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<document ID-CLB-Dataset="3346" ID-DOI="http://dx.doi.org/10.3897/alpento.5.77751" ID-GBIF-Dataset="b5931bca-640e-4550-a272-d4eee4b21d5c" ID-Pensoft-Pub="2535-0889-5-107" ID-Pensoft-UUID="6A7F0EBE16CF5F77B6838ECFD6AB0017" ID-ZooBank="873D2684880C42C58E9A0D708F22D827" ModsDocID="2535-0889-5-107" checkinTime="1637640615285" checkinUser="pensoft" docAuthor="Burckhardt, Daniel" docDate="2021" docId="A784E1AE0C9656749157C16823176AFB" docLanguage="en" docName="AlpEntomol 5: 107-113" docOrigin="Alpine Entomology 5" docPubDate="2021-11-22" docSource="http://dx.doi.org/10.3897/alpento.5.77751" docTitle="Ctenarytaina daleae Burckhardt 2020" docType="treatment" docVersion="5" id="6A7F0EBE16CF5F77B6838ECFD6AB0017" lastPageNumber="107" masterDocId="6A7F0EBE16CF5F77B6838ECFD6AB0017" masterDocTitle="On some Ctenarytaina species (Hemiptera, Psylloidea) from Gunung Kinabalu (Malaysia, Sabah)" masterLastPageNumber="113" masterPageNumber="107" pageNumber="107" updateTime="1732822687959" updateUser="ExternalLinkService">
<mods:mods id="2DE62BE2A08F089534FA9CA625DB4B89" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="46643F339D7983675524BC14D0FD6862">
<mods:title id="51866A83024E5E12C217D43EC5337B6D">On some Ctenarytaina species (Hemiptera, Psylloidea) from Gunung Kinabalu (Malaysia, Sabah)</mods:title>
</mods:titleInfo>
<mods:name id="50F1B727001F396DF2AD6EBABB674EE6" type="personal">
<mods:role id="0DAFB1C2FC3CF19D22C611AC29596627">
<mods:roleTerm id="F7C2600BE8F9227FB9C0F6F2B46AC817">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4BC6398B9395729257498EC244BCAD6D">Burckhardt, Daniel</mods:namePart>
<mods:nameIdentifier id="53459F1118374E4FB458FC25B7C64037" type="ORCID">https://orcid.org/0000-0001-8368-5268</mods:nameIdentifier>
<mods:affiliation id="B8B93C000B585EF4AE605CF733C80F6B">Naturhistorisches Museum, Augustinergasse 2, 4001 Basel, Switzerland</mods:affiliation>
<mods:nameIdentifier id="546A539ACE04912FC3137A5327B2B66D" type="email">daniel.burckhardt@bs.ch</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="8B943BFCF0C9F132FB40719E15747DCC">text</mods:typeOfResource>
<mods:relatedItem id="57962F747F44FB1AC3A901975EC167B2" type="host">
<mods:titleInfo id="4F0E2C488C9A3FB77AD26A0D6EBD2FC1">
<mods:title id="9718D2455F6267148B2D3834C52B434B">Alpine Entomology</mods:title>
</mods:titleInfo>
<mods:part id="77260C8029B0AB59C12EECBA99FF9C30">
<mods:date id="6621DD335056DFDE2D0E24060A7CB5D5">2021</mods:date>
<mods:detail id="CBBED0AFE294F9904865CC4E033688E8" type="pubDate">
<mods:number id="5D4673B309BCD6F947DC006F64A497F1">2021-11-22</mods:number>
</mods:detail>
<mods:detail id="F137A8A6E685B78B798951A9321E9BF0" type="volume">
<mods:number id="E17ADE99881E6B0550D6A5B8F56A4D74">5</mods:number>
</mods:detail>
<mods:extent id="9816FBF8672EC31148F651DEE33D1BEC" unit="page">
<mods:start id="AE9B4654155A3A6D1739C6E2B7496235">107</mods:start>
<mods:end id="1CAFAFB8A000163FBEE5F51AB0CCCC8D">113</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="4E78B9617B307F6CF84D8AEF92A93171">
<mods:url id="9FCB8AE1A82E330F0543538D0C881420">http://dx.doi.org/10.3897/alpento.5.77751</mods:url>
</mods:location>
<mods:classification id="A55EF3F030EC72627D40469D7C8464B9">journal article</mods:classification>
<mods:identifier id="E15ECE7C0053CBBCD3C34D3B8B63D72A" type="DOI">http://dx.doi.org/10.3897/alpento.5.77751</mods:identifier>
<mods:identifier id="FD2E9DC095442F7D5BC35B93BF67577C" type="Pensoft-Pub">2535-0889-5-107</mods:identifier>
<mods:identifier id="6634566D0CB484F8534241C6C41EB719" type="ZooBank">873D2684880C42C58E9A0D708F22D827</mods:identifier>
<mods:identifier id="37EBF32C85469CC012608209CFDEAC24" type="Pensoft-UUID">6A7F0EBE16CF5F77B6838ECFD6AB0017</mods:identifier>
</mods:mods>
<treatment id="A784E1AE0C9656749157C16823176AFB" ID-GBIF-Taxon="190549092" LSID="urn:lsid:plazi:treatment:A784E1AE0C9656749157C16823176AFB" httpUri="http://treatment.plazi.org/id/A784E1AE0C9656749157C16823176AFB" lastPageNumber="107" pageId="0" pageNumber="107" scope_order="Hemiptera" scope_superFamily="Psylloidea">
<subSubSection id="A7A25897691B260E9FCD63D659736DF0" pageId="0" pageNumber="107" type="nomenclature">
<paragraph id="2952FD097461BEE47C4F5A27A70CDD78" pageId="0" pageNumber="107">
<taxonomicName id="6DA848601ACB5D2C2AABE2AD39E90EE3" LSID="A784E1AE-0C96-5674-9157-C16823176AFB" authority="Burckhardt, 2020" authorityName="Burckhardt" authorityYear="2020" class="Insecta" family="Aphalaridae" genus="Ctenarytaina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenarytaina daleae" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="daleae">Ctenarytaina daleae Burckhardt, 2020</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="FA49660574D33F59A1BBCD5D4C409EE0" pageId="0" pageNumber="107" type="description">
<paragraph id="D8F0FC86E1FC256E03C9436AFAFA38F0" pageId="0" pageNumber="107">
<figureCitation id="CDB86A94CE3E291741728179C886B074" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Ctenarytaina spp., head, in dorsal view. 1. C. daleae; 2. C. lienhardi sp. nov.; 3. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures1-3" httpUri="https://binary.pensoft.net/fig/613526" pageId="0" pageNumber="107">Figs 1</figureCitation>
<figureCitation id="2455A0AF38E844C8480506C3F6608F70" captionStart="Figures 4–9" captionStartId="F2" captionText="Figures 4 - 9. Ctenarytaina spp. 4, 6, 8. Forewing; 5, 7, 9. Surface spinules in cell r 2 above bifurcation of vein M. 4, 5. C. daleae; 6, 7. C. lienhardi sp. nov.; 8, 9. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures4-9" httpUri="https://binary.pensoft.net/fig/613527" pageId="0" pageNumber="107">, 4</figureCitation>
<figureCitation id="E5AA08069D2FF194BA383E9BDDAE16AD" captionStart="Figures 4–9" captionStartId="F2" captionText="Figures 4 - 9. Ctenarytaina spp. 4, 6, 8. Forewing; 5, 7, 9. Surface spinules in cell r 2 above bifurcation of vein M. 4, 5. C. daleae; 6, 7. C. lienhardi sp. nov.; 8, 9. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures4-9" httpUri="https://binary.pensoft.net/fig/613527" pageId="0" pageNumber="107">, 5</figureCitation>
<figureCitation id="D83696EF1BE804DE4D4CA9C5DF74482A" captionStart="Figures 10–24" captionStartId="F3" captionText="Figures 10 - 24. Ctenarytaina spp., terminalia. 10, 15, 18. Male terminalia, in profile; 11, 16, 19. Distal portion of aedeagus; 12, 17, 20. Inner face of paramere; 13, 21, 23. Female terminalia, in profile; 14, 22, 24. Dorsal and ventral valvulae. 10 - 14 C. daleae; 15 - 17, 21, 22. C. lienhardi sp. nov.; 18 - 20, 23, 24. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures10-24" httpUri="https://binary.pensoft.net/fig/613528" pageId="0" pageNumber="107">, 10-14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="401D52AC5808F01FFAB36C0C680778A3" pageId="0" pageNumber="107" type="reference_group">
<paragraph id="5CB70196731BD14C014BE48B17E5F27A" pageId="0" pageNumber="107">
<taxonomicName id="74E933FC0168672BC2504B0DE33AF60C" authorityName="Burckhardt" authorityYear="2020" class="Insecta" family="Aphalaridae" genus="Ctenarytaina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenarytaina daleae" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="daleae">Ctenarytaina daleae</taxonomicName>
Burckhardt:
<bibRefCitation id="7E2066D6EF887429EC77C9871EFEC4DB" DOI="https://doi.org/10.12976/jib/2020.20.2.1" author="Burckhardt, D" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="107" pagination="35 - 58" refId="B3" refString="Burckhardt, D, Giliomee, JH, Hamilton-Attwell, VL, Queiroz, DL, 2020. A review of jumping plant lice of the genus Ctenarytaina (Hemiptera, Psylloidea, Aphalaridae) associated with Syzygium (Myrtaceae). Journal of Insect Biodiversity 20 (2): 35 - 58, DOI: https://doi.org/10.12976/jib/2020.20.2.1" title="A review of jumping plant lice of the genus Ctenarytaina (Hemiptera, Psylloidea, Aphalaridae) associated with Syzygium (Myrtaceae)." url="https://doi.org/10.12976/jib/2020.20.2.1" volume="20" year="2020">Burckhardt et al. (2020</bibRefCitation>
: 44), p.p. [description and figures ♂, key]
</paragraph>
</subSubSection>
<subSubSection id="A7519749315D8B48EEEACE3C0E400B63" pageId="0" pageNumber="107" type="type locality">
<paragraph id="D1069A69FDF666C9693B80E2C8F5E581" pageId="0" pageNumber="107">Type locality.</paragraph>
<paragraph id="2A28EE3ED9A3382A3487CD01DA3136E1" pageId="0" pageNumber="107">
Malaysia, Sabah, Ranau, Gunung Kinabalu,
<geoCoordinate id="B1A4B5616C0BE7C2470597204762B73E" degrees="6.0428" direction="north" orientation="latitude" precision="5" value="6.0428">6.0428°N</geoCoordinate>
,
<geoCoordinate id="B8094818474A07C1EB2EDB30641CB431" degrees="116.5587" direction="east" orientation="longitude" precision="5" value="116.5587">116.5587°E</geoCoordinate>
, 2600 m.
</paragraph>
</subSubSection>
<subSubSection id="9191210F44859B58066A5C3DA5E1172C" pageId="0" pageNumber="107" type="materials_examined">
<paragraph id="DD1729D7CDA61A7F62E0C1346EF850B8" pageId="0" pageNumber="107">Material examined.</paragraph>
<paragraph id="2C3AA702752676AFD37A6C77E7AB60BC" pageId="0" pageNumber="107">
<materialsCitation id="288CDDDB918CBDDDDD1ADE7E1B67E886" collectingDate="1982-04-29" collectingDateMax="1987-05-20" collectingDateMin="1982-04-29" collectingMethod="interception trap" collectorName="D. Burckhardt, I. Loebl, Layang Layang, A. Smetana, Laban Rata" country="Malaysia" elevation="2600" latitude="6.0428" location="Gunung Kinabalu" longLatPrecision="7" longitude="116.5587" municipality="Ranau" specimenCount="190" specimenCount-female="112" specimenCount-male="78" stateProvince="Sabah" typeStatus="Holotype">
<collectingCountry id="B981C446181EAD166334410476851268" name="Malaysia">Malaysia</collectingCountry>
:
<emphasis id="35A9E88559D9682EA6C4CEDDA813E05E" bold="true" italics="true" pageId="0" pageNumber="107">
<typeStatus id="D84FA08839914907672BC9CE1CC59503">Holotype</typeStatus>
</emphasis>
<specimenCount id="37733A20021F161889924A8590E39CF9">♂</specimenCount>
,
<collectingRegion id="9CA14A8FABB55A42D9BA00955815DFD3" country="Malaysia" name="Sabah">Sabah</collectingRegion>
:
<collectingMunicipality id="FCD1E20932B559A2811600FC7C8C5644">Ranau</collectingMunicipality>
,
<location id="9C333B645009A0EFFB4D3C0ED540B0F5" LSID="urn:lsid:plazi:treatment:A784E1AE0C9656749157C16823176AFB:9C333B645009A0EFFB4D3C0ED540B0F5" country="Malaysia" latitude="6.0428" longLatPrecision="7" longitude="116.5587" municipality="Ranau" name="Gunung Kinabalu" stateProvince="Sabah">Gunung Kinabalu</location>
,
<geoCoordinate id="6E027E8FB5D5D7BA9D927E53C9BF5027" degrees="6.0428" direction="north" orientation="latitude" precision="5" value="6.0428">6.0428°N</geoCoordinate>
,
<geoCoordinate id="3CF086E2A55668658512356F529592B0" degrees="116.5587" direction="east" orientation="longitude" precision="5" value="116.5587">116.5587°E</geoCoordinate>
,
<quantity id="DCE7D0A8A105D90B7108DEEEF771BAF3" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation id="59BAE95EAD34EE765D9A90D9212C3498" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<collectingDate id="FAF23092EA9020EA9B2BF0018C1A7965" value="1987-05-02">2.v.1987</collectingDate>
,
<taxonomicName id="E14ADC6791E90C32649EAA001855F1F5" class="Insecta" family="Myrtaceae" genus="Syzygium" kingdom="Animalia" lsidName="Syzygium korthalsianum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="korthalsianum">
<emphasis id="0BEB311CC64A2AA2A598A956902C08B7" italics="true" pageId="0" pageNumber="107">Syzygium korthalsianum</emphasis>
</taxonomicName>
(
<collectorName id="F73E942ADC82CB7779D7899DE2F12C1C">D. Burckhardt</collectorName>
&amp;
<collectorName id="948BAEBE5278F11EA89A5058E5796A4E">
I.
<normalizedToken id="21D779A1930069B69F460F2B548AFD31" originalValue="Löbl">Loebl</normalizedToken>
</collectorName>
) #8751 (MNHG, slide mounted). -
<collectingCountry id="B4CC8F3CD65BF9E1F7CC57157C4B6F9C" name="Malaysia">Malaysia</collectingCountry>
:
<specimenCount id="ED46A4CA201FF60ABB292BE0B0BD8B71" type="male">6 ♂</specimenCount>
,
<specimenCount id="0FB554DC9E5B376D25DB92D1214CD451" type="female">8 ♀</specimenCount>
, same but summit trail,
<quantity id="76C6AA3AA58BD2D06BB2ADB9EBB390BD" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" unit="m" value="3230.0">
<elevation id="677A0B3D968F4379443B41AA222B3C28" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" unit="m" value="3230.0">3230 m</elevation>
</quantity>
,
<taxonomicName id="B7FAE840392CC21D35157DDF499B62E2" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="3D894BD3275131A77FBBC68C6B7FFB11" italics="true" pageId="0" pageNumber="107">Leptospermum</emphasis>
</taxonomicName>
sp.,
<collectingDate id="595B97D311E08305FED14453C8E44E09" value="1982-04-29">29.iv.1982</collectingDate>
,
<taxonomicName id="1A971C627BAB19A4E0CB5364DEA27B98" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="FBB9708D847DD0A9FEE062802154042D" italics="true" pageId="0" pageNumber="107">Leptospermum</emphasis>
</taxonomicName>
scrub (
<collectorName id="6A802555A128471E326469DE77140FCE">D. Burckhardt</collectorName>
) #8277;
<specimenCount id="BA432506207C1A54E8EE0BB5DB194A61" type="male">10 ♂</specimenCount>
,
<specimenCount id="3DBCB29755DDABC72EA418AF2B09FC59" type="female">12 ♀</specimenCount>
, same but
<quantity id="CA4D2C49F60FF24B1C5132243C2878CC" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" unit="m" value="3230.0">
<elevation id="444CD7F30E8E5FD61015A3EFF7C0DB2F" metricMagnitude="3" metricUnit="m" metricValue="3.2299999999999995" unit="m" value="3230.0">3230 m</elevation>
</quantity>
,
<taxonomicName id="DABDB856D7C906C1B3E324D81A76EDEE" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum recurvum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="recurvum">
<emphasis id="1763EAF5ECAA2A721C85C1367C2807C4" italics="true" pageId="0" pageNumber="107">Leptospermum recurvum</emphasis>
</taxonomicName>
, #8278;
<specimenCount id="B52986BDCC834720D75403B22B38DFA8" type="male">1 ♂</specimenCount>
,
<specimenCount id="C5B80863F546E2CC85BEAFFD6223F066" type="female">1 ♀</specimenCount>
, same but
<quantity id="49AF2556F38D891ADE658ED425C6A741" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation id="25EB54D37C014C4208277EFB60D65543" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<collectingDate id="29B2329B69989BF32B70BDBDA6DF15A0" value="1987-05-01">1.v.1987</collectingDate>
(
<collectorName id="791BFA0A372FB98D0D7FCA3C54811D0F">D. Burckhardt</collectorName>
&amp;
<collectorName id="730D09F879B1E44D43F1C83D00A34E91">
I.
<normalizedToken id="977AD9AC3B443714ABF412BDE4B90830" originalValue="Löbl">Loebl</normalizedToken>
</collectorName>
) #8735;
<specimenCount id="944C11A8814B5F364E9E2F4FA8F3F4A8" type="male">21 ♂</specimenCount>
,
<specimenCount id="6C6A267F6F58E00F415A1813467603F7" type="female">36 ♀</specimenCount>
, same but
<quantity id="05F3B7A5B9FFC7AB0AE5BD507CD4273D" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation id="9BCCB47064D312E9695EA188375CD8F9" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<collectingDate id="360E608B2B46698DD5E7A3B0E05904C2" value="1987-05-02">2.v.1987</collectingDate>
,
<taxonomicName id="A6AF9E5FC74341D10A5C85924E3BC76A" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum recurvum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="recurvum">
<emphasis id="877630F04D09ABD57B0E51E6D39A7C60" italics="true" pageId="0" pageNumber="107">Leptospermum recurvum</emphasis>
</taxonomicName>
, #8747, #8748, #8749;
<specimenCount id="8A91A3689FDC535719F5470070D582F8" type="male">6 ♂</specimenCount>
,
<specimenCount id="FA3A7C440EBB8CDA8E9ACBDEF42C7A8E" type="female">6 ♀</specimenCount>
, same but
<quantity id="521570253C53D6FFBDF60A2008CA6E6D" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation id="07EF045513BDE06544C3103304971F3C" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<collectingDate id="042FECA8CFB8E6E0272BC80421A30B17" value="1987-05-02">2.v.1987</collectingDate>
, general sweeping of vegetation, #8752;
<specimenCount id="7E014CA9E22BC7FDD5743E7590721108" type="male">2 ♂</specimenCount>
,
<specimenCount id="06C08C56563DF51231A89DD0BD747310" type="female">9 ♀</specimenCount>
, same but
<quantity id="BCFA0F88B34A6417B8D33FF29E940E14" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation id="B2D8486953AC091283DE029648C0C1E5" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<collectingDate id="160C49FE7A48C7F1521C10DE499E15D9" value="1987-05-02">2.v.1987</collectingDate>
,
<taxonomicName id="B7C385F0B716C72F9F11716A2FC50F97" authorityName="Blume" authorityYear="1827" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum javanicum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="0D5169B9F4BFC5F5FF6852E98F77CF2C" italics="true" pageId="0" pageNumber="107">Leptospermum javanicum</emphasis>
</taxonomicName>
, #8753;
<specimenCount id="548BB3902CDBF29590942968E7C1A982" type="male">12 ♂</specimenCount>
,
<specimenCount id="FAF2FC99B255B60282BA58D2E5C4B05A" type="female">13 ♀</specimenCount>
, same but
<quantity id="498DC889CD53A6A439581C86F6783BF6" metricMagnitude="3" metricUnit="m" metricValue="3.3" unit="m" value="3300.0">
<elevation id="7529CD3FADB2E130C7DEDB0E45478F2D" metricMagnitude="3" metricUnit="m" metricValue="3.3" unit="m" value="3300.0">3300 m</elevation>
</quantity>
,
<collectingDate id="A384FDF419A8CB84CF9CD9FAD913EF10" value="1987-05-04">4.v.1987</collectingDate>
,
<taxonomicName id="FAE07E8C8AE83F98B0CF12D65EEC8CF6" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum recurvum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="recurvum">
<emphasis id="2155693D55B30132831322CC5061FEF4" italics="true" pageId="0" pageNumber="107">Leptospermum recurvum</emphasis>
</taxonomicName>
, #8756;
<specimenCount id="27E0388396C5732925FB3402587EA258" type="male">4 ♂</specimenCount>
,
<specimenCount id="3AFADF09D4EC0428E8F3D354E793F410" type="female">4 ♀</specimenCount>
, same but below
<collectorName id="FC70F6F97950D51085E5E3D5344B1DB0">Layang Layang</collectorName>
,
<quantity id="B2783FD5FDB1B7DDAAC28756EF002C94" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<elevation id="B69CBC5F83C87BB7A9BF82E7BE9FE41B" metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2600 m</elevation>
</quantity>
,
<collectingDate id="3FEB60E6E3CA4E1BDDAE0571BD7DAD4C" value="1987-05-02" valueMax="1987-05-08" valueMin="1987-05-02">2-8.v.1987</collectingDate>
,
<collectingMethod id="B69BD3A96822FC71631F34F23C79A6EE">interception trap</collectingMethod>
(
<collectorName id="52B2701C6D0CCFAF2CF69D6A2BBB46EE">A. Smetana</collectorName>
);
<specimenCount id="71761A449F442F8069B5BBD2CC8F8888" type="male">7 ♂</specimenCount>
,
<specimenCount id="A8A31057D46DECE514282173EDA8E095" type="female">11 ♀</specimenCount>
, same but below
<collectorName id="AD8C1A6C3DAC00FAFA2FDF2DF3841A80">Laban Rata</collectorName>
,
<quantity id="F5450B23018201BC258F472EEAB51F99" metricMagnitude="3" metricUnit="m" metricValue="3.155" unit="m" value="3155.0">
<elevation id="521798E78F4AFB978DD460161A1DCCB1" metricMagnitude="3" metricUnit="m" metricValue="3.155" unit="m" value="3155.0">3155 m</elevation>
</quantity>
,
<collectingDate id="B69243D2F02C67E1090E7E958F79F304" value="1987-05-05">5.v.1987</collectingDate>
;
<specimenCount id="7FB51661CC3F74B28EC8D657DC65C596" type="male">1 ♂</specimenCount>
,
<specimenCount id="EF7FC9686710B933768ED32768DA9744" type="female">5 ♀</specimenCount>
, same but
<collectorName id="1F0034E66C6EA76E72F3A70E9AAFEB35">Laban Rata</collectorName>
,
<quantity id="56E8C6503179A9F7CE9CA1403A788E6E" metricMagnitude="3" metricUnit="m" metricValue="3.2" unit="m" value="3200.0">
<elevation id="EFE961813C38FC70ABB67185E0CFBEC8" metricMagnitude="3" metricUnit="m" metricValue="3.2" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<collectingDate id="088206B0AF10276085FAEBBB29E48231" value="1987-05-04" valueMax="1987-05-08" valueMin="1987-05-04">4-8.v.1987</collectingDate>
,
<collectingMethod id="328837D8F3F51416B517FE2D27FA0341">interception trap</collectingMethod>
;
<specimenCount id="C53F42F62902E42B6538D2231E6FD3B6" type="male">8 ♂</specimenCount>
,
<specimenCount id="4B9AC0A0D373963FBF486B93EB6587D0" type="female">7 ♀</specimenCount>
, same but
<quantity id="91E916AC7F541F6947E7615312CE244F" metricMagnitude="3" metricUnit="m" metricValue="3.2" unit="m" value="3200.0">
<elevation id="A72954A19A3FD7BE2B905EFEF05E049D" metricMagnitude="3" metricUnit="m" metricValue="3.2" unit="m" value="3200.0">3200 m</elevation>
</quantity>
,
<collectingDate id="5BCCD3D551563B6E97E54CD2CCBF95E4" value="1987-05-09" valueMax="1987-05-20" valueMin="1987-05-09">9-20.v.1987</collectingDate>
,
<collectingMethod id="97255AA09B84FC6C82900C810873B00B">interception trap</collectingMethod>
(MHNG, NHMB, dry and slide mounted).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="9CDEB3EF0A71FF4958F1AB5A09B03FFF" pageId="0" pageNumber="107" type="diagnosis">
<paragraph id="E322E447C82A001B6809DE085DB1FB1E" pageId="0" pageNumber="107">Diagnosis.</paragraph>
<paragraph id="BDF402DE2EE37089C0ED5D9FC07BB0BE" pageId="0" pageNumber="107">Genal processes 0.3 times as long as vertex along mid-line, irregularly rounded anteriorly. Forewing oblong oval, widest in the middle, 2.3-2.7 times as long as broad, broadly rounded apically; vein C+Sc mostly straight, weakly concave in proximal third, cell c+sc narrow. Surface spinules present in all cells, forming cellular pattern; in cell r2 above bifurcation of vein M, the cells are irregularly hexagonal consisting of two indistinct rows of spinules. No extra pore fields developed on abdominal intersegmental membrane. Basal segment of proctiger weakly curved posteriorly, hind margin with a row of stout setae; apical segment 0.2-0.3 times as long as basal segment. Paramere digitiform. Distal segment of aedeagus cuneate. Female terminalia strongly narrowed medially, bearing each an apical process on proctiger and subgenital plate, female proctiger dorsally serrate.</paragraph>
</subSubSection>
<subSubSection id="114C2CA38C20CDDCF31AFDA68C112515" pageId="0" pageNumber="107" type="redescription">
<paragraph id="0E89AAE1F93595FF63B2F56003A3D75A" pageId="0" pageNumber="107">Redescription.</paragraph>
<paragraph id="445D352649710FFD977E21E4E6B44549" pageId="0" pageNumber="107">Adult. Colouration. Head and thorax light reddish brown. Vertex with dark brown dot in the middle of either half; genal processes dark brown at base, yellow apically. Antenna light orange brown at base, getting gradually darker from segment 6 to apex which is dark brown or black. Pronotum with each two submedian dark dots and mesopraescutum with each one submedian dark dot along fore margin on either side. Legs yellow; profemora light greyish brown. Forewings light ochreous or amber-coloured, slightly lighter along fore margin; veins concolourous with membrane. Abdomen yellow or orange; base of female proctiger light brown. Younger specimens with less extended dark colour.</paragraph>
</subSubSection>
<subSubSection id="F22E98611BEF1AC73AF5567A6A99070F" pageId="0" pageNumber="107" type="structure">
<paragraph id="04A262B161927C5CA83D7A842201766F" pageId="0" pageNumber="107">Structure.</paragraph>
<paragraph id="5771CCDC680386C9052F215CDE926325" pageId="0" pageNumber="107">
Conforming to the generic description of
<bibRefCitation id="969F9B7402C26BAA75D59DFB64A3CDDC" DOI="https://doi.org/10.12976/jib/2020.20.2.1" author="Burckhardt, D" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="107" pagination="35 - 58" refId="B3" refString="Burckhardt, D, Giliomee, JH, Hamilton-Attwell, VL, Queiroz, DL, 2020. A review of jumping plant lice of the genus Ctenarytaina (Hemiptera, Psylloidea, Aphalaridae) associated with Syzygium (Myrtaceae). Journal of Insect Biodiversity 20 (2): 35 - 58, DOI: https://doi.org/10.12976/jib/2020.20.2.1" title="A review of jumping plant lice of the genus Ctenarytaina (Hemiptera, Psylloidea, Aphalaridae) associated with Syzygium (Myrtaceae)." url="https://doi.org/10.12976/jib/2020.20.2.1" volume="20" year="2020">Burckhardt et al. (2020)</bibRefCitation>
. Body length ♂ 1.4-1.5 mm, ♀ 1.5-1.9 mm (6 ♂, 6 ♀). Head deflexed 45° from longitudinal axis of body. Vertex rhomboidal, weakly concave at base; preocular sclerite forming flat tubercule; genal processes 0.3 times as long as vertex along mid-line, irregularly rounded anteriorly, contiguous medially; eyes weakly
<normalizedToken id="58243C6C20A9EEBB5CD3281ED184B9B8" originalValue="‘stalked’">'stalked'</normalizedToken>
(Fig.
<figureCitation id="5FBF1B0FBB9D420B49FA5149DDAEC22A" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Ctenarytaina spp., head, in dorsal view. 1. C. daleae; 2. C. lienhardi sp. nov.; 3. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures1-3" httpUri="https://binary.pensoft.net/fig/613526" pageId="0" pageNumber="107">1</figureCitation>
). Antenna 0.6-1.0 times as long as head width. Metatibia 0.5-0.7 times as long as head width, weakly widening to apex, with 5 irregularly spaced apical spurs. Forewing (Fig.
<figureCitation id="11EAAC280C1C95E07EA8176252132F7A" captionStart="Figures 4–9" captionStartId="F2" captionText="Figures 4 - 9. Ctenarytaina spp. 4, 6, 8. Forewing; 5, 7, 9. Surface spinules in cell r 2 above bifurcation of vein M. 4, 5. C. daleae; 6, 7. C. lienhardi sp. nov.; 8, 9. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures4-9" httpUri="https://binary.pensoft.net/fig/613527" pageId="0" pageNumber="107">4</figureCitation>
) oblong oval, widest in the middle, 2.3-3.0 times as long as head width, 2.3-2.7 times as long as broad, broadly rounded apically; pterostigma, at base narrower than adjacent part of cell r1, regularly narrowing to apex, ending at apical fifth of wing; vein C+Sc mostly straight, weakly concave in proximal third, cell c+sc narrow; vein Rs almost straight, vein M long, with short, weakly diverging branches, vein Cu1a relatively straight, reaching the wing margin distinctly distal to bifurcation of vein M. Surface spinules present in all cells, forming cellular pattern; in cell r2 above bifurcation of vein M, the cells are irregularly hexagonal consisting of two indistinct rows of spinules (Fig.
<figureCitation id="11768C7E81B6537966CD453F17220FD9" captionStart="Figures 4–9" captionStartId="F2" captionText="Figures 4 - 9. Ctenarytaina spp. 4, 6, 8. Forewing; 5, 7, 9. Surface spinules in cell r 2 above bifurcation of vein M. 4, 5. C. daleae; 6, 7. C. lienhardi sp. nov.; 8, 9. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures4-9" httpUri="https://binary.pensoft.net/fig/613527" pageId="0" pageNumber="107">5</figureCitation>
). No extra pore fields developed on abdominal intersegmental membrane. Male terminalia as in Figs
<figureCitation id="22AFAD1C56E70499D0479D840AA942EF" captionStart="Figures 10–24" captionStartId="F3" captionText="Figures 10 - 24. Ctenarytaina spp., terminalia. 10, 15, 18. Male terminalia, in profile; 11, 16, 19. Distal portion of aedeagus; 12, 17, 20. Inner face of paramere; 13, 21, 23. Female terminalia, in profile; 14, 22, 24. Dorsal and ventral valvulae. 10 - 14 C. daleae; 15 - 17, 21, 22. C. lienhardi sp. nov.; 18 - 20, 23, 24. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures10-24" httpUri="https://binary.pensoft.net/fig/613528" pageId="0" pageNumber="107">10-12</figureCitation>
. Proctiger 0.4-0.6 times as long as head width; basal segment, in profile, weakly curved posteriorly, irregularly beset with fine setae, hind margin with a row of stout setae; apical segment tubular, 0.2-0.3 times as long as basal segment. Subgenital plate, in profile, triangular, with almost straight dorsal margin; sparsely beset with short setae. Paramere, in profile, digitiform, almost straight anteriorly, wavy posteriorly, broadly rounded apically; inner face densely coverd in moderately long bristles. Distal portion of aedeagus slender in basal half, widening towards apex which is rounded; sclerotised end tube of ductus ejaculatorius small, weakly curved. Female terminalia as in Fig.
<figureCitation id="E5116D497B69A021851D0069A96942C0" captionStart="Figures 10–24" captionStartId="F3" captionText="Figures 10 - 24. Ctenarytaina spp., terminalia. 10, 15, 18. Male terminalia, in profile; 11, 16, 19. Distal portion of aedeagus; 12, 17, 20. Inner face of paramere; 13, 21, 23. Female terminalia, in profile; 14, 22, 24. Dorsal and ventral valvulae. 10 - 14 C. daleae; 15 - 17, 21, 22. C. lienhardi sp. nov.; 18 - 20, 23, 24. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures10-24" httpUri="https://binary.pensoft.net/fig/613528" pageId="0" pageNumber="107">13</figureCitation>
. Proctiger 0.8-1.0 times as long as head width, 2.3-3.7 times as long as circumanal ring, cuneate; strongly narrowed medially, bearing an apical process, apical third serrate dorsally, subacute apically; sparsely beset with short setae, with a longitudinal lateral row of slightly longer setae in apical third and 2 longitudinal rows of peg setae near ventral margin in apical half. Subgenital plate 0.7-0.8 times as long as proctiger, in profile, bearing narrow, apically pointed process. Valvulae dorsalis and ventralis straight (Fig.
<figureCitation id="66CA04163E36A06E9AA75FF98A2D3EF0" captionStart="Figures 10–24" captionStartId="F3" captionText="Figures 10 - 24. Ctenarytaina spp., terminalia. 10, 15, 18. Male terminalia, in profile; 11, 16, 19. Distal portion of aedeagus; 12, 17, 20. Inner face of paramere; 13, 21, 23. Female terminalia, in profile; 14, 22, 24. Dorsal and ventral valvulae. 10 - 14 C. daleae; 15 - 17, 21, 22. C. lienhardi sp. nov.; 18 - 20, 23, 24. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures10-24" httpUri="https://binary.pensoft.net/fig/613528" pageId="0" pageNumber="107">14</figureCitation>
); valvula lateralis pointed apically.
</paragraph>
<paragraph id="3E7D3B53CC11AFE39AE2AF258B763A92" pageId="0" pageNumber="107">Measurements in mm (5 ♂, 6 ♀). Head width 0.44-0.58; antenna length 0.34-0.50; forewing length 1.08-1.48; length of male proctiger 0.20-0.24; paramere length 0.16-0.18; length of distal portion of aedeagus 0.10-0.14; female proctiger length 0.42-046.</paragraph>
<paragraph id="9793E0077DBEFE3A363A17590DE87FDA" pageId="0" pageNumber="107">Fifth instar immature unknown.</paragraph>
</subSubSection>
<subSubSection id="2393F5C881243C93D680BDA1E9ED1C2E" pageId="0" pageNumber="107" type="distribution">
<paragraph id="F1474B93C0FB515B1DE5C5816B431E6B" pageId="0" pageNumber="107">Distribution.</paragraph>
<paragraph id="C0B189EC0ED5108E17150D0C815F005D" pageId="0" pageNumber="107">Malaysia: Sabah, probably endemic to Gunung Kinabalu.</paragraph>
</subSubSection>
<subSubSection id="891233727B90A7E09DE552ED7FE2FBBE" pageId="0" pageNumber="107" type="host">
<paragraph id="8C7D5A3E7A80F9CC522814583E880A42" pageId="0" pageNumber="107">Host plant, biology and habitat.</paragraph>
<paragraph id="82C7FD9AE3F4D5B0388F76272447F774" pageId="0" pageNumber="107">
Adults were collected in large numbers on
<taxonomicName id="57318E4B295CB3A601F5924749F34510" authorityName="Blume" authorityYear="1827" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum javanicum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="1BC71CFB2BCFD65807BFF992AD923A7E" italics="true" pageId="0" pageNumber="107">Leptospermum javanicum</emphasis>
</taxonomicName>
Blume (=
<taxonomicName id="FEBC5288CE322A61782B40FFC9BC0A20" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="L. flavescens" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="flavescens">
<emphasis id="FF3CB76F67EA9723FC506DADBBD767BB" italics="true" pageId="0" pageNumber="107">L. flavescens</emphasis>
</taxonomicName>
auct.) and
<taxonomicName id="52C0A584C97D9C83410128358E9A2EA7" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="L. recurvum" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="recurvum">
<emphasis id="199E798BA547037C6CAC9765D83A02A8" italics="true" pageId="0" pageNumber="107">L. recurvum</emphasis>
</taxonomicName>
Hook.f. (
<taxonomicName id="B4C7B5000E45E94C9794958FB941C4EF" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) suggesting that these two species constitute hosts.
<taxonomicName id="9D7750495413E711C04E602444F7C59A" authorityName="Blume" authorityYear="1827" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum javanicum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="javanicum">
<emphasis id="D74EC668868A396067167BF405544D2F" italics="true" pageId="0" pageNumber="107">Leptospermum javanicum</emphasis>
</taxonomicName>
is widely distributed from Burma and southern Thailand to the Philippines, Malucu and Lesser Sunda Islands, and
<taxonomicName id="6DB6193D140BE1015BEBF45AFB8401BA" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="L. recurvum" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="recurvum">
<emphasis id="09DFA366E3CB2B8CAC02C81B751D86A5" italics="true" pageId="0" pageNumber="107">L. recurvum</emphasis>
</taxonomicName>
, is endemic to Gunung Kinabalu (
<bibRefCitation id="51FC1029F29628F5524777176C117742" DOI="https://doi.org/10.7751/telopea19894902" author="Thompson, J" journalOrPublisher="IntechOpen, Rijeka" pageId="0" pageNumber="107" refId="B13" refString="Thompson, J, 1989. A revision of the genus Leptospermum (Myrtaceae). Telopea 3: 301-449. https://doi.org/10.7751/telopea19894902" title="A revision of the genus Leptospermum (Myrtaceae). Telopea 3: 301 - 449." url="https://doi.org/10.7751/telopea19894902" year="1989">Thompson 1989</bibRefCitation>
). It has been suggested that
<taxonomicName id="5481081497509A848BCA4D3F28805BE0" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="L. recurvum" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="recurvum">
<emphasis id="AAB4BBDDACA8EBC3AE2E81CBEC5A9F55" italics="true" pageId="0" pageNumber="107">L. recurvum</emphasis>
</taxonomicName>
has split from the former after the last Pleistocene glaciation (
<bibRefCitation id="A5B0F82EE98669F8E08E96C3EECDE88D" DOI="https://doi.org/10.1111/j.1095-8339.1980.tb01987.x" author="Lee, DW" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="107" pagination="223 - 242" refId="B9" refString="Lee, DW, Lowry, JB, 1980. Plant speciation on tropical mountains: Leptospermum (Myrtaceae) on Mount Kinabalu, Borneo. Botanical Journal of the Linnean Society 80: 223 - 242, DOI: https://doi.org/10.1111/j.1095-8339.1980.tb01987.x" title="Plant speciation on tropical mountains: Leptospermum (Myrtaceae) on Mount Kinabalu, Borneo." url="https://doi.org/10.1111/j.1095-8339.1980.tb01987.x" volume="80" year="1980">Lee and Lowry 1980</bibRefCitation>
). The species occurs as a tree and at high altitudes as prostrate shrub on an outcrop of ultra basic rocks. It is one of the main shrub species of the summit zone above 3200 m altitude (
<bibRefCitation id="4C83CE21176771347CB363E2AB7CF50C" author="Cockburn, PF" editor="Luping, M" journalOrPublisher="Sabah Society Monograph 1978. Sabah Society, Kota Kinabalu, Sabah, Malaysia" pageId="0" pageNumber="107" pagination="179 - 190" refId="B6" refString="Cockburn, PF, 1978. Chapter 7, Flora. In: Luping, M, Wen, C, Dingley, ER, Eds., Kinabalu: Summit of Borneo. Sabah Society Monograph 1978. Sabah Society, Kota Kinabalu, Sabah, Malaysia: 179 - 190" title="Chapter 7, Flora." volumeTitle="Kinabalu: Summit of Borneo." year="1978">Cockburn 1978</bibRefCitation>
;
<bibRefCitation id="F416407A3E8384B976FFBD72F5E87D69" author="Corner, EJH" editor="Luping, M" journalOrPublisher="Sabah Society Monograph 1978. Sabah Society, Kota Kinabalu, Sabah, Malaysia" pageId="0" pageNumber="107" pagination="112 - 178" refId="B7" refString="Corner, EJH, 1978. Chapter 6, Plant life. In: Luping, M, Wen, C, Dingley, ER, Eds., Kinabalu: Summit of Borneo. Sabah Society Monograph 1978. Sabah Society, Kota Kinabalu, Sabah, Malaysia: 112 - 178" title="Chapter 6, Plant life." volumeTitle="Kinabalu: Summit of Borneo." year="1978">Corner 1978</bibRefCitation>
). A singe male was collected also on
<taxonomicName id="6F5B3AE7FF3E0D49B818EE58E3C2D3C5" class="Insecta" family="Myrtaceae" genus="Syzygium" kingdom="Animalia" lsidName="Syzygium korthalsianum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="korthalsianum">
<emphasis id="80474D93C0EEEAB5DF7659A45FBB9CF1" italics="true" pageId="0" pageNumber="107">Syzygium korthalsianum</emphasis>
</taxonomicName>
(Miq.) Miq. (
<taxonomicName id="A38C43881C43B05888EF5D13BC125736" class="Insecta" family="Myrtaceae" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="family">Myrtaceae</taxonomicName>
), which is an unlikely host contrary to the statement by
<bibRefCitation id="0FBCF15642E6B35F05974C050C2C29F1" DOI="https://doi.org/10.12976/jib/2020.20.2.1" author="Burckhardt, D" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="107" pagination="35 - 58" refId="B3" refString="Burckhardt, D, Giliomee, JH, Hamilton-Attwell, VL, Queiroz, DL, 2020. A review of jumping plant lice of the genus Ctenarytaina (Hemiptera, Psylloidea, Aphalaridae) associated with Syzygium (Myrtaceae). Journal of Insect Biodiversity 20 (2): 35 - 58, DOI: https://doi.org/10.12976/jib/2020.20.2.1" title="A review of jumping plant lice of the genus Ctenarytaina (Hemiptera, Psylloidea, Aphalaridae) associated with Syzygium (Myrtaceae)." url="https://doi.org/10.12976/jib/2020.20.2.1" volume="20" year="2020">Burckhardt et al. (2020)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="46E0EF00259C53A2ECF83A4D53F5F115" pageId="0" pageNumber="107" type="comments">
<paragraph id="1B0F581022D4851F21A036899F39159B" pageId="0" pageNumber="107">Comments.</paragraph>
<paragraph id="0FC240021A97227A9A7C916FD128B024" pageId="0" pageNumber="107">
<taxonomicName id="1700CB56D16B499DE0B67E5406CA1B4E" authorityName="Burckhardt" authorityYear="2020" class="Insecta" family="Aphalaridae" genus="Ctenarytaina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenarytaina daleae" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="daleae">
<emphasis id="264E898178F2969F70D9C02ADE91EA4B" italics="true" pageId="0" pageNumber="107">Ctenarytaina daleae</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4123FC1B9CD61FECF8072926302F8E80" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. insularis" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="insularis">
<emphasis id="3C859A65319D24777B190D50B40C8B24" italics="true" pageId="0" pageNumber="107">C. insularis</emphasis>
</taxonomicName>
Martoni &amp; Armstrong in the posteriorly weakly lobed male proctiger, the digitiform paramere, the female terminalia, which are strongly narrowed medially and bear each an apical process on the proctiger and subgenital plate as well as the dorsally serrate female proctiger. It differs from the latter in the surface spinules forming rings consisting of two rows of spinules, the distal aedeagal segment which is evenly widening to apex rather than with a slender stalk and inflated apical part, the dorsally less concave female proctiger and the host plant:
<taxonomicName id="0DFBDFE38E932185F65DA1F2E045910C" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="969D042E88D3DBC254F1E22E912112F1" italics="true" pageId="0" pageNumber="107">Leptospermum</emphasis>
</taxonomicName>
versus
<taxonomicName id="5DB4247BD876AE52C4AA668921C15323" class="Insecta" family="Myrtaceae" genus="Syzygium" kingdom="Animalia" lsidName="Syzygium" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="69E12DAD735BAE6BB681B60282DB46DF" italics="true" pageId="0" pageNumber="107">Syzygium</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="E690C2AFC03635B90E03D208848AF9EA" pageId="0" pageNumber="107">
<taxonomicName id="DF0C11783E7342A2D79CCDF24E7B5A6F" authorityName="Burckhardt" authorityYear="2020" class="Insecta" family="Aphalaridae" genus="Ctenarytaina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenarytaina daleae" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="daleae">
<emphasis id="51BDAA2C9C6DECC899B46BC2DF751879" italics="true" pageId="0" pageNumber="107">Ctenarytaina daleae</emphasis>
</taxonomicName>
was described based on the male holotype and two female paratypes (
<bibRefCitation id="2C1227224D963D5B8A8ED59569A190C7" DOI="https://doi.org/10.12976/jib/2020.20.2.1" author="Burckhardt, D" journalOrPublisher="Journal of Insect Biodiversity" pageId="0" pageNumber="107" pagination="35 - 58" refId="B3" refString="Burckhardt, D, Giliomee, JH, Hamilton-Attwell, VL, Queiroz, DL, 2020. A review of jumping plant lice of the genus Ctenarytaina (Hemiptera, Psylloidea, Aphalaridae) associated with Syzygium (Myrtaceae). Journal of Insect Biodiversity 20 (2): 35 - 58, DOI: https://doi.org/10.12976/jib/2020.20.2.1" title="A review of jumping plant lice of the genus Ctenarytaina (Hemiptera, Psylloidea, Aphalaridae) associated with Syzygium (Myrtaceae)." url="https://doi.org/10.12976/jib/2020.20.2.1" volume="20" year="2020">Burckhardt et al. 2020</bibRefCitation>
). The examination of a long series of material from the summit region of Gunung Kinabalu collected on
<taxonomicName id="92B14909D614814C5F015E01B60BB3EE" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="944F69F893D2527AEC8FCF7C52ADD418" italics="true" pageId="0" pageNumber="107">Leptospermum</emphasis>
</taxonomicName>
spp. shows that the holotype of
<taxonomicName id="74D03A8E54C7A69C9720827BD3B8D439" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. daleae" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="daleae">
<emphasis id="495C3982D4FF30E96B7759F9B0B3F581" italics="true" pageId="0" pageNumber="107">C. daleae</emphasis>
</taxonomicName>
corresponds to the species from
<taxonomicName id="0FCB77ABF9CA2F6BDC20203A792F9110" class="Insecta" family="Myrtaceae" genus="Leptospermum" kingdom="Animalia" lsidName="Leptospermum" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="31996E9DDD3753F8619CA87AD50EB6F7" italics="true" pageId="0" pageNumber="107">Leptospermum</emphasis>
</taxonomicName>
but not the females which belong to
<taxonomicName id="294BB9F90F00C13A680FEB32351A7F1C" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. smetanai" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="smetanai">
<emphasis id="62E22BEEDBE637778F7FC238C7A9D2E8" italics="true" pageId="0" pageNumber="107">C. smetanai</emphasis>
</taxonomicName>
sp. nov. described below.
<taxonomicName id="5F7DD82C042430F41BEC3D407B2E8D8D" authorityName="Burckhardt" authorityYear="2020" class="Insecta" family="Aphalaridae" genus="Ctenarytaina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenarytaina daleae" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="species" species="daleae">
<emphasis id="F995EF09211859D246B7D85B8FEAEC9D" italics="true" pageId="0" pageNumber="107">Ctenarytaina daleae</emphasis>
</taxonomicName>
and
<taxonomicName id="44BF26B90B44056A417BE6438C89E0A1" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. smetanai" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="smetanai">
<emphasis id="13074D6776BDE84BFA8A7A4B537913FE" italics="true" pageId="0" pageNumber="107">C. smetanai</emphasis>
</taxonomicName>
sp. nov. are similar in the head and forewing structure (Figs
<figureCitation id="F1E3FEB3780679EE8777F701399A6E97" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Ctenarytaina spp., head, in dorsal view. 1. C. daleae; 2. C. lienhardi sp. nov.; 3. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures1-3" httpUri="https://binary.pensoft.net/fig/613526" pageId="0" pageNumber="107">1</figureCitation>
,
<figureCitation id="8B2E70E63CAED449824EDAD5D940AB69" captionStart="Figures 1–3" captionStartId="F1" captionText="Figures 1 - 3. Ctenarytaina spp., head, in dorsal view. 1. C. daleae; 2. C. lienhardi sp. nov.; 3. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures1-3" httpUri="https://binary.pensoft.net/fig/613526" pageId="0" pageNumber="107">3</figureCitation>
,
<figureCitation id="6CDA0EB0EC0F7DDC3C7B66357EDA1747" captionStart="Figures 4–9" captionStartId="F2" captionText="Figures 4 - 9. Ctenarytaina spp. 4, 6, 8. Forewing; 5, 7, 9. Surface spinules in cell r 2 above bifurcation of vein M. 4, 5. C. daleae; 6, 7. C. lienhardi sp. nov.; 8, 9. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures4-9" httpUri="https://binary.pensoft.net/fig/613527" pageId="0" pageNumber="107">4</figureCitation>
,
<figureCitation id="059B7721C53181087DD8862E673AD050" captionStart="Figures 4–9" captionStartId="F2" captionText="Figures 4 - 9. Ctenarytaina spp. 4, 6, 8. Forewing; 5, 7, 9. Surface spinules in cell r 2 above bifurcation of vein M. 4, 5. C. daleae; 6, 7. C. lienhardi sp. nov.; 8, 9. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures4-9" httpUri="https://binary.pensoft.net/fig/613527" pageId="0" pageNumber="107">8</figureCitation>
) but differ in the male and female terminalia (Figs
<figureCitation id="A64883C70B938B44A04B4113DD05E2C6" captionStart="Figures 10–24" captionStartId="F3" captionText="Figures 10 - 24. Ctenarytaina spp., terminalia. 10, 15, 18. Male terminalia, in profile; 11, 16, 19. Distal portion of aedeagus; 12, 17, 20. Inner face of paramere; 13, 21, 23. Female terminalia, in profile; 14, 22, 24. Dorsal and ventral valvulae. 10 - 14 C. daleae; 15 - 17, 21, 22. C. lienhardi sp. nov.; 18 - 20, 23, 24. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures10-24" httpUri="https://binary.pensoft.net/fig/613528" pageId="0" pageNumber="107">10-14</figureCitation>
,
<figureCitation id="4F723198602DED10E46702018D3298C6" captionStart="Figures 10–24" captionStartId="F3" captionText="Figures 10 - 24. Ctenarytaina spp., terminalia. 10, 15, 18. Male terminalia, in profile; 11, 16, 19. Distal portion of aedeagus; 12, 17, 20. Inner face of paramere; 13, 21, 23. Female terminalia, in profile; 14, 22, 24. Dorsal and ventral valvulae. 10 - 14 C. daleae; 15 - 17, 21, 22. C. lienhardi sp. nov.; 18 - 20, 23, 24. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures10-24" httpUri="https://binary.pensoft.net/fig/613528" pageId="0" pageNumber="107">18-20</figureCitation>
,
<figureCitation id="F1FB28FF89C493C998F448EA14DAAC41" captionStart="Figures 10–24" captionStartId="F3" captionText="Figures 10 - 24. Ctenarytaina spp., terminalia. 10, 15, 18. Male terminalia, in profile; 11, 16, 19. Distal portion of aedeagus; 12, 17, 20. Inner face of paramere; 13, 21, 23. Female terminalia, in profile; 14, 22, 24. Dorsal and ventral valvulae. 10 - 14 C. daleae; 15 - 17, 21, 22. C. lienhardi sp. nov.; 18 - 20, 23, 24. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures10-24" httpUri="https://binary.pensoft.net/fig/613528" pageId="0" pageNumber="107">23</figureCitation>
,
<figureCitation id="670241393FA4C80A1BB282EB3A8DE6A0" captionStart="Figures 10–24" captionStartId="F3" captionText="Figures 10 - 24. Ctenarytaina spp., terminalia. 10, 15, 18. Male terminalia, in profile; 11, 16, 19. Distal portion of aedeagus; 12, 17, 20. Inner face of paramere; 13, 21, 23. Female terminalia, in profile; 14, 22, 24. Dorsal and ventral valvulae. 10 - 14 C. daleae; 15 - 17, 21, 22. C. lienhardi sp. nov.; 18 - 20, 23, 24. C. smetanai sp. nov." figureDoi="10.3897/alpento.5.77751.figures10-24" httpUri="https://binary.pensoft.net/fig/613528" pageId="0" pageNumber="107">24</figureCitation>
).
</paragraph>
<caption id="EAC1CA2C23DA5C7D7A58BFC421F85B68" doi="10.3897/alpento.5.77751.figures1-3" httpUri="https://binary.pensoft.net/fig/613526" pageId="0" pageNumber="107" start="Figures 1–3" startId="F1">
<paragraph id="E98B04491A0B66846402CBEF99971881" pageId="0" pageNumber="107">
<emphasis id="AFBC7B18A9696C56F98E4E30B3EE3E69" bold="true" pageId="0" pageNumber="107">Figures 1-3.</emphasis>
<taxonomicName id="02F90142F5A468B78CAD8E149B08A722" authorityName="Ferris &amp; Klyver" authorityYear="1932" class="Insecta" family="Aphalaridae" genus="Ctenarytaina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenarytaina" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="461C69F2C877A1C57984BAB94A49D611" italics="true" pageId="0" pageNumber="107">Ctenarytaina</emphasis>
</taxonomicName>
spp., head, in dorsal view.
<emphasis id="2FFE8F407B2843F44A46F1750C8AFB05" bold="true" pageId="0" pageNumber="107">1.</emphasis>
<taxonomicName id="35995CB8F2BE23E4B74EA84C2C9AF02F" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. daleae" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="daleae">
<emphasis id="DD696A269234E5A79EAAE68B8D772A93" italics="true" pageId="0" pageNumber="107">C. daleae</emphasis>
</taxonomicName>
;
<emphasis id="9C5E222A89211BC598C42C54C61898E8" bold="true" pageId="0" pageNumber="107">2.</emphasis>
<taxonomicName id="859D3DCFD6D23C3F983CA247081E5027" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. lienhardi" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="lienhardi">
<emphasis id="36ED364BB170DE3C44BF31E7C9CC2728" italics="true" pageId="0" pageNumber="107">C. lienhardi</emphasis>
</taxonomicName>
sp. nov.;
<emphasis id="2EB51CCC981EEE405392A9F3700A064C" bold="true" pageId="0" pageNumber="107">3.</emphasis>
<taxonomicName id="A0D7BC604031FEFB4A60F79D30DDFDCF" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. smetanai" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="smetanai">
<emphasis id="62A51CCC1D1A1DD83F3B6B6C1EF188C4" italics="true" pageId="0" pageNumber="107">C. smetanai</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
<caption id="6464F555613AAD5A07992BB16777964E" doi="10.3897/alpento.5.77751.figures4-9" httpUri="https://binary.pensoft.net/fig/613527" pageId="0" pageNumber="107" start="Figures 4–9" startId="F2">
<paragraph id="4D69B95E8D6651D1DE0FDDA97EAA48EB" pageId="0" pageNumber="107">
<emphasis id="8A93A6E6817FDC30BA76B668129BA9A8" bold="true" pageId="0" pageNumber="107">Figures 4-9.</emphasis>
<taxonomicName id="3C9FF39672E6F404BD77BE8DA41F1B18" authorityName="Ferris &amp; Klyver" authorityYear="1932" class="Insecta" family="Aphalaridae" genus="Ctenarytaina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ctenarytaina" order="Hemiptera" pageId="0" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="9969F4718CD2EEEDAE8892B7083A3F99" italics="true" pageId="0" pageNumber="107">Ctenarytaina</emphasis>
</taxonomicName>
spp.
<emphasis id="676FBC4660CCD1F2A6424FA1740B69A5" bold="true" pageId="0" pageNumber="107">4, 6, 8.</emphasis>
Forewing;
<emphasis id="12DC939D443431B013590953EBF38124" bold="true" pageId="0" pageNumber="107">5, 7, 9.</emphasis>
Surface spinules in cell r2 above bifurcation of vein M.
<emphasis id="9D9CDDF75AD2A16E3FB2A5EDF382EDE1" bold="true" pageId="0" pageNumber="107">4, 5.</emphasis>
<taxonomicName id="733C868A50E0EB053F6155AD2B149014" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. daleae" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="daleae">
<emphasis id="2F6B316919D00CC61CB1BB14AF544E97" italics="true" pageId="0" pageNumber="107">C. daleae</emphasis>
</taxonomicName>
;
<emphasis id="64ADD06A75C1A9FA88D66392487BFC28" bold="true" pageId="0" pageNumber="107">6, 7.</emphasis>
<taxonomicName id="663571532EDA3460BFEACCCAEBE4719E" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. lienhardi" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="lienhardi">
<emphasis id="25CC59445C224594446046F8C0E2E9D4" italics="true" pageId="0" pageNumber="107">C. lienhardi</emphasis>
</taxonomicName>
sp. nov.;
<emphasis id="43738AF37BF60FD0BC7F2BC714B89C58" bold="true" pageId="0" pageNumber="107">8, 9.</emphasis>
<taxonomicName id="64CC6678626F5A9956064A299D380D14" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" lsidName="C. smetanai" order="Myrtales" pageId="0" pageNumber="107" phylum="Tracheophyta" rank="species" species="smetanai">
<emphasis id="228CC54F5B9ED9C2727D202330424AC5" italics="true" pageId="0" pageNumber="107">C. smetanai</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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