155 lines
17 KiB
XML
155 lines
17 KiB
XML
<document id="038E034C55F8E1459873C5FAABC7D9EB" ID-CLB-Dataset="3320" ID-DOI="10.1093/zoolinnean/zlaa052" ID-GBIF-Dataset="fd4b2931-7bad-4644-b485-f365040ae01e" ID-ISSN="0024-4082" ID-Zenodo-Dep="5724192" ID-ZooBank="0810A91E-7D8D-4F56-B468-9D5B3A6115E9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619628947432" checkinUser="carolina" docAuthor="Kamiński, Marcin J., Gearner, Olivia M., Kanda, Kojun, Swichtenberg, Kali, Purchart, Luboš & Smith, Aaron D." docDate="2021" docId="BA5587AF63784A1E92A76A4C2A5BFEBD" docLanguage="en" docName="ZoolJLinnSoc.191.883-901.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Toktokkus congolensis Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith 2021" docType="treatment" docVersion="6" lastPageNumber="896" masterDocId="466CFFD763744A13920F6C4F2B04FFCB" masterDocTitle="First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group" masterLastPageNumber="901" masterPageNumber="883" pageNumber="895" updateTime="1698937967960" updateUser="ExternalLinkService">
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<mods:titleInfo id="F852A4B14356EC96F42BA79CE4137219">
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<mods:title id="F2BDACC4FCDC524AEB1D9F2D30EFA3F1">First insights into the phylogeny of tok-tokkie beetles (Tenebrionidae: Molurina, Phanerotomeina) and examination of the status of the Psammodes vialis species-group</mods:title>
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</mods:titleInfo>
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<mods:name id="64FD57FD1F9831071284C9F6747357ED" type="personal">
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<mods:role id="9DC10A37D65ACB0BC3C0E73C15259856">
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<mods:roleTerm id="55DABC461E32D5DF3B480B6B8B61FD4D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="082AE950F8D0369D5FC35589F0CF26A3">Kamiński, Marcin J.</mods:namePart>
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</mods:name>
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<mods:name id="BC948806FC00DFDEB59EF8DFE33830B7" type="personal">
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<mods:role id="B3B84AD82A101922BF9843DE4A28D74B">
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<mods:roleTerm id="9091004BE04ADFC2EFDA8FB34433A585">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="742A131071D341C358934E279DBA8696">Gearner, Olivia M.</mods:namePart>
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</mods:name>
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<mods:name id="FF06D786B68D8B117E40BDA8B87A46A7" type="personal">
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<mods:role id="53048AB66ED1DE00090525A8B85B180C">
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<mods:roleTerm id="FFB7A505DE23582295C3DFA71F3FFCEE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3A936927683134BA92D8DB97BEB709D5">Kanda, Kojun</mods:namePart>
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</mods:name>
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<mods:name id="E36F66074CBE7670E7AFC06D0BF9D1D2" type="personal">
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<mods:role id="9D1CF8B3D84E8208218D44166F6EF129">
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<mods:roleTerm id="B132B98280B1BC320347D8171EAD97C2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D4326086DC8ACCEA0BF52976CCC28BF">Swichtenberg, Kali</mods:namePart>
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</mods:name>
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</mods:role>
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<mods:namePart id="15A6CB3E33A0CA2F017D0FF7E034CBBA">Purchart, Luboš</mods:namePart>
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</mods:name>
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<mods:name id="FE4246529CE7B19EC770F609474DFB6D" type="personal">
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</mods:role>
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<mods:namePart id="CB3F762CEC848783967C1777229F49E4">Smith, Aaron D.</mods:namePart>
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<mods:typeOfResource id="F1F64E561642632708F4682B8DAB441D">text</mods:typeOfResource>
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<mods:title id="D737BDCF049E4273F586CBAD6664A8C9">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="6AC880D296BC58F85F7A822453507D6E">
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<mods:date id="EB83C6DD7D88789802D140BEFEA3A253">2021</mods:date>
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<mods:detail id="4A28709C785FBFF8447DDEE901866F19" type="pubDate">
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<mods:number id="04E2997EAE10F54217EA3EE582C29D0D">2020-06-16</mods:number>
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</mods:detail>
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<mods:detail id="A8A1F180E65372E7B46A9CECAAE38E36" type="volume">
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<mods:number id="14CFFBA1838D5E4E1AA49CDF9000B24F">191</mods:number>
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<mods:start id="3F4CD0B0C28EA160E6AB9D001518CC27">883</mods:start>
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<mods:end id="E78F0C940545F94209701078CA4027B0">901</mods:end>
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</mods:extent>
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<mods:classification id="7A8CA590D78D36ECE2F382DD494FCBF5">journal article</mods:classification>
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<mods:identifier id="C8C8890F3B07A3DFBCDA4B6A5D40AFD1" type="CLB-Dataset">3320</mods:identifier>
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<mods:identifier id="9BC173BC79F76AF623A92E5EC71BCF0C" type="DOI">10.1093/zoolinnean/zlaa052</mods:identifier>
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<mods:identifier id="718BA5B95BEDD89E9CE7F0D31DF3AC6C" type="GBIF-Dataset">fd4b2931-7bad-4644-b485-f365040ae01e</mods:identifier>
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<mods:identifier id="862CAC8B5E87A4A11D8CDD35D7591A50" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="7CB22D02FC72EE766979449ACA1831F4" type="Zenodo-Dep">5724192</mods:identifier>
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<mods:identifier id="7158DD9975F2E57D2FC873B73A1875E5" type="ZooBank">0810A91E-7D8D-4F56-B468-9D5B3A6115E9</mods:identifier>
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</mods:mods>
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<treatment id="BA5587AF63784A1E92A76A4C2A5BFEBD" ID-DOI="http://doi.org/10.5281/zenodo.5724208" ID-GBIF-Taxon="190683659" ID-Zenodo-Dep="5724208" LSID="urn:lsid:plazi:treatment:BA5587AF63784A1E92A76A4C2A5BFEBD" httpUri="http://treatment.plazi.org/id/BA5587AF63784A1E92A76A4C2A5BFEBD" lastPageId="13" lastPageNumber="896" pageId="12" pageNumber="895">
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<subSubSection id="7AE6653263784A1F92A76A4C29E5F9D1" box="[168,737,1538,1563]" pageId="12" pageNumber="895" type="nomenclature">
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||
<paragraph id="324336B963784A1F92A76A4C29E5F9D1" blockId="12.[168,737,1538,1563]" box="[168,737,1538,1563]" pageId="12" pageNumber="895">
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||
<heading id="690B81D563784A1F92A76A4C29E5F9D1" box="[168,737,1538,1563]" centered="true" fontSize="9" level="2" pageId="12" pageNumber="895" reason="2">
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||
<taxonomicName id="F5FC4D3A63784A1F92A76A4C29E5F9D1" authority="Kaminski & Gearner" authorityName="Kamiński & Gearner & Kanda & Swichtenberg & Purchart & Smith" authorityYear="2021" box="[168,737,1538,1563]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="901" phylum="Arthropoda" rank="genus" status="sp. nov.">
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TOKTOKKUS
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<emphasis id="0088EAAB63784A1F93356A492ADCF9D1" box="[314,472,1542,1562]" italics="true" pageId="12" pageNumber="895">CONGOLENSIS</emphasis>
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KAMIŃSKI & GEARNER
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="7AE6653263784A1F939E6A712A96F950" pageId="12" pageNumber="895" type="description">
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<paragraph id="324336B963784A1F939E6A712AFCF99C" blockId="12.[401,504,1598,1623]" box="[401,504,1598,1623]" pageId="12" pageNumber="895">
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(
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<figureCitation id="AAC72A3C63784A1F93956A712AF4F99C" box="[410,496,1598,1623]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="12" pageNumber="895">FIG. 3B</figureCitation>
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)
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</paragraph>
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<paragraph id="324336B963784A1F929E6A292A96F950" blockId="12.[145,761,1638,1752]" pageId="12" pageNumber="895">
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lsid
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<uri id="466D3ABB63784A1F92CC6A292A96F950" pageId="12" pageNumber="895">
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urn:lsid:zoobank.org:act:
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<uuid id="465A0C6C63784A1F93D66A292A96F950" pageId="12" pageNumber="895">86883CCF-78A5-4DD3- A2C6-0AFA496EB42A</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="7AE6653263784A1F929E6AEA2BD8F913" pageId="12" pageNumber="895" type="materials_examined">
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<paragraph id="324336B963784A1F929E6AEA2BD8F913" blockId="12.[145,761,1638,1752]" pageId="12" pageNumber="895">
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<emphasis id="0088EAAB63784A1F929E6AEA2A42F972" box="[145,326,1700,1722]" italics="true" pageId="12" pageNumber="895">Type material:</emphasis>
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<materialsCitation id="82943CE463784A1F935A6AEB2BD8F913" ID-GBIF-Occurrence="3411244305" collectionCode="BMNH" country="Democratic Republic of the Congo" location="Congo" pageId="12" pageNumber="895" specimenCount="1" typeStatus="holotype">
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<typeStatus id="ED47881B63784A1F935A6AEB2AC7F971" box="[341,451,1700,1722]" pageId="12" pageNumber="895" type="holotype">Holotype</typeStatus>
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(
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<collectionCode id="54EDAE7C63784A1F93D96AEB2935F971" box="[470,561,1700,1722]" pageId="12" pageNumber="895">BMNH</collectionCode>
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), male: ‘
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<collectingCountry id="4AEB762963784A1F90966AEB29E2F972" box="[665,742,1700,1721]" name="Democratic Republic of the Congo" pageId="12" pageNumber="895">Congo</collectingCountry>
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/ 80-32’.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="7AE6653263784A1F929E6B4D2F1DFED3" pageId="12" pageNumber="895" type="diagnosis">
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<paragraph id="324336B963784A1F929E6B4D2F1DFED3" blockId="12.[145,761,1793,1876]" lastBlockId="12.[809,1425,197,280]" pageId="12" pageNumber="895">
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<emphasis id="0088EAAB63784A1F929E6B4D2A0CF8DC" box="[145,264,1794,1815]" italics="true" pageId="12" pageNumber="895">Diagnosis:</emphasis>
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This species is most similar to
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<taxonomicName id="F5FC4D3A63784A1F90606B4D29FDF8DD" authorityName="Kaminski" authorityYear="2021" box="[623,761,1793,1815]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="895" phylum="Arthropoda" rank="species" species="waclawae">
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<emphasis id="0088EAAB63784A1F90606B4D29FDF8DD" box="[623,761,1793,1815]" italics="true" pageId="12" pageNumber="895">T. waclawae</emphasis>
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</taxonomicName>
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due to its large, dense, confluent tubercles, but can be distinguished by the apex of the elytra, which is straight and depressed with raised margins (
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<figureCitation id="AAC72A3C63784A1F973F6C8A2E86FF11" box="[1328,1410,197,219]" captionStart="Figure 3" captionStartId="5.[163,242,1678,1700]" captionTargetBox="[206,1393,199,1635]" captionTargetId="figure-29@5.[204,1401,195,1638]" captionTargetPageId="5" captionText="Figure 3. Habitus, morphological details, and labels of the newly discovered species representing Toktokkus: A, holotype of Toktokkus barclayi Kamiński & Gearner sp. nov.; B, holotype of T. congolensis Kamiński & Gearner sp. nov.; C, paratype of T. tschinkeli Kamiński & Gearner sp. nov.; D, holotype of T. herero Gearner sp. nov.; E, paratype of T. makuya Gearner sp. nov.; F, holotype of T. waclawae Kamiński sp. nov. Scale bar = 10.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5724200" httpUri="https://zenodo.org/record/5724200/files/figure.png" pageId="12" pageNumber="895">Fig. 3B</figureCitation>
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), while in
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<taxonomicName id="F5FC4D3A63784A1F91976CAB2F29FF32" authorityName="Kaminski" authorityYear="2021" box="[920,1069,228,249]" class="Insecta" family="Tenebrionidae" genus="Toktokkus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="895" phylum="Arthropoda" rank="species" species="waclawae">
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<emphasis id="0088EAAB63784A1F91976CAB2F29FF32" box="[920,1069,228,249]" italics="true" pageId="12" pageNumber="895">T. waclawae</emphasis>
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</taxonomicName>
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this structure is sloped and lacking a depression.
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</paragraph>
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</subSubSection>
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<subSubSection id="7AE6653263784A1E91266D0D2979FF31" lastPageId="13" lastPageNumber="896" pageId="12" pageNumber="895" type="description">
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<paragraph id="324336B963784A1E91266D0D2979FF31" blockId="12.[809,1425,321,1876]" lastBlockId="13.[163,779,197,250]" lastPageId="13" lastPageNumber="896" pageId="12" pageNumber="895">
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<emphasis id="0088EAAB63784A1F91266D0D28B9FE9C" box="[809,957,322,343]" italics="true" pageId="12" pageNumber="895">Description:</emphasis>
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Length 30.0 mm, width of pronotum 9.0 mm and elytra 12.0 mm.
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<emphasis id="0088EAAB63784A1F96826D2F2FD7FEBE" box="[1165,1235,352,373]" italics="true" pageId="12" pageNumber="895">Head:</emphasis>
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Hypognathous. Frons finely punctate (2–4 diameters apart) except at middle; frontoclypeal suture course, with deep groove in middle; apical clypeal margin broadly shallowly emarginate to non-emarginate; clypeus projected toward front of body; apical margin of labrum shallowly emarginate medially, scattered punctation in apical half, margin of labrum densely covered with yellowish, acuminate setae. Eye comma-shaped, with reduced ventral part, strongly emarginate around epistomal base; with deep groove on temporal side. Mentum trapezoidal, with straight base, not fully filling buccal cavity; anterior margin not emarginate. Submentum semicircular, concave basally. Antenna slender, moderately covered in recumbent acuminate goldish setae; antennomere 2 short, equal to 0.2 of antennomere 3 length; antennomere 4 about twothirds of antennomere 3 length; length of antenna equal to or slightly longer than pronotal length.
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<emphasis id="0088EAAB63784A1F91266FE928AEFC70" box="[809,938,934,955]" italics="true" pageId="12" pageNumber="895">Prothorax:</emphasis>
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Pronotal lateral margin rounded, well visible. Pronotum widest above middle. Disc dull, impunctate; anterior margin complete, basal margin absent at middle, anterior apices strongly produced. Hypomeron convex, without submarginal groove, impunctate. Prosternal process rounded in lateral view, longitudinally depressed in middle (ventral view). Anterior margin of prosternum straight.
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<emphasis id="0088EAAB63784A1F970C68322E94FB59" box="[1283,1424,1149,1170]" italics="true" pageId="12" pageNumber="895">Pterothorax:</emphasis>
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Scutellum densely covered with microtubercles. Elytra widest in basal third, slightly rounded; disc impunctate, without tubercles in middle; lateral part covered with nearly confluent tubercles organized in rows; declivous portion on each elytron with additional 3–4 tuberculate rows. Elytral slope relatively steep, elytral apex depressed with raised margin. Epipleura, impunctate, not tuberculate, clearly distinguishable from neighbouring portion of elytra, widely enfolding fifth ventrite. Mesoventrite with deep median groove and elevated sides. Metaventrite impunctate, setose. Lateral regions of metaventrite (between coxae) short. Metaepisternal suture abbreviated posteriorly.
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<emphasis id="0088EAAB63784A1F91266A642862F98B" box="[809,870,1579,1600]" italics="true" pageId="12" pageNumber="895">Legs:</emphasis>
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Covered with goldish setae. Procoxa exposed basally. Apex of protibia with prominent denticle on outer margin, lateral carina terminating at middle to basal third; median spur reduced, about half of outer lateral spur length. Spurs on meso- and metatibiae of equal length. Tarsi narrowed laterally.
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<emphasis id="0088EAAB63784A1F97146A8B2E94F912" box="[1307,1424,1732,1753]" italics="true" pageId="12" pageNumber="895">Abdomen:</emphasis>
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Ventrites 1–4 medially densely covered with goldish setae, weakly punctate and weakly rugulose; ventrite 5 without submarginal sulcus, densely punctured (~0.5 diameters apart), each puncture with single goldish setae.
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<emphasis id="0088EAAB63794A1E934A6C8A2ACFFF11" box="[325,459,197,218]" italics="true" pageId="13" pageNumber="896">Terminalia:</emphasis>
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Due to scarcity of available specimens, terminalia were not dissected.
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</paragraph>
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</subSubSection>
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<subSubSection id="7AE6653263794A1E92AC6D6D2A5BFEBD" pageId="13" pageNumber="896" type="etymology">
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<paragraph id="324336B963794A1E92AC6D6D2A5BFEBD" blockId="13.[163,779,290,374]" pageId="13" pageNumber="896">
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<emphasis id="0088EAAB63794A1E92AC6D6D2A28FEFC" box="[163,300,290,311]" italics="true" pageId="13" pageNumber="896">Etymology:</emphasis>
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Specific epithet is derived from
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<typeStatus id="ED47881B63794A1E90D96D6C280EFEF3" box="[726,778,291,312]" pageId="13" pageNumber="896">type</typeStatus>
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locality,
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<collectingCountry id="4AEB762963794A1E930E6D0E2A4EFE9D" box="[257,330,321,342]" name="Democratic Republic of the Congo" pageId="13" pageNumber="896">Congo</collectingCountry>
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, an area in Central Africa surrounding the
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<collectingCountry id="4AEB762963794A1E92C06D2F2A12FEBE" box="[207,278,352,373]" name="Democratic Republic of the Congo" pageId="13" pageNumber="896">Congo</collectingCountry>
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River.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |