308 lines
35 KiB
XML
308 lines
35 KiB
XML
<document id="B04FDAD8AFA86FB8CC69D9A82247F50D" ID-CLB-Dataset="3320" ID-DOI="10.1093/zoolinnean/zlaa052" ID-GBIF-Dataset="fd4b2931-7bad-4644-b485-f365040ae01e" ID-ISSN="0024-4082" ID-Zenodo-Dep="5724192" ID-ZooBank="0810A91E-7D8D-4F56-B468-9D5B3A6115E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619628947432" checkinUser="carolina" docAuthor="Kamiński, Marcin J., Gearner, Olivia M., Kanda, Kojun, Swichtenberg, Kali, Purchart, Luboš & Smith, Aaron D." docDate="2021" docId="BA5587AF637A4A1C91A36D782E0CFDC4" docLanguage="en" docName="ZoolJLinnSoc.191.883-901.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Toktokkus makuya Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith 2021" docType="treatment" docVersion="6" lastPageNumber="898" masterDocId="466CFFD763744A13920F6C4F2B04FFCB" masterDocTitle="First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group" masterLastPageNumber="901" masterPageNumber="883" pageNumber="897" updateTime="1698937967960" updateUser="ExternalLinkService">
|
||
<mods:mods id="733CBD39D0052F014D8E3A356AE0681C" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="80658B9BDBFD025844750779029AEBA6">
|
||
<mods:title id="7F4EEA99C6E52C696C0DADE39A161912">First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="7D36441A9E0442E03D80F85C6CB14B46" type="personal">
|
||
<mods:role id="611A8C43D510422677218966B30E84DC">
|
||
<mods:roleTerm id="B36243C33662FB8045EBB3B8B404A6E6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="47E1645A51884D0BBEC929431E2D8F01">Kamiński, Marcin J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="DFDDFA853C0C337A100631BFA1821799" type="personal">
|
||
<mods:role id="092DCD9D59862D97B6915C1A75DBA1A2">
|
||
<mods:roleTerm id="56C59DEDA95599EA0E04E880982DC329">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8E212E76C997AF04C5947AC20DD3F279">Gearner, Olivia M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8C70866F78436F551F54FF5A6A5989BF" type="personal">
|
||
<mods:role id="757D94B6DC74818F62760F5429804B63">
|
||
<mods:roleTerm id="35A3A5C94B4126F1947589D52C74A388">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="7F57EAEA8CC5A10860508C13CE753F63">Kanda, Kojun</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="340DA793DF2138BB0E3F6AE29265375F" type="personal">
|
||
<mods:role id="978A785C16A8C0AC9193D3405FF9F536">
|
||
<mods:roleTerm id="C3E68CED4372DD4EFD9FC178A4853381">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="93CF9CB13613953625DE90C870A1F060">Swichtenberg, Kali</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="9FCCD451162CAC7CC8C2FF9DD91FF13C" type="personal">
|
||
<mods:role id="FA99F836FF0C164367C5A6DD770581C4">
|
||
<mods:roleTerm id="0FD57E923FAD57AFDABE0CCADF946CAA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="679A71C36CAC0FECBF902E725284C8BD">Purchart, Luboš</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="18C981F0EC0FF90622A614BFAA04196A" type="personal">
|
||
<mods:role id="6300CB4D04009DB01FEACC99CD1F6DBB">
|
||
<mods:roleTerm id="40A17C5DBECBB044C714E3AE82979D61">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C062B61827BB6799418A5DC787647F64">Smith, Aaron D.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="9D2DB5DF7DB78673132DF0D33950FA7D">text</mods:typeOfResource>
|
||
<mods:relatedItem id="DA6A978CF9B4BAAA52CA37F9FB741B3D" type="host">
|
||
<mods:titleInfo id="21F0C41AB053D157E210786BB77FF07F">
|
||
<mods:title id="40296FADE749D590003584C18416CACF">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="62395FF8F06D8F9D0DBE461585B89FC0">
|
||
<mods:date id="A926923F3D810295FB06743494E2BE57">2021</mods:date>
|
||
<mods:detail id="3C623E80E91576D6A678108E1171B609" type="pubDate">
|
||
<mods:number id="451D2382621657D962A249BF907D4DF7">2020-06-16</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="2350FDC340EDAC2FF78C87DBBF22AE60" type="volume">
|
||
<mods:number id="BC71899A8E4336D0B5FD83DBE9EF6A6E">191</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FB1F76F8ECC360EE9E9776CE1603ECE3" unit="page">
|
||
<mods:start id="765A439C2872E40131F89FAB7E02E64B">883</mods:start>
|
||
<mods:end id="BE0306B6A785CBB93A4198F49260BFA4">901</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F9BCD73719B5D00CC1B6397851C4C0B4">journal article</mods:classification>
|
||
<mods:identifier id="5E619C568C8A3797756205E9EB758D83" type="CLB-Dataset">3320</mods:identifier>
|
||
<mods:identifier id="7DB9C60ABC196AF0E5DC8B18230CDA9E" type="DOI">10.1093/zoolinnean/zlaa052</mods:identifier>
|
||
<mods:identifier id="9B16A177112921BF73A85309D9ECBC1C" type="GBIF-Dataset">fd4b2931-7bad-4644-b485-f365040ae01e</mods:identifier>
|
||
<mods:identifier id="FEF8BB983EC89A9C37CF89317826BF9A" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="BAFB0AA3DAAD9F25E4A5CC8CABE615A1" type="Zenodo-Dep">5724192</mods:identifier>
|
||
<mods:identifier id="6E9964E59D082DBA6DDC20661B7E6B1A" type="ZooBank">0810A91E-7D8D-4F56-B468-9D5B3A6115E9</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="BA5587AF637A4A1C91A36D782E0CFDC4" ID-DOI="http://doi.org/10.5281/zenodo.5724212" ID-GBIF-Taxon="190683657" ID-Zenodo-Dep="5724212" LSID="urn:lsid:plazi:treatment:BA5587AF637A4A1C91A36D782E0CFDC4" httpUri="http://treatment.plazi.org/id/BA5587AF637A4A1C91A36D782E0CFDC4" lastPageId="15" lastPageNumber="898" pageId="14" pageNumber="897">
|
||
<subSubSection id="7AE66532637A4A1D91A36D782F6BFE58" pageId="14" pageNumber="897" type="nomenclature">
|
||
<paragraph id="324336B9637A4A1D91A36D782E0AFE85" blockId="14.[940,1294,310,335]" box="[940,1294,310,335]" pageId="14" pageNumber="897">
|
||
<heading id="690B81D5637A4A1D91A36D782E0AFE85" box="[940,1294,310,335]" centered="true" fontSize="9" level="2" pageId="14" pageNumber="897" reason="2">
|
||
<taxonomicName id="F5FC4D3A637A4A1D91A36D782E0AFE85" authority="Gearner" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" box="[940,1294,310,335]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="901" phylum="Arthropoda" rank="genus" status="sp. nov.">
|
||
TOKTOKKUS
|
||
<emphasis id="0088EAAB637A4A1D96326D752F9DFE85" box="[1085,1177,314,334]" italics="true" pageId="14" pageNumber="897">MAKUYA</emphasis>
|
||
GEARNER
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="324336B9637A4A1D91266D112F6BFE58" blockId="14.[809,1426,350,1630]" pageId="14" pageNumber="897">
|
||
lsid
|
||
<uri id="466D3ABB637A4A1D91696D112F6BFE58" pageId="14" pageNumber="897">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="465A0C6C637A4A1D96AF6D112F6BFE58" pageId="14" pageNumber="897">7265FD25-E2AF- 49BD-947A-A4C6D80E0C97</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7AE66532637A4A1C91266DD22E0CFDC4" lastPageId="15" lastPageNumber="898" pageId="14" pageNumber="897" type="materials_examined">
|
||
<paragraph id="324336B9637A4A1D91266DD22E6DF996" blockId="14.[809,1426,350,1630]" pageId="14" pageNumber="897">
|
||
<emphasis id="0088EAAB637A4A1D91266DD228C9FE7A" box="[809,973,412,434]" italics="true" pageId="14" pageNumber="897">Type material:</emphasis>
|
||
<typeStatus id="ED47881B637A4A1D91D66DD32F3BFE79" box="[985,1087,412,434]" pageId="14" pageNumber="897" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<collectionCode id="54EDAE7C637A4A1D96416DD32FA4FE79" box="[1102,1184,412,434]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="897" type="Museum">USNM</collectionCode>
|
||
), male: ‘South Africa: Limpopo: Makuya / N.
|
||
<collectionCode id="54EDAE7C637A4A1D963E6DF42F43FE1B" box="[1073,1095,443,464]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="897">R</collectionCode>
|
||
., Mutale Falls Safari Camp, /
|
||
<quantity id="F5049B5C637A4A1D91336D962886FE25" box="[828,898,473,494]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="897" unit="m" value="300.0">300m</quantity>
|
||
,
|
||
<geoCoordinate id="57C8507E637A4A1D919C6D9628F7FE25" box="[915,1011,473,494]" degrees="22.42" direction="south" orientation="latitude" pageId="14" pageNumber="897" precision="555" value="-22.42">22.42°S</geoCoordinate>
|
||
<geoCoordinate id="57C8507E637A4A1D91F06D962F60FE24" box="[1023,1124,473,495]" degrees="31.05" direction="east" orientation="longitude" pageId="14" pageNumber="897" precision="555" value="31.05">31.05°E</geoCoordinate>
|
||
, /
|
||
<date id="46421079637A4A1D96886D962FF3FE24" box="[1159,1271,473,495]" pageId="14" pageNumber="897" value="2018-02-02">2.ii.2018</date>
|
||
. KK18_064. K. Kanda, /
|
||
<collectionCode id="54EDAE7C637A4A1D91CC6DB728DDFDC6" box="[963,985,504,525]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="897">R</collectionCode>
|
||
. A. Gomez, J. M. Pflug’.
|
||
<emphasis id="0088EAAB637A4A1D971F6DB72E8DFDC6" box="[1296,1417,504,525]" italics="true" pageId="14" pageNumber="897">
|
||
<typeStatus id="ED47881B637A4A1D971F6DB72E8DFDC6" box="[1296,1417,504,525]" pageId="14" pageNumber="897" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
: Four males and
|
||
<specimenCount id="24FAFD30637A4A1D91E46E592F89FDE7" box="[1003,1165,534,556]" pageId="14" pageNumber="897" type="female">three females</specimenCount>
|
||
(Kojun Kanda): same data as
|
||
<typeStatus id="ED47881B637A4A1D91876E7A28EAFD80" box="[904,1006,565,587]" pageId="14" pageNumber="897" type="holotype">holotype</typeStatus>
|
||
; male (
|
||
<collectionCode id="54EDAE7C637A4A1D964B6E7A2F94FD80" box="[1092,1168,565,587]" pageId="14" pageNumber="897">PERC</collectionCode>
|
||
): same data but with: ‘Tenebrionid Base / Aaron D Smith / Catalog # 22651’; female (
|
||
<collectionCode id="54EDAE7C637A4A1D918B6E3C28C9FD42" box="[900,973,627,649]" pageId="14" pageNumber="897">CASC</collectionCode>
|
||
): ‘
|
||
<collectingCountry id="4AEB7629637A4A1D91ED6E3C2F9FFD43" box="[994,1179,627,648]" name="Mozambique" pageId="14" pageNumber="897">MOZAMBIQUE</collectingCountry>
|
||
/ Lourenco Marques’, ‘
|
||
<date id="46421079637A4A1D91216EDE2883FD6C" box="[814,903,657,679]" pageId="14" pageNumber="897" value="1957-02">ii 1957</date>
|
||
<emphasis id="0088EAAB637A4A1D919B6EDE289FFD6C" bold="true" box="[916,923,657,679]" pageId="14" pageNumber="897">/</emphasis>
|
||
NLHKrauss’; male (
|
||
<collectionCode id="54EDAE7C637A4A1D96A86EDE2FF3FD6C" box="[1191,1271,657,679]" pageId="14" pageNumber="897">CASC</collectionCode>
|
||
): ‘
|
||
<geoCoordinate id="57C8507E637A4A1D971C6EDE2E57FD6D" box="[1299,1363,657,678]" degrees="28" direction="east" orientation="longitude" pageId="14" pageNumber="897" precision="55555" value="28.0">28°E</geoCoordinate>
|
||
<geoCoordinate id="57C8507E637A4A1D975D6EDE2E8EFD6D" box="[1362,1418,657,678]" degrees="18" direction="south" orientation="latitude" pageId="14" pageNumber="897" precision="55555" value="-18.0">18°S</geoCoordinate>
|
||
, Rhodesia / Hostes Nicolle Inst. / Wildlife Res. Jan. / 1974 M. B. Fenton’; male and
|
||
<specimenCount id="24FAFD30637A4A1D968A6E812E27FD2F" box="[1157,1315,718,740]" pageId="14" pageNumber="897" type="female">three females</specimenCount>
|
||
(
|
||
<collectionCode id="54EDAE7C637A4A1D973C6E802E40FD2F" box="[1331,1348,719,740]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="897">T</collectionCode>
|
||
. Keith Philips): ‘SO.
|
||
<collectionCode id="54EDAE7C637A4A1D91DD6EA22F38FCC9" box="[978,1084,749,770]" pageId="14" pageNumber="897">AFRICA</collectionCode>
|
||
: Northern Prov. just NW of Sukses, Madikela / Game Res.,
|
||
<date id="46421079637A4A1D968B6F432E2DFCEA" box="[1156,1321,780,801]" pageId="14" pageNumber="897" value="1999-03-12" valueMax="1999-03-14" valueMin="1999-03-12">12–14.III.1999</date>
|
||
/ Philips, Gerofsky, and Kryger /
|
||
<geoCoordinate id="57C8507E637A4A1D96216F652F80FC8B" box="[1070,1156,810,832]" degrees="24" direction="south" minutes="05" orientation="latitude" pageId="14" pageNumber="897" precision="925" value="-24.083334">S24°05’</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="57C8507E637A4A1D969F6F642FE6FC8B" box="[1168,1250,811,832]" degrees="28" direction="east" minutes="18" orientation="longitude" pageId="14" pageNumber="897" precision="925" value="28.3">E28°18</geoCoordinate>
|
||
’’; male (
|
||
<collectionCode id="54EDAE7C637A4A1D97316F642E83FC8A" box="[1342,1415,811,833]" pageId="14" pageNumber="897">CASC</collectionCode>
|
||
): ‘S.
|
||
<collectionCode id="54EDAE7C637A4A1D91466F0628AEFC95" box="[841,938,841,862]" pageId="14" pageNumber="897">AFRICA</collectionCode>
|
||
/ Bechuanaland / Tsuagara /
|
||
<date id="46421079637A4A1D96F56F052E87FC94" box="[1274,1411,841,863]" pageId="14" pageNumber="897" value="1965-01-07">Jan. 7, 1965</date>
|
||
/ John W. Neal’; male (
|
||
<collectionCode id="54EDAE7C637A4A1D96156F272F60FCB5" box="[1050,1124,872,894]" pageId="14" pageNumber="897">CASC</collectionCode>
|
||
): ‘Naawpoort; / Pbg 21-11- 21 / G.
|
||
<collectionCode id="54EDAE7C637A4A1D91756FC82888FC57" box="[890,908,903,924]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="897" type="Museum">V</collectionCode>
|
||
. Son’, ‘
|
||
<taxonomicName id="F5FC4D3A637A4A1D91D76FC82F87FC57" box="[984,1155,903,924]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="901" phylum="Arthropoda" rank="genus">PSAMMODES</taxonomicName>
|
||
/ vialis / Burchell / det. Dr. C. Koch’;
|
||
<specimenCount id="24FAFD30637A4A1D91B56FEA2F57FC70" box="[954,1107,933,955]" pageId="14" pageNumber="897" type="female">three females</specimenCount>
|
||
(LuboŠ Purchart): ‘
|
||
<collectionCode id="54EDAE7C637A4A1D97246FEA2E89FC71" box="[1323,1421,933,954]" pageId="14" pageNumber="897">AFRICA</collectionCode>
|
||
,
|
||
<collectingCountry id="4AEB7629637A4A1D91266F8B28E2FC12" box="[809,998,964,985]" name="Mozambique" pageId="14" pageNumber="897">MOZAMBIQUE</collectingCountry>
|
||
/
|
||
<geoCoordinate id="57C8507E637A4A1D91F06F8B2F91FC11" box="[1023,1173,964,986]" degrees="22" direction="south" minutes="04.963" orientation="latitude" pageId="14" pageNumber="897" precision="1" value="-22.082716">S 22°04.963’</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="57C8507E637A4A1D96AC6F8B2E3DFC11" box="[1187,1337,964,986]" degrees="33" direction="east" minutes="55.577" orientation="longitude" pageId="14" pageNumber="897" precision="1" value="33.926285">E 33°55.577’</geoCoordinate>
|
||
/ camp site /
|
||
<date id="46421079637A4A1D91616FAC2F0AFC33" box="[878,1038,994,1016]" pageId="14" pageNumber="897" value="2011-03-03" valueMax="2011-03-04" valueMin="2011-03-03">3–4. iii. 2011</date>
|
||
’, ‘
|
||
<collectionCode id="54EDAE7C637A4A1D962A6FAC2F3FFC33" box="[1061,1083,995,1016]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="897">R</collectionCode>
|
||
. Blažek lgt.’; male (
|
||
<collectionCode id="54EDAE7C637A4A1D97236FAC2E81FC32" box="[1324,1413,995,1017]" pageId="14" pageNumber="897">BMNH</collectionCode>
|
||
): ‘Naauwpoort, / Petersburg / distr. /
|
||
<date id="46421079637A4A1D96A1684E2E1EFBDC" box="[1198,1306,1025,1047]" pageId="14" pageNumber="897" value="1928-10-25">25/x.1928</date>
|
||
/ G.
|
||
<collectionCode id="54EDAE7C637A4A1D974A684E2E5DFBDD" box="[1349,1369,1025,1046]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="897" type="Museum">V</collectionCode>
|
||
.Son’; male (
|
||
<collectionCode id="54EDAE7C637A4A1D917E686F28CCFBFD" box="[881,968,1056,1078]" pageId="14" pageNumber="897">BMNH</collectionCode>
|
||
): ‘Naauwpoort, / Petersburg Distr. / 25/ xii. 27. / G. van Son’; female (
|
||
<collectionCode id="54EDAE7C637A4A1D967A68712FCFFB9F" box="[1141,1227,1086,1108]" pageId="14" pageNumber="897">BMNH</collectionCode>
|
||
): ‘Lake / Nigami’; male (
|
||
<collectionCode id="54EDAE7C637A4A1D917E681228CCFBB8" box="[881,968,1117,1139]" pageId="14" pageNumber="897">BMNH</collectionCode>
|
||
): ‘
|
||
<taxonomicName id="F5FC4D3A637A4A1D91EF68122FACFBB8" authority="Burchell" authorityName="Burchell" box="[992,1192,1117,1139]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="897" phylum="Arthropoda" rank="species" species="vialis">P. vialis Burchell</taxonomicName>
|
||
/ = pierreti Amyot / det. K.G.Blair.’, ‘
|
||
<taxonomicName id="F5FC4D3A637A4A1D91EA68332F74FB59" box="[997,1136,1148,1170]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="901" phylum="Arthropoda" rank="genus">Psammodes</taxonomicName>
|
||
tuberculipennis’, ‘L. M. / 20-1-09’, ‘Pres. by / Imp. Bur. Ent. / Brit. Mus. 1925–93.’, ‘Lourenco Marques: /
|
||
<date id="46421079637A4A1D961268F62F8DFB05" box="[1053,1161,1209,1230]" pageId="14" pageNumber="897" value="1909-01-20">20.1.1909</date>
|
||
/ Howard Coll.’; female (
|
||
<collectionCode id="54EDAE7C637A4A1D913F6897288CFB25" box="[816,904,1240,1262]" pageId="14" pageNumber="897">BMNH</collectionCode>
|
||
): ‘
|
||
<collectingCountry id="4AEB7629637A4A1D919168972F03FB26" box="[926,1031,1240,1261]" name="Zambia" pageId="14" pageNumber="897">ZAMBIA</collectingCountry>
|
||
<quantity id="F5049B5C637A4A1D961F68972F5AFB26" box="[1040,1118,1240,1261]" metricMagnitude="3" metricUnit="m" metricValue="1.147" pageId="14" pageNumber="897" unit="m" value="1147.0">1147m</quantity>
|
||
/ Lukwakwa, West Lunga N.P. /
|
||
<geoCoordinate id="57C8507E637A4A1D916368B928ECFAC0" box="[876,1000,1270,1291]" degrees="12" direction="south" minutes="39" orientation="latitude" pageId="14" pageNumber="897" precision="15" seconds="40" value="-12.661111">S12°39’40”</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="57C8507E637A4A1D91FB68B92F75FAC7" box="[1012,1137,1270,1292]" degrees="24" direction="east" minutes="26" orientation="longitude" pageId="14" pageNumber="897" precision="15" seconds="13" value="24.436943">E24°26’13”</geoCoordinate>
|
||
/ 28–29.iv.14. Light Trap / leg. Smith,
|
||
<collectionCode id="54EDAE7C637A4A1D91B2695A28D7FAE1" box="[957,979,1301,1322]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="897">R</collectionCode>
|
||
., Takano, H., / Chmurova, L, & Smith, L.’, ‘
|
||
<taxonomicName id="F5FC4D3A637A4A1D9158697B28E7FA81" box="[855,995,1332,1354]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="901" phylum="Arthropoda" rank="genus">Psammodes</taxonomicName>
|
||
/ vialis / E. Ruzzier det. 2015’; male (
|
||
<collectionCode id="54EDAE7C637A4A1D913F691D2883FAA3" box="[816,903,1362,1384]" pageId="14" pageNumber="897">BMNH</collectionCode>
|
||
): ‘Damara Land’, ‘F. Bates / 81–19’; male and
|
||
<specimenCount id="24FAFD30637A4A1D9126693E28B5FA4C" box="[809,945,1393,1415]" pageId="14" pageNumber="897" type="female">two females</specimenCount>
|
||
(Ted MacRae): ‘
|
||
<collectionCode id="54EDAE7C637A4A1D9668693E2F79FA4D" box="[1127,1149,1393,1414]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="897">R</collectionCode>
|
||
.S. Africa Northern Prov. / waterberg, Goelhoutbosh /
|
||
<geoCoordinate id="57C8507E637A4A1D966469DF2FEAFA6E" box="[1131,1262,1424,1445]" degrees="24" direction="south" minutes="22" orientation="latitude" pageId="14" pageNumber="897" precision="15" seconds="34" value="-24.376112">24°22’34’’ S</geoCoordinate>
|
||
, 27°33’ 64’’ E’, ‘
|
||
<date id="46421079637A4A1D912269E128A7FA0F" box="[813,931,1454,1476]" pageId="14" pageNumber="897" value="1999-11-29">29.xi.1999</date>
|
||
,
|
||
<collectionCode id="54EDAE7C637A4A1D91A469E128B8FA08" box="[939,956,1454,1475]" name="Tavera, Department of Geology and Geophysics" pageId="14" pageNumber="897">T</collectionCode>
|
||
.C. MacRae / Nocturnally in sandy / ground in open woodland’; male (
|
||
<collectionCode id="54EDAE7C637A4A1D964169822F84FA29" box="[1102,1152,1485,1506]" pageId="14" pageNumber="897">MIZ</collectionCode>
|
||
<collectionCode id="54EDAE7C637A4A1D968869822FBFFA28" box="[1159,1211,1485,1507]" country="Indonesia" lsid="urn:lsid:biocol.org:col:15187" name="Java Sugar Experimental Station" pageId="14" pageNumber="897" type="Herbarium">PAS</collectionCode>
|
||
): same data;
|
||
<specimenCount id="24FAFD30637A4A1D975B69822880F9C9" pageId="14" pageNumber="897" type="female">three females</specimenCount>
|
||
(
|
||
<collectionCode id="54EDAE7C637A4A1D919A69A328E5F9C9" box="[917,993,1516,1538]" pageId="14" pageNumber="897">PERC</collectionCode>
|
||
): same data; female (
|
||
<collectionCode id="54EDAE7C637A4A1D96D069A32E1FF9C9" box="[1247,1307,1516,1538]" pageId="14" pageNumber="897">XXX</collectionCode>
|
||
): ‘
|
||
<collectingCountry id="4AEB7629637A4A1D973E69A32895F9D4" name="South Africa" pageId="14" pageNumber="897">SOUTH AFRICA</collectingCountry>
|
||
1965 / Bechuanaland 5 IV / Ngamiland Nokaneng’;
|
||
<specimenCount id="24FAFD30637A4A1D91A26A662F1BF9F4" box="[941,1055,1577,1599]" pageId="14" pageNumber="897" type="male">two males</specimenCount>
|
||
and a female (
|
||
<collectionCode id="54EDAE7C637A4A1D96CD6A662E0EF9F4" box="[1218,1290,1577,1599]" pageId="14" pageNumber="897">CASC</collectionCode>
|
||
): ‘SW Africa /
|
||
<geoCoordinate id="57C8507E637A4A1D91396A072896F996" box="[822,914,1608,1629]" degrees="19" direction="south" minutes="14" orientation="latitude" pageId="14" pageNumber="897" precision="925" value="-19.233334">19° 14’S</geoCoordinate>
|
||
/
|
||
<geoCoordinate id="57C8507E637A4A1D91A96A072F0DF996" box="[934,1033,1608,1629]" degrees="20" direction="east" minutes="14" orientation="longitude" pageId="14" pageNumber="897" precision="925" value="20.233334">20° 14’E’</geoCoordinate>
|
||
, ‘CO Handley Jr /
|
||
<date id="46421079637A4A1D96D86A072E65F996" box="[1239,1377,1607,1629]" pageId="14" pageNumber="897" value="1952-11-28">XI. 28, 1952</date>
|
||
’.
|
||
</paragraph>
|
||
<paragraph id="324336B9637A4A1C91266AC829DBFED3" blockId="14.[809,1425,1670,1907]" lastBlockId="15.[163,779,197,280]" lastPageId="15" lastPageNumber="898" pageId="14" pageNumber="897">
|
||
<emphasis id="0088EAAB637A4A1D91266AC828BFF957" box="[809,955,1671,1692]" italics="true" pageId="14" pageNumber="897">Diagnosis:</emphasis>
|
||
The presence of microtubercles between the tuberculate rows on the elytra of this species distinguishes it from the following species:
|
||
<taxonomicName id="F5FC4D3A637A4A1D91266AAC28C0F93C" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1896" box="[809,964,1762,1784]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="897" phylum="Arthropoda" rank="species" species="mashunus">
|
||
<emphasis id="0088EAAB637A4A1D91266AAC28C0F93C" box="[809,964,1762,1784]" italics="true" pageId="14" pageNumber="897">T. mashunus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F5FC4D3A637A4A1D91DA6AAC2F49F93C" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1899" box="[981,1101,1762,1784]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="897" phylum="Arthropoda" rank="species" species="mulleri">
|
||
<emphasis id="0088EAAB637A4A1D91DA6AAC2F49F93C" box="[981,1101,1762,1784]" italics="true" pageId="14" pageNumber="897">T. mulleri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F5FC4D3A637A4A1D96516AAC2FE3F93C" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" baseAuthorityName="Peringuey" baseAuthorityYear="1908" box="[1118,1255,1762,1784]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="897" phylum="Arthropoda" rank="species" species="schultzei">
|
||
<emphasis id="0088EAAB637A4A1D96516AAC2FE3F93C" box="[1118,1255,1762,1784]" italics="true" pageId="14" pageNumber="897">T. schultzei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F5FC4D3A637A4A1D96F76AAC2E5EF93C" box="[1272,1370,1762,1784]" class="Insecta" family="Tenebrionidae" genus="Psammodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="897" phylum="Arthropoda" rank="species" species="vialis">
|
||
<emphasis id="0088EAAB637A4A1D96F76AAC2E5EF93C" box="[1272,1370,1762,1784]" italics="true" pageId="14" pageNumber="897">T. vialis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F5FC4D3A637A4A1D91266B4D28BCF8DD" authorityName="Kaminski" authorityYear="2021" box="[809,952,1793,1815]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="897" phylum="Arthropoda" rank="species" species="waclawae">
|
||
<emphasis id="0088EAAB637A4A1D91266B4D28BCF8DD" box="[809,952,1793,1815]" italics="true" pageId="14" pageNumber="897">T. waclawae</emphasis>
|
||
</taxonomicName>
|
||
. This species can be differentiated from
|
||
<taxonomicName id="F5FC4D3A637A4A1D91266B6F28ABF8FE" authorityName="Kaminski & Gearner" authorityYear="2021" box="[809,943,1824,1845]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="897" phylum="Arthropoda" rank="species" species="tschinkeli">
|
||
<emphasis id="0088EAAB637A4A1D91266B6F28ABF8FE" box="[809,943,1824,1845]" italics="true" pageId="14" pageNumber="897">T.tschinkeli</emphasis>
|
||
</taxonomicName>
|
||
by the lack of a prominent lip-like structure on the margin of the prosternum, and the elytra, while round, are more elongate than
|
||
<emphasis id="0088EAAB637A4A1D96C06B112E5BF8B9" box="[1231,1375,1885,1907]" italics="true" pageId="14" pageNumber="897">
|
||
<taxonomicName id="F5FC4D3A637A4A1D96C06B112E5FF8B9" authorityName="Kaminski & Gearner" authorityYear="2021" box="[1231,1371,1885,1907]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="897" phylum="Arthropoda" rank="species" species="tschinkeli">T. tschinkeli</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The tubercles in this species are taller and less dense than those of
|
||
<taxonomicName id="F5FC4D3A637B4A1C930D6CAA2ACAFF32" box="[258,462,228,250]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="898" phylum="Arthropoda" rank="species" species="tuberculipennis">
|
||
<emphasis id="0088EAAB637B4A1C930D6CAA2ACAFF32" box="[258,462,228,250]" italics="true" pageId="15" pageNumber="898">T. tuberculipennis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="F5FC4D3A637B4A1C90086CAA296EFF32" authorityName="Gearner" authorityYear="2021" box="[519,618,228,250]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="898" phylum="Arthropoda" rank="species" species="herero">
|
||
<emphasis id="0088EAAB637B4A1C90086CAA296EFF32" box="[519,618,228,250]" italics="true" pageId="15" pageNumber="898">T. herero</emphasis>
|
||
</taxonomicName>
|
||
, with those of
|
||
<taxonomicName id="F5FC4D3A637B4A1C92AC6D4C2A6AFEDC" box="[163,366,258,280]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="898" phylum="Arthropoda" rank="species" species="tuberculipennis">
|
||
<emphasis id="0088EAAB637B4A1C92AC6D4C2A6AFEDC" box="[163,366,258,280]" italics="true" pageId="15" pageNumber="898">T. tuberculipennis</emphasis>
|
||
</taxonomicName>
|
||
being nearly confluent at times.
|
||
</paragraph>
|
||
<paragraph id="324336B9637B4A1C92AC6D0D2E90FE59" blockId="15.[163,779,321,1907]" lastBlockId="15.[827,1444,197,403]" pageId="15" pageNumber="898">
|
||
<emphasis id="0088EAAB637B4A1C92AC6D0D2A2EFE9C" box="[163,298,322,343]" italics="true" pageId="15" pageNumber="898">Description:</emphasis>
|
||
Length 28.0–33.0 mm, width of pronotum 10.0–13.0 mm and elytra 17.0–22.0 mm.
|
||
<emphasis id="0088EAAB637B4A1C90CE6D2F280EFEBE" box="[705,778,352,373]" italics="true" pageId="15" pageNumber="898">Head:</emphasis>
|
||
Hypognathous. Frons finely punctate (2–4 diameters apart); frontoclypeal suture course, with deep groove in middle; apical clypeal margin broadly shallowly emarginate to non-emarginate; clypeus projected toward front of body; apical margin of labrum shallowly to sharply emarginate medially, densely punctate (although punctures often fine) in apical half, margin of labrum densely covered with yellowish, acuminate setae. Eye comma-shaped, with reduced ventral part, strongly emarginate around epistomal base; with deep groove on temporal side. Mentum trapezoidal, with straight base, not fully filling buccal cavity; anterior margin not emarginate. Submentum semicircular, concave basally. Antenna slender, moderately covered in recumbent acuminate goldish setae; antennomere 2 short, equal to 0.1–0.2 of antennomere 3 length; antennomere 4 about half of antennomere 3 length; length of antenna slightly longer than pronotal length.
|
||
<emphasis id="0088EAAB637B4A1C92AC6FE92A1EFC70" box="[163,282,934,955]" italics="true" pageId="15" pageNumber="898">Prothorax:</emphasis>
|
||
Pronotal lateral margin rounded, well visible. Pronotum widest at middle. Disc dull, impunctate; anterior fully marginate, posterior margin occasionally absent at middle, anterior apices strongly produced. Hypomeron convex, without submarginal groove, impunctate but weakly rugulose in places. Prosternal process rounded in lateral view with small projection by coxa, longitudinally depressed in middle (ventral view). Anterior margin of prosternum straight to slightly projecting ventrally (lateral view).
|
||
<emphasis id="0088EAAB637B4A1C908F68F5280FFB04" box="[640,779,1210,1231]" italics="true" pageId="15" pageNumber="898">Pterothorax:</emphasis>
|
||
Scutellum densely covered with microtubercles. Elytra widest at middle, slightly rounded; disc impunctate, not covered by tubercles; lateral part (below humerus) covered with tubercles (organized in ~7–8 more or less regular rows on each elytron), with microtubercles scattered between rows; declivous portion on each elytron with additional 3–4 tuberculate rows with microtubercles in between; elytral margin not visible dorsally except in apical quarter, tuberculate rows extend more or less to lateral margin. Elytral slope relatively steep, elytral apex flattened. Epipleura, impunctate, not tuberculate, clearly distinguishable from neighbouring portion of elytra, widely enfolding fifth ventrite. Mesoventrite with deep median groove and elevated sides. Metaventrite impunctate, often densely setose (in males). Lateral regions of metaventrite (between coxae) short. Metaepisternal suture abbreviated posteriorly.
|
||
<emphasis id="0088EAAB637B4A1C902C6AAC2964F933" box="[547,608,1763,1784]" italics="true" pageId="15" pageNumber="898">Legs:</emphasis>
|
||
Covered with dense gold setae. Procoxa exposed basally. Apex of protibia with prominent denticle on outer margin, lateral carina terminating at middle to basal third; median spur reduced, reaching 0.5 of outer lateral spur length. Spurs on meso- and metatibiae of equal length. Tarsi narrowed laterally.
|
||
<emphasis id="0088EAAB637B4A1C96756CAB2FF6FF32" box="[1146,1266,228,249]" italics="true" pageId="15" pageNumber="898">Abdomen:</emphasis>
|
||
Ventrites 1–3 medially densely covered with goldish setae (males), moderately punctate and weakly rugulose; ventrite 5 densely punctate; ventrite 5 without submarginal sulcus.
|
||
<emphasis id="0088EAAB637B4A1C91906D112F32FEB8" box="[927,1078,350,371]" italics="true" pageId="15" pageNumber="898">Terminalia:</emphasis>
|
||
Aedeagus as in
|
||
<figureCitation id="AAC72A3C637B4A1C97176D112E9FFEB8" box="[1304,1435,350,372]" captionStart="Figure 1" captionStartId="2.[145,227,1253,1275]" captionTargetBox="[146,1421,201,1212]" captionTargetId="figure-279@2.[145,1425,195,1213]" captionTargetPageId="2" captionText="Figure 1. Phylogeny of Sepidiini imposed with new generic classification. Presented topology is the majority rule consensus of post-burn-in trees obtained in Bayesian analysis of the concatenated CAD, wg, COI JP, COI BC, COII, and 28S matrix. Posterior probabilities are displayed above branches, while the ultrafast IQTree bootstrap values are shown below.Morphological details shown on the right side of the figure corresponds to clades a–e on the tree. Habitus photographs presented on the left illustrates morphological incoherence of the genus Psammodes." figureDoi="http://doi.org/10.5281/zenodo.5724194" httpUri="https://zenodo.org/record/5724194/files/figure.png" pageId="15" pageNumber="898">Figure 1E</figureCitation>
|
||
. Ovipositor similar to others in the genus (
|
||
<figureCitation id="AAC72A3C637B4A1C971D6D322E80FE58" box="[1298,1412,381,403]" captionStart="Figure 6" captionStartId="10.[145,225,1641,1663]" captionTargetBox="[154,1423,208,1588]" captionTargetId="figure-81@10.[145,1425,195,1601]" captionTargetPageId="10" captionText="Figure 6. Female terminalia morphology of selected Sepidiini and Adelostomini: A, B, Toktokkus tuberculipennis; C–H, T. waclawae; I, Psammodes pinguis; J, K, Lepidochora kahani; L, M, Eurychora dilatata. Abbreviations: bc, vacuous of valvifer; bp, vacuous of paraproct;c, coxites; c2–c4, coxite plates 2–4; ip, inner plate of paraproct; op, outer plate of paraproct; pa, paraproct; pr, proctiger; ss, sensory setae; v, valvifer." figureDoi="http://doi.org/10.5281/zenodo.5724206" httpUri="https://zenodo.org/record/5724206/files/figure.png" pageId="15" pageNumber="898">Fig. 6A–H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="324336B9637B4A1C91346DF42E0CFDC4" blockId="15.[827,1443,443,527]" pageId="15" pageNumber="898">
|
||
<emphasis id="0088EAAB637B4A1C91346DF428B8FE1B" box="[827,956,443,464]" italics="true" pageId="15" pageNumber="898">Etymology:</emphasis>
|
||
Named after the locality from which the
|
||
<typeStatus id="ED47881B637B4A1C91346D952899FE3B" box="[827,925,474,496]" pageId="15" pageNumber="898" type="holotype">holotype</typeStatus>
|
||
was collected and is also the name of a tribe in the
|
||
<collectingRegion id="F038F85B637B4A1C918B6DB62F4BFDC4" box="[900,1103,505,527]" country="South Africa" name="Limpopo" pageId="15" pageNumber="898">Limpopo province</collectingRegion>
|
||
of
|
||
<collectingCountry id="4AEB7629637B4A1C967E6DB62E00FDC4" box="[1137,1284,505,527]" name="South Africa" pageId="15" pageNumber="898">South Africa</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |