174 lines
21 KiB
XML
174 lines
21 KiB
XML
<document id="F20AC70E326D7B03CB50E9AE05E7352F" ID-CLB-Dataset="26049" ID-DOI="10.11646/zootaxa.4654.1.1" ID-GBIF-Dataset="9ce1d103-9234-4fbe-9544-f1c294bb8783" ID-ISSN="1175-5326" ID-Zenodo-Dep="3365803" ID-ZooBank="CFD82CC0-3110-472E-972B-7ADC0C523A04" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565592061838" checkinUser="plazi" docAuthor="Hansen, Thomas" docDate="2019" docId="BF18F633A9A0FF452B9CC1D4FD94F9E2" docLanguage="en" docName="zootaxa.4654.1.1.pdf" docOrigin="Zootaxa 4654 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Ravniella danica" docType="treatment" docVersion="23" lastPageNumber="161" masterDocId="43218E4BA900FFE52B0BC14DFFA9FFE3" masterDocTitle="Gastropods from the Cretaceous-Palaeogene boundary in Denmark" masterLastPageNumber="196" masterPageNumber="1" pageNumber="161" updateTime="1698750564626" updateUser="ExternalLinkService">
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<mods:title id="6D7FFE50AB63896AE35A8E18C5E8BC91">Gastropods from the Cretaceous-Palaeogene boundary in Denmark</mods:title>
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<mods:namePart id="2CB85E440C39174B5F20BC1734199C12">Hansen, Thomas</mods:namePart>
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<mods:title id="66183B751F97C07F7F2EE7C903C31C65">Zootaxa</mods:title>
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<mods:part id="730E3AFBBE306F46FE8A6DCB3D50E3AB">
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<mods:date id="88167BD72749DED6583F86509BF6C4D0">2019</mods:date>
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<mods:number id="91F35780924D478BB4EF73570DEE41BB">2019-08-12</mods:number>
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<mods:number id="B3FFF815A5000D3CCF86A25575676C11">4654</mods:number>
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<mods:identifier id="7D2E1773FC5F5B936904D37EB1298030" type="DOI">10.11646/zootaxa.4654.1.1</mods:identifier>
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<treatment id="BF18F633A9A0FF452B9CC1D4FD94F9E2" ID-DOI="http://doi.org/10.5281/zenodo.5582985" ID-GBIF-Taxon="159074653" ID-Zenodo-Dep="5582985" LSID="urn:lsid:plazi:treatment:BF18F633A9A0FF452B9CC1D4FD94F9E2" httpUri="http://treatment.plazi.org/id/BF18F633A9A0FF452B9CC1D4FD94F9E2" lastPageNumber="161" pageId="160" pageNumber="161">
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<subSubSection id="7FAB14AEA9A0FF452B9CC1D4FDA2FF50" box="[151,523,153,180]" pageId="160" pageNumber="161" type="nomenclature">
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<paragraph id="370E4725A9A0FF452B9CC1D4FDA2FF50" blockId="160.[151,523,153,215]" box="[151,523,153,180]" pageId="160" pageNumber="161">
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<heading id="6C46F049A9A0FF452B9CC1D4FDA2FF50" bold="true" box="[151,523,153,180]" fontSize="11" level="1" pageId="160" pageNumber="161" reason="1">
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<taxonomicName id="F0B13CA6A9A0FF452B9CC1D4FDA2FF50" authority="(Ravn, 1902)" baseAuthorityName="Ravn" baseAuthorityYear="1902" box="[151,523,153,180]" class="Gastropoda" family="Acteonidae" genus="Ravniella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="160" pageNumber="161" phylum="Mollusca" rank="species" species="danica">
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<emphasis id="05C59B37A9A0FF452B9CC1D4FDA2FF50" bold="true" box="[151,523,153,180]" pageId="160" pageNumber="161">
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<emphasis id="05C59B37A9A0FF452B9CC1D4FECAFF57" bold="true" box="[151,355,153,180]" italics="true" pageId="160" pageNumber="161">Ravniella danica</emphasis>
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(
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<bibRefCitation id="53203AD4A9A0FF452A7EC1D7FDABFF50" author="Ravn, J. P. J." box="[373,514,153,180]" pageId="160" pageNumber="161" pagination="205 - 270" refId="ref138257" refString="Ravn, J. P. J. (1902) Molluskerne i Danmarks Kridtaflejringer. II, Scaphopoder, Gastropoder og Cephalopoder. Det Kongelige Danske Videnskabernes Selskab Skrifter, 6. Raekke, Naturvidenskabelig og Mathematisk Afdeling XI, 4, 205 - 270, pls. 1 - 5." type="journal article" year="1902">Ravn, 1902</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7FAB14AEA9A0FF452B9CC1F0FC67FEDB" pageId="160" pageNumber="161" type="description">
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<paragraph id="370E4725A9A0FF452B9CC1F0FEBEFF34" blockId="160.[151,523,153,215]" box="[151,279,189,215]" pageId="160" pageNumber="161">
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<figureCitation id="AF8A5BA0A9A0FF452B9CC1F0FF40FF34" box="[151,233,189,215]" captionStart="FIGURE 34" captionStartId="159.[151,250,188,213]" captionTargetId="figure@158.[151,1435,179,1918]" captionTargetPageId="158" captionText="FIGURE 34. A–E. Admetula sp. A–D. Silicone cast and internal mould of MGUH 33297. Widths 6.6, 7.2, 7.0 and 6.1 mm respectively. E. Internal mould of MGUH 33298. Width 5.6 mm. F. Kroisbachia sp., Silicone cast of MGUH 33299. Width 2.4 mm. G. Mataxa s.l. sp., Silicone cast of MGUH 33300. Width 3.9 mm. H–I. Neogastropoda genus et sp. indet., Silicone cast of MGUH 33301. Width 0.7 mm. J–N. Ravniella danica (Ravn, 1902). J–K. Silicone cast of MGUH 33202. Widths 4.7 and 4.3 respectively. L. Silicone cast of lectotype MGUH 126. Width 4.3 mm. Photograph by S.L. Jakobsen. M–N. Silicone cast of MGUH 33303. Width 4.1 and 3.8 mm. O–S. Disculus selandicum (Ravn, 1902). O. Silicone cast of MGUH 33304. Width 8.7 mm. P–Q. Silicone cast of paralectotype MGUH 80 figured by J.P.J. Ravn (1902, Pl. I: 6). Width 7.1, 6.9 mm respectively. R. Silicone cast of lectotype MGUH 79. Width 6.0 mm. Photograph by S.L. Jakobsen. S. Silicone cast of MGUH 33305. Width 5.9 mm." figureDoi="http://doi.org/10.5281/zenodo.3442124" httpUri="https://zenodo.org/record/3442124/files/figure.png" pageId="160" pageNumber="161">Figs 34</figureCitation>
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J–N
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</paragraph>
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<paragraph id="370E4725A9A0FF452B9CC04FFD7CFEFA" blockId="160.[151,974,257,313]" box="[151,725,257,282]" pageId="160" pageNumber="161">
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1902
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<taxonomicName id="F0B13CA6A9A0FF452BDBC04FFEC1FEF9" authority="Hansen, 2019" authorityName="Hansen" authorityYear="2019" box="[208,360,258,282]" class="Gastropoda" family="Ringiculidae" genus="Cinulia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="160" pageNumber="161" phylum="Mollusca" rank="species" species="danica" status="sp. nov.">
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<emphasis id="05C59B37A9A0FF452BDBC04FFEC1FEF9" box="[208,360,258,282]" italics="true" pageId="160" pageNumber="161">Cinulia danica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="1EF6264CA9A0FF452A65C04FFE0CFEFA" box="[366,421,258,281]" pageId="160" pageNumber="161" rank="species">n. sp.</taxonomicNameLabel>
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Ravn: 238–239, pl. III, fig. 7.
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</paragraph>
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<paragraph id="370E4725A9A0FF452B9CC06DFC67FEDB" blockId="160.[151,974,257,313]" box="[151,974,288,313]" pageId="160" pageNumber="161">
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1970
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<taxonomicName id="F0B13CA6A9A0FF452BDBC06DFDF6FEDA" authority="(Ravn)" baseAuthorityName="Ravn" box="[208,607,288,313]" class="Gastropoda" family="Acteonidae" genus="Tornatellaea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="160" pageNumber="161" phylum="Mollusca" rank="species" species="danica" subGenus="Ravniella">
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<emphasis id="05C59B37A9A0FF452BDBC06DFEF8FEDB" box="[208,337,288,312]" italics="true" pageId="160" pageNumber="161">Tornatellaea</emphasis>
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(
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<emphasis id="05C59B37A9A0FF452A6BC06DFE68FEDB" box="[352,449,288,312]" italics="true" pageId="160" pageNumber="161">Ravniella</emphasis>
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)
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<emphasis id="05C59B37A9A0FF452AC4C06DFDBDFEDB" box="[463,532,288,312]" italics="true" pageId="160" pageNumber="161">danica</emphasis>
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(Ravn)
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</taxonomicName>
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—Rosenkrantz: 431–433, fig. 10: 1.
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</paragraph>
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</subSubSection>
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<subSubSection id="7FAB14AEA9A0FF452B9CC02BFD94F9E2" pageId="160" pageNumber="161" type="materials_examined">
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<paragraph id="370E4725A9A0FF452B9CC02BFD60FE2A" blockId="160.[151,1437,358,1537]" pageId="160" pageNumber="161">
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<emphasis id="05C59B37A9A0FF452B9CC02BFEBBFE62" bold="true" box="[151,274,358,385]" pageId="160" pageNumber="161">Diagnosis.</emphasis>
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Protoconch heterostrophic, of 1½ smooth whorls. Teleoconch low-spired with strongly developed columellar folds and around eight teeth on outer lip. Whorl sculpture of widely spaced, finely pitted spiral grooves, generally numbering three to five on spire whorls.
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</paragraph>
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<paragraph id="370E4725A9A0FF452BCCC09EFB68FE0E" blockId="160.[151,1437,358,1537]" box="[199,1217,466,493]" pageId="160" pageNumber="161">
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<materialsCitation id="87D94D78A9A0FF452BCCC09EFB14FE0E" ID-GBIF-Occurrence="2331574344" box="[199,1213,466,493]" country="Denmark" location="Stevns Klint" pageId="160" pageNumber="161" specimenCount="2" typeStatus="holotype">
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<emphasis id="05C59B37A9A0FF452BCCC09EFD93FE0E" bold="true" box="[199,570,466,493]" pageId="160" pageNumber="161">
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||
<typeStatus id="E80AF987A9A0FF452BCCC09EFEABFE0E" box="[199,258,467,493]" pageId="160" pageNumber="161">Type</typeStatus>
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stratum and
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<typeStatus id="E80AF987A9A0FF452AA8C09EFE7FFE0E" box="[419,470,467,493]" pageId="160" pageNumber="161">type</typeStatus>
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locality.
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</emphasis>
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<taxonomicName id="F0B13CA6A9A0FF45294AC09EFC3CFE0E" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[577,917,467,493]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="160" pageNumber="161" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
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at
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<location id="326E11FEA9A0FF4528B3C09EFBECFE0E" LSID="urn:lsid:plazi:treatment:BF18F633A9A0FF452B9CC1D4FD94F9E2:326E11FEA9A0FF4528B3C09EFBECFE0E" box="[952,1093,467,493]" country="Denmark" name="Stevns Klint" pageId="160" pageNumber="161">Stevns Klint</location>
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,
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<collectingCountry id="4FA607B5A9A0FF452F5AC09EFB14FE0E" box="[1105,1213,467,493]" name="Denmark" pageId="160" pageNumber="161">Denmark</collectingCountry>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="370E4725A9A0FF452BCCC0BBFE73FC9A" blockId="160.[151,1437,358,1537]" pageId="160" pageNumber="161">
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<emphasis id="05C59B37A9A0FF452BCCC0BBFE9FFDF2" bold="true" box="[199,310,502,529]" pageId="160" pageNumber="161">Material.</emphasis>
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Of the original
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<specimenCount id="21B78CACA9A0FF452AEEC0BAFD9EFDF2" box="[485,567,503,529]" pageId="160" pageNumber="161" type="generic">12 type</specimenCount>
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specimens
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<bibRefCitation id="53203AD4A9A0FF4529B3C0BAFCEDFDF2" author="Ravn, J. P. J." box="[696,836,503,529]" pageId="160" pageNumber="161" pagination="205 - 270" refId="ref138257" refString="Ravn, J. P. J. (1902) Molluskerne i Danmarks Kridtaflejringer. II, Scaphopoder, Gastropoder og Cephalopoder. Det Kongelige Danske Videnskabernes Selskab Skrifter, 6. Raekke, Naturvidenskabelig og Mathematisk Afdeling XI, 4, 205 - 270, pls. 1 - 5." type="journal article" year="1902">Ravn (1902)</bibRefCitation>
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used for his description, only the one figured,
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<collectionCode id="51A0DFE0A9A0FF452E4CC0BAFA34FDF2" box="[1351,1437,503,529]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="160" pageNumber="161">MGUH</collectionCode>
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126, is known today. The others were not given a number and are no longer possible to separate from the rest of the specimens found in the old collections. For this reason
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<collectionCode id="51A0DFE0A9A0FF452849C372FC31FDBA" box="[834,920,575,601]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="160" pageNumber="161">MGUH</collectionCode>
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126 is here selected as
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<typeStatus id="E80AF987A9A0FF452FA4C372FAB0FDBA" box="[1199,1305,575,601]" pageId="160" pageNumber="161" type="lectotype">lectotype</typeStatus>
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||
. Additional material consists of
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<collectionCode id="51A0DFE0A9A0FF452A88C32EFE70FD9E" box="[387,473,611,637]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="160" pageNumber="161">MGUH</collectionCode>
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||
33302,
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<collectionCode id="51A0DFE0A9A0FF452930C32EFD38FD9E" box="[571,657,611,637]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="160" pageNumber="161">MGUH</collectionCode>
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||
33303,
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||
<collectionCode id="51A0DFE0A9A0FF4529F8C32FFC9DFD9E" box="[755,820,610,637]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="160" pageNumber="161">ØSM</collectionCode>
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||
.10042-88-a–b,
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<collectionCode id="51A0DFE0A9A0FF4528EDC32FFB8EFD9E" box="[998,1063,610,637]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="160" pageNumber="161">ØSM</collectionCode>
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.10042-89-a–b,
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||
<collectionCode id="51A0DFE0A9A0FF452FD3C32FFAB0FD9E" box="[1240,1305,610,637]" country="USA" httpUri="http://grbio.org/cool/2e2p-880n" name="Ohio State University Museum" pageId="160" pageNumber="161">ØSM</collectionCode>
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||
.10042-106, more than
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<specimenCount id="21B78CACA9A0FF452A1FC3CAFE15FD42" box="[276,444,647,673]" pageId="160" pageNumber="161" type="generic">130 specimens</specimenCount>
|
||
without numbers from the old collections at the Geological Museum, Natural History Museum of Copenhagen, and 43 new specimens with the informal sample numbers SH.106.B, SH.142, SH.148, SH.161, SH.186, SH.197.A, SH.205, SH.206, SH.285.A–B, SH.285.A, SH.289 (
|
||
<specimenCount id="21B78CACA9A0FF452F3EC382FB6FFD09" box="[1077,1222,719,746]" pageId="160" pageNumber="161" type="generic">2 specimens</specimenCount>
|
||
), SH.290, SH.303. A (
|
||
<specimenCount id="21B78CACA9A0FF452BB1C3BEFEE3FCED" box="[186,330,755,782]" pageId="160" pageNumber="161" type="generic">2 specimens</specimenCount>
|
||
), SH.324, SH.325.B–C, SH.325.C, SH.334.A–B, SH.334.C, SH.352.A–B (
|
||
<specimenCount id="21B78CACA9A0FF452F96C3BEFA85FCED" box="[1181,1324,755,782]" pageId="160" pageNumber="161" type="generic">6 specimens</specimenCount>
|
||
), SH.384. A, SH.410.A–B, SH.414.B–C, SH.416, SH.418.A–B, SR.168, SR.225, SR.278, SR.283, SR.286, SR.307.A–B, SR.400, SR.516 (
|
||
<specimenCount id="21B78CACA9A0FF452A6BC276FE5BFCB5" box="[352,498,827,854]" pageId="160" pageNumber="161" type="generic">2 specimens</specimenCount>
|
||
), SR.570.A–C, SR.642.B, SO.110.A–B, DN.26.A–B, DN.30.A–B, DN.33.A–B (
|
||
<specimenCount id="21B78CACA9A0FF452E86C276FEA4FC99" pageId="160" pageNumber="161" type="generic">2 specimens</specimenCount>
|
||
) and DN.52.A–B.
|
||
</paragraph>
|
||
<paragraph id="370E4725A9A0FF452BCCC2CFFE69FC22" blockId="160.[151,1437,358,1537]" pageId="160" pageNumber="161">
|
||
<emphasis id="05C59B37A9A0FF452BCCC2CFFEF3FC7E" bold="true" box="[199,346,898,925]" pageId="160" pageNumber="161">Occurrence.</emphasis>
|
||
Common in the
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||
<taxonomicName id="F0B13CA6A9A0FF45291EC2CEFCC0FC7E" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[533,873,899,925]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="160" pageNumber="161" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
|
||
at Stevns Klint and in the contemporaneous ‘dead layer’ in Northern Jutland.
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||
</paragraph>
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||
<paragraph id="370E4725A9A0FF452BCCC287FF5EFAAE" blockId="160.[151,1437,358,1537]" pageId="160" pageNumber="161">
|
||
<emphasis id="05C59B37A9A0FF452BCCC287FEF0FC06" bold="true" box="[199,345,970,997]" pageId="160" pageNumber="161">Description.</emphasis>
|
||
Shell subovate, small, generally below
|
||
<quantity id="F049EAC0A9A0FF45282BC286FCD9FC06" box="[800,880,971,997]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="160" pageNumber="161" unit="mm" value="10.0">10 mm</quantity>
|
||
high, width/height ratio varying between 57 and 70 %, generally decreasing with size. Protoconch heterostrophic with nearly 1.5 loosely coiled, smooth whorls. Transition to teleoconch sharp, marked by appearance of spiral grooves. Protoconch between 0.6 and
|
||
<quantity id="F049EAC0A9A0FF452E08C55EFAF0FBCE" box="[1283,1369,1043,1069]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="160" pageNumber="161" unit="mm" value="0.8">0.8 mm</quantity>
|
||
wide. Teleoconch with rather weakly convex whorls and distinct suture. Spire sides nearly straight to at times slightly concave due to an increase in whorl expansion on later whorls. Last whorl corresponding to between 75 and 85 % of shell height. Aperture narrowly oval-rhombic, width approximating 2/3 the height. Inner lip with two widely spaced, oblique strong columellar folds and a weaker parietal fold slightly above columella. Outer lip moderately sharp, slightly thickened, carrying around eight sharp teeth, although adapical and abapical ones often appears effaced or missing. Whorl sculpture of widely spaced, finely pitted spiral grooves, generally numbering three to five on spire whorls, adapical groove markedly stronger than succeeding ones. Last whorl typically with around 15 spiral grooves.
|
||
</paragraph>
|
||
<paragraph id="370E4725A9A0FF452BCCC41AFF42FA76" blockId="160.[151,1437,358,1537]" pageId="160" pageNumber="161">
|
||
<emphasis id="05C59B37A9A0FF452BCCC41AFED7FA92" bold="true" box="[199,382,1367,1393]" pageId="160" pageNumber="161">Measurements.</emphasis>
|
||
<typeStatus id="E80AF987A9A0FF452A8CC41AFE5EFA92" box="[391,503,1367,1393]" pageId="160" pageNumber="161" type="lectotype">Lectotype</typeStatus>
|
||
<collectionCode id="51A0DFE0A9A0FF45290AC41AFDFEFA92" box="[513,599,1367,1393]" country="Denmark" httpUri="http://grbio.org/cool/c1ae-v64p" name="Museum Geologicum Universitatis Hafniensis" pageId="160" pageNumber="161">MGUH</collectionCode>
|
||
126 is approximately
|
||
<quantity id="F049EAC0A9A0FF452850C41BFC1DFA92" box="[859,948,1366,1393]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="160" pageNumber="161" unit="mm" value="6.5">6.5 mm</quantity>
|
||
high and
|
||
<quantity id="F049EAC0A9A0FF452F21C41AFB2AFA92" box="[1066,1155,1367,1393]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="160" pageNumber="161" unit="mm" value="4.3">4.3 mm</quantity>
|
||
wide, consisting of 5 ½ whorls.
|
||
</paragraph>
|
||
<paragraph id="370E4725A9A0FF452BCCC4D2FD94F9E2" blockId="160.[151,1437,358,1537]" pageId="160" pageNumber="161">
|
||
<emphasis id="05C59B37A9A0FF452BCCC4D2FE92FA5A" bold="true" box="[199,315,1439,1465]" pageId="160" pageNumber="161">Remarks.</emphasis>
|
||
This gastropod, which most likely was a vermivore (
|
||
<bibRefCitation id="53203AD4A9A0FF4528ADC4D2FB18FA5A" author="Jeffery, P. & Tracey, S." box="[934,1201,1439,1465]" pageId="160" pageNumber="161" pagination="75 - 137" refId="ref132882" refString="Jeffery, P. & Tracey, S. (1997) The Early Eocene London Clay Formation mollusc fauna of the former Bursledon Brickworks, Lower Swanwick, Hampshire. Tertiary Research, 17 (3 - 4), 75 - 137." type="journal article" year="1997">Jeffery & Tracey 1997</bibRefCitation>
|
||
), is one of the more dominant gastropod species in the
|
||
<taxonomicName id="F0B13CA6A9A0FF452917C48EFCDAFA3E" authority="Limestone Member" authorityName="Gilleaudeau et al." authorityYear="2018" box="[540,883,1475,1501]" class="Gastropoda" family="Cerithiidae" genus="Cerithium" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="160" pageNumber="161" phylum="Mollusca" rank="genus">Cerithium Limestone Member</taxonomicName>
|
||
, although it generally does not exceed 2 % of the total number of gastropod specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |