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<mods:title id="B74679C3ED52DB749E81452C0BFDA2EA">Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species</mods:title>
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<treatment id="A703878CA948FFBDFF26FE2D9B0D0AE2" ID-DOI="http://doi.org/10.5281/zenodo.4340400" ID-GBIF-Taxon="167366584" ID-Zenodo-Dep="4340400" LSID="urn:lsid:plazi:treatment:A703878CA948FFBDFF26FE2D9B0D0AE2" httpUri="http://treatment.plazi.org/id/A703878CA948FFBDFF26FE2D9B0D0AE2" lastPageNumber="8" pageId="7" pageNumber="8">
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<paragraph id="2F15369AA948FFBDFF26FE2D99E00DD2" blockId="7.[151,908,441,468]" box="[151,908,441,468]" pageId="7" pageNumber="8">
<heading id="745D81F6A948FFBDFF26FE2D99E00DD2" bold="true" box="[151,908,441,468]" fontSize="11" level="2" pageId="7" pageNumber="8" reason="4">
<emphasis id="1DDEEA88A948FFBDFF26FE2D99E00DD2" bold="true" box="[151,908,441,468]" pageId="7" pageNumber="8">
<taxonomicName id="E8AA4D19A948FFBDFF26FE2D98D30DD2" ID-CoL="8CSXB" authority="(Sasakawa, 1997)" authorityName="Vilkamaa &amp; Menzel" authorityYear="2019" baseAuthorityName="Sasakawa" baseAuthorityYear="1997" box="[151,703,441,468]" class="Insecta" family="Sciaridae" genus="Merizomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora">
<emphasis id="1DDEEA88A948FFBDFF26FE2D9B8C0DD5" bold="true" box="[151,480,441,468]" italics="true" pageId="7" pageNumber="8">Merizomma codonopsivora</emphasis>
(
<bibRefCitation id="4B3B4B6BA948FFBDFE5EFE2D98DB0DD2" author="Sasakawa, M." box="[495,695,441,467]" pageId="7" pageNumber="8" pagination="171 - 176" refId="ref45374" refString="Sasakawa, M. (1997) New leaf miner and stem borer of Sciaridae (Diptera). Japanese Journal of Entomology, 65 (1), 171 - 176." type="journal article" year="1997">Sasakawa, 1997</bibRefCitation>
)
</taxonomicName>
stat. et comb. n.
</emphasis>
</heading>
</paragraph>
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<subSubSection id="67B06511A948FFBDFF26FD9498AC0E62" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="2F15369AA948FFBDFF26FD9498AC0E62" blockId="7.[151,1437,512,1763]" pageId="7" pageNumber="8">
<emphasis id="1DDEEA88A948FFBDFF26FD949B7F0E1A" bold="true" box="[151,275,512,539]" pageId="7" pageNumber="8">Literature</emphasis>
.
<taxonomicName id="E8AA4D19A948FFBDFE94FD959E0B0E1A" ID-CoL="899G7" authority="Sasakawa-Sasakawa (1997)" authorityName="Sasakawa-Sasakawa" authorityYear="1997" box="[293,1127,513,539]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora" subGenus="Chorizomma">
<emphasis id="1DDEEA88A948FFBDFE94FD959BF60E1A" box="[293,410,513,539]" italics="true" pageId="7" pageNumber="8">Lycoriella</emphasis>
(
<emphasis id="1DDEEA88A948FFBDFE1EFD95982E0E1A" box="[431,578,513,539]" italics="true" pageId="7" pageNumber="8">Chorizomma</emphasis>
)
<emphasis id="1DDEEA88A948FFBDFDE9FD95996E0E1A" box="[600,770,513,539]" italics="true" pageId="7" pageNumber="8">codonopsivora</emphasis>
Sasakawa—Sasakawa (1997)
</taxonomicName>
: 171, figs 13.
<taxonomicName id="E8AA4D19A948FFBDFA96FD95992D0E3E" ID-CoL="899G7" authority="Sasakawa-Sasakawa (2003)" authorityName="Sasakawa-Sasakawa" authorityYear="2003" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora" subGenus="Merizomma">
<emphasis id="1DDEEA88A948FFBDFA96FD959FF00E1A" box="[1319,1436,513,539]" italics="true" pageId="7" pageNumber="8">Lycoriella</emphasis>
(
<emphasis id="1DDEEA88A948FFBDFF10FDB29B450E3E" box="[161,297,550,575]" italics="true" pageId="7" pageNumber="8">Merizomma</emphasis>
)
<emphasis id="1DDEEA88A948FFBDFE8BFDB19B880E3E" box="[314,484,549,575]" italics="true" pageId="7" pageNumber="8">codonopsivora</emphasis>
Sasakawa—Sasakawa (2003)
</taxonomicName>
: 119, 128.
<taxonomicName id="E8AA4D19A948FFBDFC0DFDB19B280E62" ID-CoL="899G7" authority="Sasakawa-Sasakawa (2008)" authorityName="Sasakawa-Sasakawa" authorityYear="2008" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora">
<emphasis id="1DDEEA88A948FFBDFC0DFDB19E8E0E3E" box="[956,1250,549,575]" italics="true" pageId="7" pageNumber="8">Lycoriella codonopsivora</emphasis>
Sasakawa—Sa-sakawa (2008)
</taxonomicName>
: 128;
<bibRefCitation id="4B3B4B6BA948FFBDFE39FDDD98EF0E65" author="Eiseman, C. S. &amp; Heller, K. &amp; Rulik, B." box="[392,643,585,612]" pageId="7" pageNumber="8" pagination="519 - 532" refId="ref39995" refString="Eiseman, C. S., Heller, K. &amp; Rulik, B. (2016) A new leaf-mining Dark-Winged Fungus Gnat (Diptera: Sciaridae), with notes on other insect associates of marsh marigold (Ranunculaceae: Caltha palustris L.). Proceedings of the Entomological Society of Washington, 118 (4), 519 - 532. https: // doi. org / 10.4289 / 0013 - 8797.118.4.519" type="journal article" year="2016">
Eiseman
<emphasis id="1DDEEA88A948FFBDFE43FDDE98400E62" box="[498,556,585,611]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2016)
</bibRefCitation>
: 527.
</paragraph>
</subSubSection>
<subSubSection id="67B06511A948FFBDFF76FDF8994D0F5E" pageId="7" pageNumber="8" type="discussion">
<paragraph id="2F15369AA948FFBDFF76FDF8994D0F5E" blockId="7.[151,1437,512,1763]" pageId="7" pageNumber="8">
<emphasis id="1DDEEA88A948FFBDFF76FDF89B290E86" bold="true" box="[199,325,620,647]" pageId="7" pageNumber="8">Discussion</emphasis>
.
<bibRefCitation id="4B3B4B6BA948FFBDFEE7FDF9983F0E86" author="Sasakawa, M." box="[342,595,621,647]" pageId="7" pageNumber="8" pagination="171 - 176" refId="ref45374" refString="Sasakawa, M. (1997) New leaf miner and stem borer of Sciaridae (Diptera). Japanese Journal of Entomology, 65 (1), 171 - 176." type="journal article" year="1997">Sasakawa (1997: 171)</bibRefCitation>
described the distinctive
<taxonomicName id="E8AA4D19A948FFBDFCCBFDF99F300E86" authority="Sasakawa, 1997" authorityName="Sasakawa-Sasakawa" authorityYear="1997" box="[890,1372,621,647]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora">
<emphasis id="1DDEEA88A948FFBDFCCBFDF99EF00E86" box="[890,1180,621,647]" italics="true" pageId="7" pageNumber="8">Lycoriella codonopsivora</emphasis>
<bibRefCitation id="4B3B4B6BA948FFBDFB17FDF99F300E86" author="Sasakawa, M." box="[1190,1372,621,647]" pageId="7" pageNumber="8" pagination="171 - 176" refId="ref45374" refString="Sasakawa, M. (1997) New leaf miner and stem borer of Sciaridae (Diptera). Japanese Journal of Entomology, 65 (1), 171 - 176." type="journal article" year="1997">Sasakawa, 1997</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="ED6EF878A948FFBDFF26FD059B690EAA" box="[151,261,657,683]" country="Japan" name="Hokkaido" pageId="7" pageNumber="8">Hokkaido</collectingRegion>
(
<collectingCountry id="57BD760AA948FFBDFEA5FD059B3B0EAA" box="[276,343,657,683]" name="Japan" pageId="7" pageNumber="8">Japan</collectingCountry>
) whose larvae feed on deodeok leaves (
<taxonomicName id="E8AA4D19A948FFBDFCA3FD059E7E0EAA" box="[786,1042,657,683]" class="Magnoliopsida" family="Campanulaceae" genus="Codonopsis" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="species" species="lanceolata">
<emphasis id="1DDEEA88A948FFBDFCA3FD059E7E0EAA" box="[786,1042,657,683]" italics="true" pageId="7" pageNumber="8">Codonopsis lanceolata</emphasis>
</taxonomicName>
,
<taxonomicName id="E8AA4D19A948FFBDFB91FD059EBB0EAD" box="[1056,1239,657,684]" class="Magnoliopsida" family="Campanulaceae" kingdom="Plantae" order="Asterales" pageId="7" pageNumber="8" phylum="Tracheophyta" rank="family">Campanulaceae</taxonomicName>
). For this species, he erected the monotypic subgenus
<taxonomicName id="E8AA4D19A948FFBDFD89FD2198A50ECE" box="[568,713,693,719]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Chorizomma">
<emphasis id="1DDEEA88A948FFBDFD89FD2198A50ECE" box="[568,713,693,719]" italics="true" pageId="7" pageNumber="8">Chorizomma</emphasis>
</taxonomicName>
of
<taxonomicName id="E8AA4D19A948FFBDFD45FD219E0B0ECE" authority="Sasakawa, 1997" box="[756,1127,693,719]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFD45FD21990A0ECE" box="[756,870,693,719]" italics="true" pageId="7" pageNumber="8">Lycoriella</emphasis>
(
<bibRefCitation id="4B3B4B6BA948FFBDFCC8FD219E0B0ECE" author="Sasakawa, M." box="[889,1127,693,719]" pageId="7" pageNumber="8" pagination="171 - 176" refId="ref45374" refString="Sasakawa, M. (1997) New leaf miner and stem borer of Sciaridae (Diptera). Japanese Journal of Entomology, 65 (1), 171 - 176." type="journal article" year="1997">Sasakawa 1997: 174</bibRefCitation>
</taxonomicName>
[preocc., not
<taxonomicName id="E8AA4D19A948FFBDFABDFD219B450EF2" authority="Simon, 1872" authorityName="Simon" authorityYear="1872" class="Insecta" family="Sciaridae" genus="Chorizomma" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFABDFD219FF10ECE" box="[1292,1437,693,719]" italics="true" pageId="7" pageNumber="8">Chorizomma</emphasis>
Simon, 1872
</taxonomicName>
;
<taxonomicName id="E8AA4D19A948FFBDFE83FD4D9BFE0EF2" box="[306,402,729,755]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="order">Araneae</taxonomicName>
,
<taxonomicName id="E8AA4D19A948FFBDFE2CFD4C98770EF2" box="[413,539,728,755]" class="Arachnida" family="Dictynidae" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Dictynidae</taxonomicName>
] and renamed the subgenus later as
<taxonomicName id="E8AA4D19A948FFBDFC1BFD4E9E5C0EF2" authorityName="Sasakawa" authorityYear="2003" box="[938,1072,730,755]" class="Insecta" family="Sciaridae" genus="Merizomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFC1BFD4E9E5C0EF2" box="[938,1072,730,755]" italics="true" pageId="7" pageNumber="8">Merizomma</emphasis>
</taxonomicName>
(
<bibRefCitation id="4B3B4B6BA948FFBDFB8FFD4D9F490EF2" author="Sasakawa, M." box="[1086,1317,729,755]" pageId="7" pageNumber="8" pagination="119 - 133" refId="ref45407" refString="Sasakawa, M. (2003) Notes on the Japanese Diptera Part 2. Japanese Journal of Entomology, new series, 6 (3), 119 - 133." type="journal article" year="2003">Sasakawa 2003: 119</bibRefCitation>
, 128). The subgenus and the
<typeStatus id="F0118838A948FFBDFED2FD699BFF0F16" box="[355,403,765,791]" pageId="7" pageNumber="8">type</typeStatus>
species are not included in the monograph of the Palaearctic
<taxonomicName id="E8AA4D19A948FFBDFBE5FD699AB00F3A" authority="(Menzel &amp; Mohrig 2000)" baseAuthorityName="Menzel &amp; Mohrig" baseAuthorityYear="2000" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">
Sciaridae (
<bibRefCitation id="4B3B4B6BA948FFBDFB7DFD699AB80F3A" author="Menzel, F. &amp; Mohrig, W." pageId="7" pageNumber="8" refId="ref43072" refString="Menzel, F. &amp; Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement, 6 (1999). Ampyx-Verlag, Halle (Saale), 761 pp." type="book" year="2000">Menzel &amp; Mohrig 2000</bibRefCitation>
)
</taxonomicName>
. Therefore, these taxa are discussed here in detail on the basis of the original descriptions by
<bibRefCitation id="4B3B4B6BA948FFBDFB6CFCB59FF00F3A" author="Sasakawa, M." box="[1245,1436,801,827]" pageId="7" pageNumber="8" pagination="171 - 176" refId="ref45374" refString="Sasakawa, M. (1997) New leaf miner and stem borer of Sciaridae (Diptera). Japanese Journal of Entomology, 65 (1), 171 - 176." type="journal article" year="1997">Sasakawa (1997)</bibRefCitation>
and included in the identification key and in the checklist.
</paragraph>
</subSubSection>
<subSubSection id="67B06511A948FFBDFF26FCFD9E1108A2" pageId="7" pageNumber="8" type="reference_group">
<paragraph id="2F15369AA948FFBDFF26FCFD9F750FA6" blockId="7.[151,1437,512,1763]" pageId="7" pageNumber="8">
<treatmentCitationGroup id="0FBA11B4A948FFBDFF26FCFD9F750FA6" pageId="7" pageNumber="8">
<taxonomicName id="E8AA4D19A948FFBDFF26FCFD98150F82" ID-CoL="899G7" authority="Sasakawa, 1997" authorityName="Sasakawa-Sasakawa" authorityYear="1997" box="[151,633,873,899]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora">
<emphasis id="1DDEEA88A948FFBDFF26FCFD9BD50F82" box="[151,441,873,899]" italics="true" pageId="7" pageNumber="8">Lycoriella codonopsivora</emphasis>
<bibRefCitation id="4B3B4B6BA948FFBDFE73FCFD98150F82" author="Sasakawa, M." box="[450,633,873,899]" pageId="7" pageNumber="8" pagination="171 - 176" refId="ref45374" refString="Sasakawa, M. (1997) New leaf miner and stem borer of Sciaridae (Diptera). Japanese Journal of Entomology, 65 (1), 171 - 176." type="journal article" year="1997">Sasakawa, 1997</bibRefCitation>
</taxonomicName>
has characters which are not present in this combination in any other genus and which distinctly differ from those given to
<emphasis id="1DDEEA88A948FFBDFD41FC1999FD0FA6" box="[752,913,909,935]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="E8AA4D19A948FFBDFD41FC19990E0FA6" ID-CoL="5HLM" authority="Sasakawa, 1997" box="[752,866,909,935]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus" sensu="lato">Lycoriella</taxonomicName>
<taxonomicNameLabel id="06ED57F3A948FFBDFCD8FC1A99FD0FA6" box="[873,913,909,935]" pageId="7" pageNumber="8" sensu="lato">s. l.</taxonomicNameLabel>
</emphasis>
by
<treatmentCitation id="AE0B108BA948FFBDFC0AFC199F790FA6" author="Menzel, F. &amp; Mohrig, W." box="[955,1301,909,935]" page="377" pageId="7" pageNumber="8" year="2000">
<bibRefCitation id="4B3B4B6BA948FFBDFC0AFC199F790FA6" author="Menzel, F. &amp; Mohrig, W." box="[955,1301,909,935]" pageId="7" pageNumber="8" refId="ref43072" refString="Menzel, F. &amp; Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement, 6 (1999). Ampyx-Verlag, Halle (Saale), 761 pp." type="book" year="2000">Menzel &amp; Mohrig (2000: 377)</bibRefCitation>
</treatmentCitation>
:
</treatmentCitationGroup>
</paragraph>
<paragraph id="2F15369AA948FFBDFF76FC259E1108A2" blockId="7.[151,1437,512,1763]" pageId="7" pageNumber="8">
Eye bridge not united at middle, without ommatidia; maxillary palpus 2-segmented, without pit of sensilla on 1
<superScript id="D8DF9BD2A948FFBDFF14FC409ADC0FE3" attach="left" box="[165,176,980,994]" fontSize="6" pageId="7" pageNumber="8">st</superScript>
segment; scutellum without long lateral setae, only with two unregular rows of short setae; R
<subScript id="B32E34DFA948FFBDFB56FC729E830FF5" attach="left" box="[1255,1263,998,1012]" fontSize="6" pageId="7" pageNumber="8">1</subScript>
longer than R, ending beyond the base of M-fork; fore tibia with small, bow-like bordered tibial organ; tarsal claws untoothed; hypopygium with wide gonocoxa; gonocoxae united in v-form and with short setae, intergonocoxal area not modified, gonostylus elongate-triangular and with apical megaseta, without apical tooth; medial side of gonostylus without megasetae and at apical half strongly impressed; basal part [at 1/5 of the medial side of gonostylus] with long, upcurved seta on long basal body; tegmen roundish, without apical or lateral modifications.
</paragraph>
</subSubSection>
<subSubSection id="67B06511A948FFBDFF76FB399B0D0AE2" pageId="7" pageNumber="8" type="description">
<paragraph id="2F15369AA948FFBDFF76FB399B0D0AE2" blockId="7.[151,1437,512,1763]" pageId="7" pageNumber="8">
On the basis of the above characters,
<taxonomicName id="E8AA4D19A948FFBDFDD8FB3A99CE08C6" authority="Sasakawa" authorityName="Sasakawa-Sasakawa" authorityYear="1997" box="[617,930,1197,1223]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora">
<emphasis id="1DDEEA88A948FFBDFDD8FB3A994108C6" box="[617,813,1197,1223]" italics="true" pageId="7" pageNumber="8">L. codonopsivora</emphasis>
Sasakawa
</taxonomicName>
must be excluded from the
<taxonomicName id="E8AA4D19A948FFBDFB6AFB399F2108C6" box="[1243,1357,1197,1223]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFB6AFB399F2108C6" box="[1243,1357,1197,1223]" italics="true" pageId="7" pageNumber="8">Lycoriella</emphasis>
</taxonomicName>
group. With its short gonocoxa, lacking intergonocoxal lobe and apical megaseta, the species resembles some species of the
<taxonomicName id="E8AA4D19A948FFBDFF73FB629B36090E" authorityName="Mohrig" authorityYear="1987" box="[194,346,1269,1295]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nepalensis">
<emphasis id="1DDEEA88A948FFBDFF73FB629B36090E" box="[194,346,1269,1295]" italics="true" pageId="7" pageNumber="8">K. nepalensis</emphasis>
</taxonomicName>
group of
<taxonomicName id="E8AA4D19A948FFBDFE7FFB619898090E" authority="Mohrig, 1987" authorityName="Mohrig" authorityYear="1987" box="[462,756,1269,1295]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFE7FFB619827090E" box="[462,587,1269,1295]" italics="true" pageId="7" pageNumber="8">Keilbachia</emphasis>
<bibRefCitation id="4B3B4B6BA948FFBDFDE4FB619898090E" author="Mohrig, W. &amp; Mamaev, B. &amp; Krivosheina, N." box="[597,756,1269,1295]" pageId="7" pageNumber="8" pagination="91 - 104" refId="ref44180" refString="Mohrig, W., Mamaev, B. &amp; Krivosheina, N. (1987) Beitrage zur Kenntnis der Trauermucken der Sowjetunion (Diptera, Sciaridae). Teil XI. Neue Arten aus der zentral-asiatischen autonomen Sowjetrepublik Tuwa. Zoologische Jahrbucher, Abteilung fur Systematik, Okologie und Geographie der Tiere, 114 (1), 91 - 104." type="journal article" year="1987">Mohrig, 1987</bibRefCitation>
</taxonomicName>
[compare here
<bibRefCitation id="4B3B4B6BA948FFBDFC1DFB619F76090E" author="Menzel, F. &amp; Martens, J." box="[940,1306,1268,1295]" pageId="7" pageNumber="8" pagination="97 - 129" refId="ref42863" refString="Menzel, F. &amp; Martens, J. (1995) Die Sciaridae (Diptera, Nematocera) des Nepal-Himalaya. Teil l. Die blutenbesuchenden Trauermucken an Aronstabgewuchsen der Gattung Arisaema (Araceae Juss.). Studia dipterologica, 2 (1), 97 - 129." type="journal article" year="1995">Menzel &amp; Martens (1995: 107)</bibRefCitation>
,
<bibRefCitation id="4B3B4B6BA948FFBDFA99FB619B120932" author="Menzel, F. &amp; Mohrig, W." pageId="7" pageNumber="8" refId="ref43072" refString="Menzel, F. &amp; Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement, 6 (1999). Ampyx-Verlag, Halle (Saale), 761 pp." type="book" year="2000">Menzel &amp; Mohrig (2000: 348)</bibRefCitation>
,
<bibRefCitation id="4B3B4B6BA948FFBDFE39FA8D98DA0932" author="Vilkamaa, P. &amp; Komarova, L. A. &amp; Hippa, H." box="[392,694,1305,1331]" pageId="7" pageNumber="8" pagination="39 - 55" refId="ref47204" refString="Vilkamaa, P., Komarova, L. A. &amp; Hippa, H. (2006) The genus Keilbachia Mohrig (Diptera: Sciaridae) in a biodiversity hot spot: new sympatric species from Kambaiti, Burma. Zootaxa, 1123, 39 - 55. https: // doi. org / 10.11646 / zootaxa. 1123.1.3" type="journal article" year="2006">
Vilkamaa
<emphasis id="1DDEEA88A948FFBDFE4CFA8E985A0932" box="[509,566,1305,1331]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2006: 40)
</bibRefCitation>
and
<bibRefCitation id="4B3B4B6BA948FFBDFD5CFA8D9E7C0932" author="Vilkamaa, P. &amp; Menzel, F. &amp; Hippa, H." box="[749,1040,1304,1331]" pageId="7" pageNumber="8" pagination="1 - 20" refId="ref47266" refString="Vilkamaa, P., Menzel, F. &amp; Hippa, H. (2009) Review of the genus Keilbachia Mohrig (Diptera: Sciaridae), with the description of eleven new species. Zootaxa, 2272, 1 - 20." type="journal article" year="2009">
Vilkamaa
<emphasis id="1DDEEA88A948FFBDFCD0FA8E99F70932" box="[865,923,1305,1331]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2009: 3)
</bibRefCitation>
]. However, a strongly reduced eye bridge, a 2-segmented palpus, an unbordered patch of sensilla on 1
<superScript id="D8DF9BD2A948FFBDFC22FAA899F2094B" attach="left" box="[915,926,1340,1354]" fontSize="6" pageId="7" pageNumber="8">st</superScript>
segment of palpus, a long R
<subScript id="B32E34DFA948FFBDFB55FADA9E80095D" attach="left" box="[1252,1260,1358,1372]" fontSize="6" pageId="7" pageNumber="8">1</subScript>
and the lack of a spiral-formed megaseta on the medial side of gonostylus differ from the characters of
<taxonomicName id="E8AA4D19A948FFBDFBC9FAF59E9F097A" authorityName="Mohrig" authorityYear="1987" box="[1144,1267,1377,1403]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFBC9FAF59E9F097A" box="[1144,1267,1377,1403]" italics="true" pageId="7" pageNumber="8">Keilbachia</emphasis>
</taxonomicName>
. These characters are typical for the
<taxonomicName id="E8AA4D19A948FFBDFE26FA129869099E" box="[407,517,1413,1439]" class="Insecta" family="Sciaridae" genus="Merizomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hippai">
<emphasis id="1DDEEA88A948FFBDFE26FA129869099E" box="[407,517,1413,1439]" italics="true" pageId="7" pageNumber="8">M. hippai</emphasis>
</taxonomicName>
group of the genus
<taxonomicName id="E8AA4D19A948FFBDFD57FA119999099E" authority="Menzel, 1995" authorityName="Menzel" authorityYear="1995" box="[742,1013,1412,1439]" class="Insecta" family="Sciaridae" genus="Mohrigia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFD57FA11993C099E" box="[742,848,1413,1439]" italics="true" pageId="7" pageNumber="8">Mohrigia</emphasis>
<bibRefCitation id="4B3B4B6BA948FFBDFCE9FA119999099E" author="Menzel, F. &amp; Martens, J." box="[856,1013,1412,1439]" pageId="7" pageNumber="8" pagination="97 - 129" refId="ref42863" refString="Menzel, F. &amp; Martens, J. (1995) Die Sciaridae (Diptera, Nematocera) des Nepal-Himalaya. Teil l. Die blutenbesuchenden Trauermucken an Aronstabgewuchsen der Gattung Arisaema (Araceae Juss.). Studia dipterologica, 2 (1), 97 - 129." type="journal article" year="1995">Menzel, 1995</bibRefCitation>
</taxonomicName>
. Speaking against the combination of
<taxonomicName id="E8AA4D19A948FFBDFF26FA3E9BBC09C2" authority="Sasakawa" authorityName="Sasakawa-Sasakawa" authorityYear="1997" box="[151,464,1449,1475]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora">
<emphasis id="1DDEEA88A948FFBDFF26FA3E9B3609C2" box="[151,346,1449,1475]" italics="true" pageId="7" pageNumber="8">L. codonopsivora</emphasis>
Sasakawa
</taxonomicName>
with
<taxonomicName id="E8AA4D19A948FFBDFDBEFA3D981509C2" authorityName="Menzel" authorityYear="1995" box="[527,633,1449,1475]" class="Insecta" family="Sciaridae" genus="Mohrigia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFDBEFA3D981509C2" box="[527,633,1449,1475]" italics="true" pageId="7" pageNumber="8">Mohrigia</emphasis>
</taxonomicName>
are the
<taxonomicName id="E8AA4D19A948FFBDFD60FA3D992009C2" authorityName="Mohrig" authorityYear="1987" box="[721,844,1449,1475]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFD60FA3D992009C2" box="[721,844,1449,1475]" italics="true" pageId="7" pageNumber="8">Keilbachia</emphasis>
</taxonomicName>
-like form of the gonostylus, the missing long medial setae on the apical half of the gonostylus, the missing intergonocoxal lobe of hypopygium and the much smaller, weakly setose fore tibial organ [compare here
<bibRefCitation id="4B3B4B6BA948FFBDFD12FA659E6A0A0A" author="Menzel, F. &amp; Martens, J." box="[675,1030,1520,1547]" pageId="7" pageNumber="8" pagination="97 - 129" refId="ref42863" refString="Menzel, F. &amp; Martens, J. (1995) Die Sciaridae (Diptera, Nematocera) des Nepal-Himalaya. Teil l. Die blutenbesuchenden Trauermucken an Aronstabgewuchsen der Gattung Arisaema (Araceae Juss.). Studia dipterologica, 2 (1), 97 - 129." type="journal article" year="1995">Menzel &amp; Martens (1995: 102)</bibRefCitation>
,
<bibRefCitation id="4B3B4B6BA948FFBDFBA2FA659F000A0A" author="Menzel, F. &amp; Mohrig, W." box="[1043,1388,1520,1547]" pageId="7" pageNumber="8" refId="ref43072" refString="Menzel, F. &amp; Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement, 6 (1999). Ampyx-Verlag, Halle (Saale), 761 pp." type="book" year="2000">Menzel &amp; Mohrig (2000: 415)</bibRefCitation>
and
<bibRefCitation id="4B3B4B6BA948FFBDFF26F9819BF60A2E" author="Rudzinski, H. - G." box="[151,410,1556,1583]" pageId="7" pageNumber="8" pagination="449 - 476" refId="ref45124" refString="Rudzinski, H. - G. (2006) Beitrage zur Trauermuckenfauna Taiwans. Teil IV: Gattungen Lycoriella, Mohrigia, Chaetosciara, Scythropochoa und Pseudoaerumnosa gen. nov. (Diptera Nematocera: Sciaridae). Entomofauna, 27 (37), 449 - 476." type="journal article" year="2006">Rudzinski (2006: 450)</bibRefCitation>
]. Furthermore, neither
<taxonomicName id="E8AA4D19A948FFBDFD2DF981997B0A2E" authorityName="Mohrig" authorityYear="1987" box="[668,791,1557,1583]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFD2DF981997B0A2E" box="[668,791,1557,1583]" italics="true" pageId="7" pageNumber="8">Keilbachia</emphasis>
</taxonomicName>
nor
<taxonomicName id="E8AA4D19A948FFBDFCFDF98199DA0A2E" authorityName="Menzel" authorityYear="1995" box="[844,950,1557,1583]" class="Insecta" family="Sciaridae" genus="Mohrigia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFCFDF98199DA0A2E" box="[844,950,1557,1583]" italics="true" pageId="7" pageNumber="8">Mohrigia</emphasis>
</taxonomicName>
have the lateral setae of scutellum reduced [both have two long and strong setae] nor an unmodified tegmen [the tegmen of
<taxonomicName id="E8AA4D19A948FFBDFB9BF9AD9EF80A52" authorityName="Menzel" authorityYear="1995" box="[1066,1172,1593,1619]" class="Insecta" family="Sciaridae" genus="Mohrigia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFB9BF9AD9EF80A52" box="[1066,1172,1593,1619]" italics="true" pageId="7" pageNumber="8">Mohrigia</emphasis>
</taxonomicName>
has a straight, strongly sclerotized dorsomedial structure; the tegmen of
<taxonomicName id="E8AA4D19A948FFBDFD71F9C999570A76" authorityName="Mohrig" authorityYear="1987" box="[704,827,1629,1655]" class="Insecta" family="Sciaridae" genus="Keilbachia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFD71F9C999570A76" box="[704,827,1629,1655]" italics="true" pageId="7" pageNumber="8">Keilbachia</emphasis>
</taxonomicName>
is sclerotized laterally and often with a narrow apical process]. On the above grounds,
<taxonomicName id="E8AA4D19A948FFBDFDBDF91699230A9A" authority="Sasakawa, 2003" authorityName="Sasakawa" authorityYear="2003" box="[524,847,1665,1691]" class="Insecta" family="Sciaridae" genus="Merizomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFDBDF91698FE0A9A" box="[524,658,1666,1691]" italics="true" pageId="7" pageNumber="8">Merizomma</emphasis>
<bibRefCitation id="4B3B4B6BA948FFBDFD28F91599230A9A" author="Sasakawa, M." box="[665,847,1665,1691]" pageId="7" pageNumber="8" pagination="119 - 133" refId="ref45407" refString="Sasakawa, M. (2003) Notes on the Japanese Diptera Part 2. Japanese Journal of Entomology, new series, 6 (3), 119 - 133." type="journal article" year="2003">Sasakawa, 2003</bibRefCitation>
</taxonomicName>
<emphasis id="1DDEEA88A948FFBDFCE6F91599C40A9A" bold="true" box="[855,936,1665,1691]" pageId="7" pageNumber="8">stat. n.</emphasis>
with the
<typeStatus id="F0118838A948FFBDFBA5F9159E280A9A" box="[1044,1092,1665,1691]" pageId="7" pageNumber="8">type</typeStatus>
species
<taxonomicName id="E8AA4D19A948FFBDFB14F9169BC10ABE" authority="(Sasakawa, 1997)" authorityName="Vilkamaa &amp; Menzel" authorityYear="2019" baseAuthorityName="Sasakawa" baseAuthorityYear="1997" class="Insecta" family="Sciaridae" genus="Merizomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="codonopsivora" status="comb. nov.">
<emphasis id="1DDEEA88A948FFBDFB14F9169AB10ABE" italics="true" pageId="7" pageNumber="8">Merizomma codonopsivora</emphasis>
(
<bibRefCitation id="4B3B4B6BA948FFBDFF5CF9319BC90ABE" author="Sasakawa, M." box="[237,421,1701,1727]" pageId="7" pageNumber="8" pagination="171 - 176" refId="ref45374" refString="Sasakawa, M. (1997) New leaf miner and stem borer of Sciaridae (Diptera). Japanese Journal of Entomology, 65 (1), 171 - 176." type="journal article" year="1997">Sasakawa, 1997</bibRefCitation>
)
</taxonomicName>
<emphasis id="1DDEEA88A948FFBDFE04F93198700ABE" bold="true" box="[437,540,1701,1727]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="06ED57F3A948FFBDFE04F93198700ABE" box="[437,540,1701,1727]" pageId="7" pageNumber="8" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
is raised to generic rank and excluded from the
<taxonomicName id="E8AA4D19A948FFBDFB8EF9319EDD0ABE" box="[1087,1201,1701,1727]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="1DDEEA88A948FFBDFB8EF9319EDD0ABE" box="[1087,1201,1701,1727]" italics="true" pageId="7" pageNumber="8">Lycoriella</emphasis>
</taxonomicName>
group
<taxonomicName id="E8AA4D19A948FFBDFAB3F9329B310AE2" authority="Menzel &amp; Mohrig (2000)" authorityName="Menzel &amp; Mohrig" authorityYear="2000" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sensu">
<emphasis id="1DDEEA88A948FFBDFAB3F9329F2C0ABE" box="[1282,1344,1702,1727]" italics="true" pageId="7" pageNumber="8">sensu</emphasis>
<bibRefCitation id="4B3B4B6BA948FFBDFAF9F9319B310AE2" author="Menzel, F. &amp; Mohrig, W." pageId="7" pageNumber="8" refId="ref43072" refString="Menzel, F. &amp; Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement, 6 (1999). Ampyx-Verlag, Halle (Saale), 761 pp." type="book" year="2000">Menzel &amp; Mohrig (2000)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>