
, Mariepskop Forest Reserve, Picnic Trail,
|
||
<geoCoordinate id="2CDB64F2FFB9FFEEFBF99872FB4FF42B" box="[1081,1229,313,340]" degrees="24.56847" direction="south" orientation="latitude" pageId="7" pageNumber="8" precision="1" value="-24.56847">24.56847° S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="2CDB64F2FFB9FFEEFB1B9872FAF3F42B" box="[1243,1393,313,340]" degrees="30.85920" direction="east" orientation="longitude" pageId="7" pageNumber="8" precision="1" value="30.8592">30.85920° E</geoCoordinate>
|
||
,
|
||
<quantity id="8E17AFD0FFB9FFEEFF7D9817FE9BF408" box="[189,281,348,375]" metricMagnitude="3" metricUnit="m" metricValue="1.545" pageId="7" pageNumber="8" unit="m" value="1545.0">1545 m</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="49500235FFB9FFEEFF7D98D5FEE7F4C7" blockId="7.[189,357,414,440]" box="[189,357,414,440]" pageId="7" pageNumber="8">
|
||
<heading id="1218B559FFB9FFEEFF7D98D5FEE7F4C7" bold="true" box="[189,357,414,440]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
|
||
<emphasis id="7B9BDE27FFB9FFEEFF7D98D5FEE7F4C7" bold="true" box="[189,357,414,440]" pageId="7" pageNumber="8">Identification</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFB9FFEEFF7D9886FD57F72F" blockId="7.[189,1399,461,593]" pageId="7" pageNumber="8">
|
||
A small species of
|
||
<taxonomicName id="8EEF79B6FFB9FFEEFE569886FE6CF498" authorityName="Pfeiffer" authorityYear="1856" box="[406,494,461,487]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7B9BDE27FFB9FFEEFE569886FE6CF498" box="[406,494,461,487]" italics="true" pageId="7" pageNumber="8">Gulella</emphasis>
|
||
</taxonomicName>
|
||
with a smooth, glossy, pupiform shell and 3-fold apertural dentition including a very large, oblique, tongue-like parietal lamella projecting far beyond the peristome; a mid-labral plate, the upper edge of which forms a narrow ridge-like tooth that runs obliquely inward and downward; a horizontal in-running tooth at columella base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="1D9052BDFFB9FFEEFF7D9EC1FEECF293" ID-DOI="http://doi.org/10.5281/zenodo.3847852" ID-Zenodo-Dep="3847852" httpUri="https://zenodo.org/record/3847852/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1930,1956]" targetBox="[193,1394,777,1884]" targetPageId="7">
|
||
<paragraph id="49500235FFB9FFEEFF7D9EC1FEECF293" blockId="7.[189,1399,1930,2028]" pageId="7" pageNumber="8">
|
||
<emphasis id="7B9BDE27FFB9FFEEFF7D9EC1FE89F2DB" bold="true" box="[189,267,1930,1956]" pageId="7" pageNumber="8">Fig. 3.</emphasis>
|
||
<taxonomicName id="8EEF79B6FFB9FFEEFED39EC1FD81F2DB" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[275,515,1930,1956]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="hadroglossa" status="sp. nov.">
|
||
<emphasis id="7B9BDE27FFB9FFEEFED39EC1FD81F2DB" box="[275,515,1930,1956]" italics="true" pageId="7" pageNumber="8">Gulella hadroglossa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB9FFEEFDCB9EC0FDE5F2DA" box="[523,615,1931,1957]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="7B9BDE27FFB9FFEEFDAE9EC1FD27F2DB" bold="true" box="[622,677,1930,1956]" pageId="7" pageNumber="8">A–E</emphasis>
|
||
. Holotype, height 3.0 mm, diameter 1.5 mm (NMSA P0230/ T4084).
|
||
<emphasis id="7B9BDE27FFB9FFEEFEE89EE6FEDDF2B8" bold="true" box="[296,351,1965,1991]" pageId="7" pageNumber="8">F–G</emphasis>
|
||
. Juvenile specimen showing apertural dentition, diameter 1.5 mm (paratype, NMSA P0417/T4085).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="01F551BEFFB6FFE1FF7D9840FC8BF1E4" pageId="8" pageNumber="9" type="description">
|
||
<paragraph id="49500235FFB6FFE1FF7D9840FECCF45A" blockId="8.[189,334,267,293]" box="[189,334,267,293]" pageId="8" pageNumber="9">
|
||
<heading id="1218B559FFB6FFE1FF7D9840FECCF45A" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="7B9BDE27FFB6FFE1FF7D9840FECCF45A" bold="true" box="[189,334,267,293]" pageId="8" pageNumber="9">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFB6FFE1FF7D9871FD7BF125" blockId="8.[189,1399,313,1114]" pageId="8" pageNumber="9">
|
||
SHELL (
|
||
<figureCitation id="D1D41EB0FFB6FFE1FED69872FEDDF42B" box="[278,351,313,340]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[193,1394,777,1884]" captionTargetId="figure@7.[189,1399,765,1889]" captionTargetPageId="7" captionText="Fig. 3. Gulella hadroglossa sp. nov. A–E. Holotype, height 3.0 mm, diameter 1.5 mm (NMSA P0230/ T4084). F–G. Juvenile specimen showing apertural dentition, diameter 1.5 mm (paratype, NMSA P0417/T4085)." figureDoi="http://doi.org/10.5281/zenodo.3847852" httpUri="https://zenodo.org/record/3847852/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
). Small, pupiform; adult height 3.0–
|
||
<quantity id="8E17AFD0FFB6FFE1FCDE9871FCFFF42B" box="[798,893,314,340]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="8" pageNumber="9" unit="mm" value="3.1">3.1 mm</quantity>
|
||
, diameter
|
||
<quantity id="8E17AFD0FFB6FFE1FC3C9872FB2FF42B" box="[1020,1197,313,340]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.55" metricValueMin="1.45" pageId="8" pageNumber="9" unit="mm" value="1.5" valueMax="1.55" valueMin="1.45">1.45–1.55 mm</quantity>
|
||
; H:D 1.96–2.09, with up to 7.25 whorls. Protoconch diameter ~
|
||
<quantity id="8E17AFD0FFB6FFE1FD2F9816FCD2F408" box="[751,848,349,375]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="9" unit="mm" value="0.7">0.7 mm</quantity>
|
||
, comprising 1.5–1.75 whorls beyond nucleus, smooth and glossy when fresh; junction with teleoconch indistinct. Teleoconch comprising 5.0–5.5 whorls; whorls weakly convex, lacking a shoulder and suture weakly indented (may appear somewhat channelled in eroded specimens); smooth and glossy, sculptured only by occasional weak growthlines, some specimens with faint traces of subsutural pliculae (not evident in
|
||
<typeStatus id="9654BC97FFB6FFE1FBA098A2FB4EF77B" box="[1120,1228,489,516]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
); growth-lines stronger and more close-set on apertural tube behind peristome. Aperture ovate, but truncated where peristome interrupted in parietal region; aperture rim flaring and strongly recurved, forming a nearly closed channel behind lip; apertural dentition 3-fold (
|
||
<figureCitation id="D1D41EB0FFB6FFE1FCF69B18FCF9F712" box="[822,891,595,621]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[193,1394,777,1884]" captionTargetId="figure@7.[189,1399,765,1889]" captionTargetPageId="7" captionText="Fig. 3. Gulella hadroglossa sp. nov. A–E. Holotype, height 3.0 mm, diameter 1.5 mm (NMSA P0230/ T4084). F–G. Juvenile specimen showing apertural dentition, diameter 1.5 mm (paratype, NMSA P0417/T4085)." figureDoi="http://doi.org/10.5281/zenodo.3847852" httpUri="https://zenodo.org/record/3847852/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
C–D): 1) a very large parietal lamella with a tongue-like anterior element that curves outward and to the right, well beyond plane of aperture and then backward towards outer lip; interval between lamella and outer lip partially filled with an overarching shelf, leaving a U-shaped notch behind lamella; 2) a mid-labral plate running inward from a low thickening of outer lip, its upper margin raised to form a narrow ridge that runs obliquely inward and downward (hidden by parietal lamella in frontal view), basal margin usually ill-defined; 3) a narrow in-running tooth near base of columella; a columella lamella is not evident, though in some specimens a very deep-set vertical ledge may be discerned where apertural tube kinks around to join rounded portion of last whorl. A juvenile specimen of ~2.25 teleoconch whorls possessed 4-fold apertural dentition (
|
||
<figureCitation id="D1D41EB0FFB6FFE1FF069ADBFE88F6D5" box="[198,266,912,938]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[193,1394,777,1884]" captionTargetId="figure@7.[189,1399,765,1889]" captionTargetPageId="7" captionText="Fig. 3. Gulella hadroglossa sp. nov. A–E. Holotype, height 3.0 mm, diameter 1.5 mm (NMSA P0230/ T4084). F–G. Juvenile specimen showing apertural dentition, diameter 1.5 mm (paratype, NMSA P0417/T4085)." figureDoi="http://doi.org/10.5281/zenodo.3847852" httpUri="https://zenodo.org/record/3847852/files/figure.png" pageId="8" pageNumber="9">Fig. 3</figureCitation>
|
||
F–G) comprising: two well-developed, flattened, semi-circular teeth set transversely at left and right of basal lip, that on the right more deep-set; a curved parietal tooth just beyond limit of aperture, behind and to the right of which arises a fourth, more rounded tooth; a second, identical set of teeth visible by transparency one half-whorl behind aperture. Umbilicus closed, base grooved beneath basal columella tooth (
|
||
<figureCitation id="D1D41EB0FFB6FFE1FE499D57FE64F148" box="[393,486,1052,1079]" captionStart="Fig" captionStartId="7.[189,232,1930,1956]" captionTargetBox="[193,1394,777,1884]" captionTargetId="figure@7.[189,1399,765,1889]" captionTargetPageId="7" captionText="Fig. 3. Gulella hadroglossa sp. nov. A–E. Holotype, height 3.0 mm, diameter 1.5 mm (NMSA P0230/ T4084). F–G. Juvenile specimen showing apertural dentition, diameter 1.5 mm (paratype, NMSA P0417/T4085)." figureDoi="http://doi.org/10.5281/zenodo.3847852" httpUri="https://zenodo.org/record/3847852/files/figure.png" pageId="8" pageNumber="9">Fig. 3E</figureCitation>
|
||
). Empty shells translucent greyish-white when fresh; yellow colour of dried body showing through in live-collected material.
|
||
</paragraph>
|
||
<paragraph id="49500235FFB6FFE1FF7D9DC9FC8BF1E4" blockId="8.[189,777,1153,1180]" box="[189,777,1153,1180]" pageId="8" pageNumber="9">LIVING ANIMAL. No preserved specimens available.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="01F551BEFFB6FFE1FF7D9D89FC5CF0E1" pageId="8" pageNumber="9" type="distribution">
|
||
<paragraph id="49500235FFB6FFE1FF7D9D89FED4F1A3" blockId="8.[189,342,1218,1244]" box="[189,342,1218,1244]" pageId="8" pageNumber="9">
|
||
<heading id="1218B559FFB6FFE1FF7D9D89FED4F1A3" bold="true" box="[189,342,1218,1244]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="7B9BDE27FFB6FFE1FF7D9D89FED4F1A3" bold="true" box="[189,342,1218,1244]" pageId="8" pageNumber="9">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFB6FFE1FF7D9DBAFAC8F050" blockId="8.[189,1399,1265,1327]" pageId="8" pageNumber="9">
|
||
A narrow-range endemic, currently recorded only from Mariepskop Forest Reserve on the edge of the Drakensberg Escarpment in northern
|
||
<collectingRegion id="8B2BCCD7FFB6FFE1FDB49C5FFC90F051" box="[628,786,1300,1326]" country="South Africa" name="Mpumalanga" pageId="8" pageNumber="9">Mpumalanga</collectingRegion>
|
||
,
|
||
<collectingCountry id="31F842A5FFB6FFE1FCDE9C5FFC35F051" box="[798,951,1300,1326]" name="South Africa" pageId="8" pageNumber="9">South Africa</collectingCountry>
|
||
; at altitudes from
|
||
<quantity id="8E17AFD0FFB6FFE1FB4A9C5FFAC4F050" box="[1162,1350,1300,1327]" metricMagnitude="3" metricUnit="m" metricValue="1.5025" metricValueMax="1.545" metricValueMin="1.46" pageId="8" pageNumber="9" unit="m" value="1502.5" valueMax="1545.0" valueMin="1460.0">
|
||
<date id="3D5124F5FFB6FFE1FB4A9C5FFAA7F050" box="[1162,1317,1300,1327]" pageId="8" pageNumber="9" value="1460" valueMax="1545">1460 to 1545</date>
|
||
m
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="49500235FFB6FFE1FF7D9C1DFE9FF00F" blockId="8.[189,285,1366,1392]" box="[189,285,1366,1392]" pageId="8" pageNumber="9">
|
||
<heading id="1218B559FFB6FFE1FF7D9C1DFE9FF00F" bold="true" box="[189,285,1366,1392]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="7B9BDE27FFB6FFE1FF7D9C1DFE9FF00F" bold="true" box="[189,285,1366,1392]" pageId="8" pageNumber="9">Habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFB6FFE1FF7D9CCFFC5CF0E1" blockId="8.[189,990,1412,1439]" box="[189,990,1412,1439]" pageId="8" pageNumber="9">
|
||
Northern mist-belt forest (
|
||
<bibRefCitation id="2D7E7FC4FFB6FFE1FE309CCFFCBAF0E0" author="Mucina L. & Rutherford M. C." box="[496,824,1412,1439]" pageId="8" pageNumber="9" refId="ref14287" refString="Mucina L. & Rutherford M. C. (eds) 2006. The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia 19, South African National Biodiversity Institute, Pretoria." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
), in leaf-litter.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="01F551BEFFB6FFE1FF7D9C8DFC65F3A2" pageId="8" pageNumber="9" type="discussion">
|
||
<paragraph id="49500235FFB6FFE1FF7D9C8DFEADF09F" blockId="8.[189,303,1478,1504]" box="[189,303,1478,1504]" pageId="8" pageNumber="9">
|
||
<heading id="1218B559FFB6FFE1FF7D9C8DFEADF09F" bold="true" box="[189,303,1478,1504]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
|
||
<emphasis id="7B9BDE27FFB6FFE1FF7D9C8DFEADF09F" bold="true" box="[189,303,1478,1504]" pageId="8" pageNumber="9">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="49500235FFB6FFE1FF7D9CBFFBBCF3E3" blockId="8.[189,1399,1524,1692]" pageId="8" pageNumber="9">
|
||
<taxonomicName id="8EEF79B6FFB6FFE1FF7D9CBFFE2FF371" authority="Herbert, 2016" authorityName="Herbert" authorityYear="2016" box="[189,429,1524,1550]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="hadroglossa" status="sp. nov.">
|
||
<emphasis id="7B9BDE27FFB6FFE1FF7D9CBFFE2FF371" box="[189,429,1524,1550]" italics="true" pageId="8" pageNumber="9">Gulella hadroglossa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="60A8635CFFB6FFE1FE759CBEFD93F370" box="[437,529,1525,1551]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is rendered distinctive amongst small, smooth-shelled
|
||
<taxonomicName id="8EEF79B6FFB6FFE1FB5C9CBFFB76F371" authorityName="Pfeiffer" authorityYear="1856" box="[1180,1268,1524,1550]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
|
||
<emphasis id="7B9BDE27FFB6FFE1FB5C9CBFFB76F371" box="[1180,1268,1524,1550]" italics="true" pageId="8" pageNumber="9">Gulella</emphasis>
|
||
</taxonomicName>
|
||
species on account of its large recurved parietal lamella. It is comparable in size (slightly smaller) and superficially similar to
|
||
<taxonomicName id="8EEF79B6FFB6FFE1FEF59F70FD09F32A" authority="Bruggen, 1966" authorityName="Bruggen" authorityYear="1966" box="[309,651,1595,1621]" class="Gastropoda" family="Streptaxidae" genus="Gulella" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="verdcourti">
|
||
<emphasis id="7B9BDE27FFB6FFE1FEF59F70FE50F32A" box="[309,466,1595,1621]" italics="true" pageId="8" pageNumber="9">G. verdcourti</emphasis>
|
||
Bruggen, 1966
|
||
</taxonomicName>
|
||
, from the neighbouring Wolkberg massif, but that species has a distinct subsutural shoulder resulting in an incised suture, and its apertural dentition is also different, the parietal lamella not projecting as far and the mid-labral tooth being stronger.
|
||
</paragraph>
|
||
<paragraph id="49500235FFB6FFE1FD8D9F88FC65F3A2" blockId="8.[589,999,1731,1757]" box="[589,999,1731,1757]" pageId="8" pageNumber="9">
|
||
Family
|
||
<taxonomicName id="8EEF79B6FFB6FFE1FD689F88FC65F3A2" authority="Simroth, 1889" authorityName="Simroth" authorityYear="1889" box="[680,999,1731,1757]" class="Gastropoda" family="Urocyclidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="family">Urocyclidae Simroth, 1889</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |