165 lines
19 KiB
XML
165 lines
19 KiB
XML
<document id="46F76BF548E881701BF98F72E4ED9FA5" ID-CLB-Dataset="21114" ID-DOI="10.11646/zootaxa.4665.1.1" ID-GBIF-Dataset="8053cc5c-900d-490b-84e9-1ca29a46aa3c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4000553" ID-ZooBank="78F3F422-DE06-4C61-AC47-69AA684EB5AB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598382139973" checkinUser="felipe" docAuthor="Vilkamaa, Pekka & Menzel, Frank" docDate="2019" docId="A703878CA945FFB1FF26F9AD99D80DB1" docLanguage="en" docName="zootaxa.4665.1.1.pdf.imf" docOrigin="Zootaxa 4665 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hemineurina unguicauda" docType="treatment" docVersion="12" lastPageNumber="11" masterDocId="5B3AFFF4A94FFFBAFFB1FF949A6C0C01" masterDocTitle="Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species" masterLastPageNumber="67" masterPageNumber="1" pageNumber="11" updateTime="1698845439498" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:title id="3AD7D90FB5A77B6930ADB5CED75A8C0E">Re-classification of Lycoriella Frey sensu lato (Diptera, Sciaridae), with description of Trichocoelina gen. n. and twenty new species</mods:title>
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<mods:namePart id="BB27E3015443DE4576D8E041065ED976">Vilkamaa, Pekka</mods:namePart>
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<mods:namePart id="4E1DA59ED35F037D2210DA031F0B8062">Menzel, Frank</mods:namePart>
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<mods:typeOfResource id="899AA51D9DDD2726EE276B965357150F">text</mods:typeOfResource>
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<mods:title id="CB32F4496F52BD650DFF225EEAB9065B">Zootaxa</mods:title>
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<mods:date id="F3BB131095F6C37749BDE89C690C56A0">2019</mods:date>
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<mods:number id="853EF5D82EB8F0EB44A1491EE3474681">2019-09-06</mods:number>
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<mods:number id="DE924A1099D092B384F958E0BF71D111">4665</mods:number>
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<mods:identifier id="226AD21EC6112E4FE4DF7AA93DBEC26C" type="DOI">10.11646/zootaxa.4665.1.1</mods:identifier>
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<treatment id="A703878CA945FFB1FF26F9AD99D80DB1" ID-DOI="http://doi.org/10.5281/zenodo.4340462" ID-GBIF-Taxon="167366595" ID-Zenodo-Dep="4340462" LSID="urn:lsid:plazi:treatment:A703878CA945FFB1FF26F9AD99D80DB1" httpUri="http://treatment.plazi.org/id/A703878CA945FFB1FF26F9AD99D80DB1" lastPageId="11" lastPageNumber="11" pageId="10" pageNumber="11">
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<subSubSection id="67B06511A945FFB0FF26F9AD99110A52" box="[151,893,1593,1620]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="2F15369AA945FFB0FF26F9AD99110A52" blockId="10.[151,893,1593,1620]" box="[151,893,1593,1620]" pageId="10" pageNumber="11">
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<heading id="745D81F6A945FFB0FF26F9AD99110A52" bold="true" box="[151,893,1593,1620]" fontSize="11" level="2" pageId="10" pageNumber="11" reason="4">
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<emphasis id="1DDEEA88A945FFB0FF26F9AD99110A52" bold="true" box="[151,893,1593,1620]" pageId="10" pageNumber="11">
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<taxonomicName id="E8AA4D19A945FFB0FF26F9AD98F10A52" ID-CoL="894TL" authority="(Malloch, 1923)" baseAuthorityName="Malloch" baseAuthorityYear="1923" box="[151,669,1593,1620]" class="Insecta" family="Sciaridae" genus="Hemineurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="unguicauda">
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<emphasis id="1DDEEA88A945FFB0FF26F9AD9BBF0A55" bold="true" box="[151,467,1593,1620]" italics="true" pageId="10" pageNumber="11">Hemineurina unguicauda</emphasis>
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(
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<bibRefCitation id="4B3B4B6BA945FFB0FE53F9AD98F90A52" author="Malloch, J. R." box="[482,661,1593,1619]" pageId="10" pageNumber="11" pagination="170 - 227" refId="ref42516" refString="Malloch, J. R. (1923) Diptera (except Tipulidae, Rhyphidae and Calliphoridae). In: A biological survey of the Pribilof Islands, Alaska. Part II. Insects, arachnids, and chilopods of the Pribilof Islands, Alaska. North American Fauna, 46, Government Printing Office, Washington D. C., pp. 170 - 227." type="book chapter" year="1923">Malloch, 1923</bibRefCitation>
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)
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</taxonomicName>
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restit. et comb. n.
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="67B06511A945FFB0FF26F9149EA80AE2" pageId="10" pageNumber="11" type="reference_group">
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<paragraph id="2F15369AA945FFB0FF26F9149EA80AE2" blockId="10.[151,1437,1664,2015]" pageId="10" pageNumber="11">
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<emphasis id="1DDEEA88A945FFB0FF26F9149B7F0A9A" bold="true" box="[151,275,1664,1691]" pageId="10" pageNumber="11">Literature</emphasis>
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.
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<taxonomicName id="E8AA4D19A945FFB0FE93F91599480A9D" ID-CoL="733CT" authority="(Holmgren)" baseAuthorityName="Holmgren" baseAuthorityYear="1883" box="[290,804,1665,1692]" class="Insecta" family="Sciaridae" genus="Lycoriella" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riparia" subGenus="Hemineurina">
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<emphasis id="1DDEEA88A945FFB0FE93F9159BF80A9A" box="[290,404,1665,1691]" italics="true" pageId="10" pageNumber="11">Lycoriella</emphasis>
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(
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<emphasis id="1DDEEA88A945FFB0FE17F91698560A9A" box="[422,570,1666,1691]" italics="true" pageId="10" pageNumber="11">Hemineurina</emphasis>
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)
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<emphasis id="1DDEEA88A945FFB0FDFAF91698F60A9A" box="[587,666,1666,1691]" italics="true" pageId="10" pageNumber="11">riparia</emphasis>
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(Holmgren)
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</taxonomicName>
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—
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<bibRefCitation id="4B3B4B6BA945FFB0FCF0F9159E420A9A" author="Mohrig, W. & Heller, K. & Hippa, H. & Vilkamaa, P. & Menzel, F." box="[833,1070,1665,1692]" pageId="10" pageNumber="11" pagination="141 - 286" refId="ref43950" refString="Mohrig, W., Heller, K., Hippa, H., Vilkamaa, P. & Menzel, F. (2013) Revision of Black Fungus Gnats (Diptera: Sciaridae) of North America. Studia dipterologica, 19 (1 - 2) (2012), 141 - 286." type="journal article" year="2013">
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Mohrig
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<emphasis id="1DDEEA88A945FFB0FC2EF91699B40A9A" box="[927,984,1665,1691]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
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(2013)
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</bibRefCitation>
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: 214, fig. 38 a, b [misidentification].
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<taxonomicName id="E8AA4D19A945FFB0FF6CF93198830ABE" ID-CoL="894TL" authority="Malloch-Malloch (1923)" authorityName="Malloch-Malloch" authorityYear="1923" box="[221,751,1701,1727]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="unguicauda">
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<emphasis id="1DDEEA88A945FFB0FF6CF9319BD90ABE" box="[221,437,1701,1727]" italics="true" pageId="10" pageNumber="11">Sciara unguicauda</emphasis>
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Malloch—Malloch (1923)
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</taxonomicName>
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: 180, pl. 13, fig. 3.
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<taxonomicName id="E8AA4D19A945FFB0FC6DF9319AA30AE5" authority="(Malloch)" baseAuthorityName="Malloch" baseAuthorityYear="1923" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="unguicauda" subGenus="Bradysia">
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<emphasis id="1DDEEA88A945FFB0FC6DF9319E2D0ABE" box="[988,1089,1701,1727]" italics="true" pageId="10" pageNumber="11">Bradysia</emphasis>
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(
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<emphasis id="1DDEEA88A945FFB0FBE4F9319ED60ABE" box="[1109,1210,1701,1727]" italics="true" pageId="10" pageNumber="11">Bradysia</emphasis>
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)
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<emphasis id="1DDEEA88A945FFB0FB7FF9319F3E0ABE" box="[1230,1362,1701,1727]" italics="true" pageId="10" pageNumber="11">unguicauda</emphasis>
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(Malloch)
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</taxonomicName>
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—
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<bibRefCitation id="4B3B4B6BA945FFB0FF5CF95D986C0AE2" author="Stone, A. & Laffoon, J. L." box="[237,512,1736,1764]" pageId="10" pageNumber="11" pagination="229 - 236" refId="ref46400" refString="Stone, A. & Laffoon, J. L. (1965) Family Sciaridae (Lycoriidae). In: Stone, A., Sabrosky, C. W., Wirth, W. W., Foote, R. H. & Coulson, J. R. (Eds.), A catalog of the Diptera of America north of Mexico. Agriculture Handbook, 276, United States Department of Agriculture, Washington D. C., pp. 229 - 236." type="book chapter" year="1965">Stone & Laffoon (1965)</bibRefCitation>
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: 234.
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<taxonomicName id="E8AA4D19A945FFB0FDF3F95D99CA0AE5" authority="(Malloch)" baseAuthorityName="Malloch" baseAuthorityYear="1923" box="[578,934,1737,1764]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="unguicauda">
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<emphasis id="1DDEEA88A945FFB0FDF3F95D995E0AE2" box="[578,818,1737,1763]" italics="true" pageId="10" pageNumber="11">Bradysia unguicauda</emphasis>
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(Malloch)
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</taxonomicName>
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—
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<bibRefCitation id="4B3B4B6BA945FFB0FC72F95D9E000AE5" author="Steffan, W. A." box="[963,1132,1737,1764]" pageId="10" pageNumber="11" pagination="1 - 77" refId="ref45952" refString="Steffan, W. A. (1966) A generic revision of the family Sciaridae (Diptera) of America north of Mexico. University of California Publications in Entomology, 44, 1 - 77." type="journal article" year="1966">Steffan (1966)</bibRefCitation>
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: 37, 54.
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</paragraph>
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</subSubSection>
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<subSubSection id="67B06511A945FFB1FF76F97899D80DB1" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
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<paragraph id="2F15369AA945FFB1FF76F97899D80DB1" blockId="10.[151,1437,1664,2015]" lastBlockId="11.[151,1437,152,432]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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<emphasis id="1DDEEA88A945FFB0FF76F9789B290B06" bold="true" box="[199,325,1772,1799]" pageId="10" pageNumber="11">Discussion</emphasis>
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. When the figure of the
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<typeStatus id="F0118838A945FFB0FDE7F97998D10B06" box="[598,701,1773,1799]" pageId="10" pageNumber="11" type="lectotype">lectotype</typeStatus>
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of
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<taxonomicName id="E8AA4D19A945FFB0FD50F9799F070B06" authority="Holmgren in Menzel & Mohrig (2000: 407" authorityName="Holmgren in Menzel & Mohrig" authorityYear="2000" box="[737,1387,1773,1799]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riparia">
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<emphasis id="1DDEEA88A945FFB0FD50F97999EC0B06" box="[737,896,1773,1799]" italics="true" pageId="10" pageNumber="11">Sciara riparia</emphasis>
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Holmgren in
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<bibRefCitation id="4B3B4B6BA945FFB0FBADF9799F070B06" author="Menzel, F. & Mohrig, W." box="[1052,1387,1773,1799]" pageId="10" pageNumber="11" refId="ref43072" refString="Menzel, F. & Mohrig, W. (2000) Revision der palaarktischen Trauermucken (Diptera: Sciaridae). Studia dipterologica Supplement, 6 (1999). Ampyx-Verlag, Halle (Saale), 761 pp." type="book" year="2000">Menzel & Mohrig (2000: 407</bibRefCitation>
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</taxonomicName>
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, fig. 375) is compared with the figure of the
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<typeStatus id="F0118838A945FFB0FDFCF88598C30B2A" box="[589,687,1809,1835]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="E8AA4D19A945FFB0FD63F88599B90B2A" authority="Malloch" authorityName="Malloch" box="[722,981,1809,1835]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="unguicauda">
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<emphasis id="1DDEEA88A945FFB0FD63F885991D0B2A" box="[722,881,1809,1835]" italics="true" pageId="10" pageNumber="11">S. unguicauda</emphasis>
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Malloch
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</taxonomicName>
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by
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<bibRefCitation id="4B3B4B6BA945FFB0FC4CF8859F7E0B2A" author="Mohrig, W. & Heller, K. & Hippa, H. & Vilkamaa, P. & Menzel, F." box="[1021,1298,1808,1835]" pageId="10" pageNumber="11" pagination="141 - 286" refId="ref43950" refString="Mohrig, W., Heller, K., Hippa, H., Vilkamaa, P. & Menzel, F. (2013) Revision of Black Fungus Gnats (Diptera: Sciaridae) of North America. Studia dipterologica, 19 (1 - 2) (2012), 141 - 286." type="journal article" year="2013">
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Mohrig
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<emphasis id="1DDEEA88A945FFB0FBE6F8869EE30B2A" box="[1111,1167,1809,1835]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
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(2013: 215
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</bibRefCitation>
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: fig. 38 a), it is clear that the species are distinct.
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<taxonomicName id="E8AA4D19A945FFB0FD94F8A299B20B4E" authority="(Holmgren, 1883)" authorityName="Vilkamaa & Menzel" authorityYear="2019" baseAuthorityName="Holmgren" baseAuthorityYear="1883" box="[549,990,1845,1871]" class="Insecta" family="Sciaridae" genus="Hemineurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="riparia" status="comb. nov.">
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<emphasis id="1DDEEA88A945FFB0FD94F8A299620B4E" box="[549,782,1846,1871]" italics="true" pageId="10" pageNumber="11">Hemineurina riparia</emphasis>
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(
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<bibRefCitation id="4B3B4B6BA945FFB0FCADF8A199B90B4E" author="Holmgren, A. E." box="[796,981,1845,1871]" pageId="10" pageNumber="11" pagination="141 - 190" refId="ref41272" refString="Holmgren, A. E. (1883) Insecta a viris doctissimis Nordenskiold illum ducem sequentibus in insulis Waigatsch et Novaja Semlia anno 1875 collecta. Hymenoptera et Diptera. Entomologisk Tidskrift, 4 (3 - 4), 141 - 190." type="journal article" year="1883">Holmgren, 1883</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="06ED57F3A945FFB0FC55F8A19E280B4E" box="[996,1092,1845,1871]" pageId="10" pageNumber="11" rank="species">comb. n.</taxonomicNameLabel>
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has a narrower gonostylus and a slightly impressed part on the basal side of the apical tooth, where there are three megasetae [two very close to the apical tooth, one of which on its dorsal, one on its ventral side and one megaseta medially at the apical third]. The medial part of the gonostylus between the two groups of megasetae has only short setae, the apical tooth is not on a lobe-like process, and the whiplash seta is on the middle of the medial side. Furthermore, all megasetae are narrow, hyalinous, and on short basal bodies. In comparison,
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<taxonomicName id="E8AA4D19A944FFB1FD5AFF0E9EAB0CB2" authority="(Malloch, 1923)" baseAuthorityName="Malloch" baseAuthorityYear="1923" box="[747,1223,153,179]" class="Insecta" family="Sciaridae" genus="Hemineurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="unguicauda">
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<emphasis id="1DDEEA88A944FFB1FD5AFF0E9E650CB2" box="[747,1033,153,179]" italics="true" pageId="11" pageNumber="12">Hemineurina unguicauda</emphasis>
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(
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<bibRefCitation id="4B3B4B6BA944FFB1FBA9FF0D9ED20CB2" author="Malloch, J. R." box="[1048,1214,153,179]" pageId="11" pageNumber="12" pagination="170 - 227" refId="ref42516" refString="Malloch, J. R. (1923) Diptera (except Tipulidae, Rhyphidae and Calliphoridae). In: A biological survey of the Pribilof Islands, Alaska. Part II. Insects, arachnids, and chilopods of the Pribilof Islands, Alaska. North American Fauna, 46, Government Printing Office, Washington D. C., pp. 170 - 227." type="book chapter" year="1923">Malloch, 1923</bibRefCitation>
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)
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</taxonomicName>
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<emphasis id="1DDEEA88A944FFB1FB7FFF0C9FF10CB2" bold="true" box="[1230,1437,152,179]" pageId="11" pageNumber="12">restit. et comb. n.</emphasis>
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has a more voluminous gonostylus, the medial side of which is strongly impressed ventrally, and the medial margin between the apex of gonostylus and the medial megasetae has strikingly long setae. Of the four megasetae, three are much stronger than those of
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<taxonomicName id="E8AA4D19A944FFB1FE4EFE92981D0D1E" authorityName="Vilkamaa & Menzel" authorityYear="2019" baseAuthorityName="Holmgren" baseAuthorityYear="1883" box="[511,625,262,287]" class="Insecta" family="Sciaridae" genus="Hemineurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="riparia">
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<emphasis id="1DDEEA88A944FFB1FE4EFE92981D0D1E" box="[511,625,262,287]" italics="true" pageId="11" pageNumber="12">H. riparia</emphasis>
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</taxonomicName>
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and two of the megasetae are close to each other on high basal bodies at the middle of the gonostylus. Furthermore, the whiplash seta of
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<taxonomicName id="E8AA4D19A944FFB1FCD0FEBE9E6B0D42" baseAuthorityName="Malloch" baseAuthorityYear="1923" box="[865,1031,297,323]" class="Insecta" family="Sciaridae" genus="Hemineurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="unguicauda">
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<emphasis id="1DDEEA88A944FFB1FCD0FEBE9E6B0D42" box="[865,1031,297,323]" italics="true" pageId="11" pageNumber="12">H. unguicauda</emphasis>
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</taxonomicName>
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is more basally placed (on the basal third of the gonostylus), one slender megaseta is on the margin of the impressed area and one, stronger, megaseta just on the basal side of the apical tooth. Strikingly, the long apical tooth and the subapical megaseta are at the apex of a long non-setose lobe, which is absent from
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<taxonomicName id="E8AA4D19A944FFB1FD01FE0299DC0DB1" authority="(Holmgren)" authorityName="Vilkamaa & Menzel" authorityYear="2019" baseAuthorityName="Holmgren" baseAuthorityYear="1883" box="[688,944,405,432]" class="Insecta" family="Sciaridae" genus="Hemineurina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="riparia">
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<emphasis id="1DDEEA88A944FFB1FD01FE02994D0DAE" box="[688,801,406,431]" italics="true" pageId="11" pageNumber="12">H. riparia</emphasis>
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(Holmgren)
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |