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<document ID-DOI="http://doi.org/10.5281/zenodo.3792392" ID-GBIF-Dataset="125703f8-ddaa-4f28-9673-97b7c469b05c" ID-GBIF-Taxon="163812734" ID-ISSN="13132970" ID-Zenodo-Dep="3792392" checkinTime="1588014657767" checkinUser="plazi" docAuthor="Chamorro-Lacayo, Maria &amp; Konstantinov, Alexander" docDate="2009" docId="552287F6FFB4FFAEFF538EBBFC2E6A34" docLanguage="en" docName="ZK_article_1978.pdf" docOrigin="ZooKeys 8 (8)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@60a754c6" docStyleName="zookeys.2008.journal_article" docTitle="Chlamisus Rafinesque 1815" docType="treatment" docVersion="10" lastPageId="9" lastPageNumber="72" masterDocId="A91BFF8EFFBCFFA7FFDF8F7BFF946F13" masterDocTitle="Synopsis of warty leaf beetle genera of the world (Coleoptera, Chrysomelidae, Cryptocephalinae, Chlamisini)" masterLastPageNumber="88" masterPageNumber="63" pageId="8" pageNumber="71" updateTime="1636502541170" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:titleInfo>
<mods:title>Synopsis of warty leaf beetle genera of the world (Coleoptera, Chrysomelidae, Cryptocephalinae, Chlamisini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chamorro-Lacayo, Maria</mods:namePart>
<mods:affiliation>National Museum of Natural History, Washington D. C. ,,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Konstantinov, Alexander</mods:namePart>
<mods:affiliation>Systematic Entomology Laboratory, MRC- 168 Washington, United States of America</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-04-28</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>8</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>8</mods:number>
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<mods:start>63</mods:start>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.8.90</mods:identifier>
<mods:identifier type="GBIF-Dataset">125703f8-ddaa-4f28-9673-97b7c469b05c</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576444</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3792392" ID-GBIF-Taxon="163812734" ID-Zenodo-Dep="3792392" LSID="urn:lsid:plazi:treatment:552287F6FFB4FFAEFF538EBBFC2E6A34" httpUri="http://treatment.plazi.org/id/552287F6FFB4FFAEFF538EBBFC2E6A34" lastPageId="9" lastPageNumber="72" pageId="8" pageNumber="71">
<subSubSection box="[140,403,448,475]" pageId="8" pageNumber="71" type="nomenclature">
<paragraph blockId="8.[140,583,448,510]" box="[140,403,448,475]" pageId="8" pageNumber="71">
<heading box="[140,403,448,475]" fontSize="11" level="2" pageId="8" pageNumber="71" reason="3">
<taxonomicName authority="Rafinesque" authorityName="Rafinesque" authorityYear="1815" box="[140,403,448,475]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[140,403,448,475]" pageId="8" pageNumber="71">
<emphasis bold="true" box="[140,264,448,475]" italics="true" pageId="8" pageNumber="71">Chlamisus</emphasis>
Rafinesque
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[140,583,483,510]" pageId="8" pageNumber="71" type="description">
<paragraph blockId="8.[140,583,448,510]" box="[140,583,483,510]" pageId="8" pageNumber="71">
(
<figureCitation box="[148,256,483,510]" captionStart="Figure 1" captionStartId="4.[140,212,1526,1548]" captionTargetBox="[166,1087,177,1506]" captionTargetId="figure@4.[137,1109,165,1507]" captionTargetPageId="4" captionText="Figure 1. Chlamisini, dorsal view. A, Aulacochlamys distincta (Achard). B, A. costicollis (Lacordaire. C, Carcinobaena pilula (Klug). D, Chlamisus foveolatus (Knoch). E, Diplacaspis prosternalis (Schaeffer). F, Exema elliptica Karren. G, Fulcidax coelestina (Lacordaire). H, Hymetes javana Lacordaire. I, Melittochlamys specula (Klug). Į, Neochlamisus insularis (Schaeffer). K, N. velutinus Karren. L, Pseudochlamys megalostomoides Lacordaire ♀." figureDoi="http://doi.org/10.5281/zenodo.3788012" httpUri="https://zenodo.org/record/3788012/files/figure.png" pageId="8" pageNumber="71">Figs. 1 D</figureCitation>
;
<figureCitation box="[268,313,484,510]" captionStart="Figure 2" captionStartId="5.[140,212,1526,1548]" captionTargetBox="[158,1106,521,1494]" captionTargetId="figure@5.[140,1108,503,1506]" captionTargetPageId="5" captionText="Figure 2. Chlamisini, lateral view. A, Aulacochlamys distincta (Achard). B, A. costicollis (Lacordaire). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren.F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770841" httpUri="https://zenodo.org/record/3770841/files/figure.png" pageId="8" pageNumber="71">2 C</figureCitation>
;
<figureCitation box="[325,370,484,510]" captionStart="Figure 3" captionStartId="7.[140,212,1526,1548]" captionTargetBox="[148,1095,175,1500]" captionTargetId="figure@7.[136,1110,165,1507]" captionTargetPageId="7" captionText="Figure 3. Chlamisini, ventral view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770843" httpUri="https://zenodo.org/record/3770843/files/figure.png" pageId="8" pageNumber="71">3 C</figureCitation>
;
<figureCitation box="[382,427,484,510]" captionStart="Figure 4" captionStartId="10.[140,212,1526,1548]" captionTargetBox="[140,1106,199,1486]" captionTargetId="figure@10.[140,1108,199,1506]" captionTargetPageId="10" captionText="Figure 4. Chlamisini, frontal view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). J, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♁. L, P. megalostomoides ♀." figureDoi="http://doi.org/10.5281/zenodo.3770847" httpUri="https://zenodo.org/record/3770847/files/figure.png" pageId="8" pageNumber="71">4 C</figureCitation>
;
<figureCitation box="[440,485,483,510]" captionStart="Figure 5" captionStartId="12.[140,213,1526,1548]" captionTargetBox="[142,1102,207,1465]" captionTargetId="figure@12.[136,1110,180,1486]" captionTargetPageId="12" captionText="Figure 5. Chlamisini, caudal view. A, Aulacochlamys costicollis (Lacordaire). B, Carcinobaena pilula (Klug). C, Chlamisus foveolatus (Knoch). D, Diplacaspis prosternalis (Schaeffer). E, Exema elliptica Karren. F, Fulcidax coelestina (Lacordaire). G, Hymetes javana Lacordaire. H, Melittochlamys specula (Klug). I, Neochlamisus insularis (Schaeffer). Į, N. velutinus Karren. K, Pseudochlamys megalostomoides Lacordaire ♀." figureDoi="http://doi.org/10.5281/zenodo.3770849" httpUri="https://zenodo.org/record/3770849/files/figure.png" pageId="8" pageNumber="71">5 C</figureCitation>
;
<figureCitation box="[497,539,484,510]" captionStart="Figure 7" captionStartId="19.[140,212,1068,1090]" captionTargetBox="[249,998,170,1037]" captionTargetId="figure@19.[235,1013,165,1048]" captionTargetPageId="19" captionText="Figure 7. Genitalia of Chlamisini. A, Male ejaculatory guide of Neochlamisus velutinus Karren, dorsal and lateral views. B, Male ejaculatory guide of Chlamisus maculipes (Chevrolat), dorsal and lateral views. C, Spermatheca of Neochlamisus velutinus. D, Spermatheca of Chlamisus maculipes." figureDoi="http://doi.org/10.5281/zenodo.3770851" httpUri="https://zenodo.org/record/3770851/files/figure.png" pageId="8" pageNumber="71">7 B</figureCitation>
, D)
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="71" type="reference_group">
<paragraph blockId="8.[140,1108,554,651]" pageId="8" pageNumber="71">
<treatmentCitationGroup pageId="8" pageNumber="71">
<treatmentCitation author="Knoch AW" box="[140,441,554,580]" page="122" pageId="8" pageNumber="71" year="1801">
<taxonomicName authority="Knoch, 1801: 122" authorityName="Knoch" authorityPageNumber="122" authorityYear="1801" box="[140,441,554,580]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinoida" pageId="8" pageNumber="71" phylum="Mollusca" rank="genus">
<emphasis box="[140,231,554,580]" italics="true" pageId="8" pageNumber="71">Chlamys</emphasis>
<bibRefCitation author="Knoch" box="[237,441,554,580]" firstAuthor="Knoch" pageId="8" pageNumber="71" refId="ref10144" refString="Knoch AW (1801) Neue Beytrage zur Insectenkunde, vol. 1. Schwinkert, Leipzig, 208 pp." type="book" year="1801">Knoch, 1801: 122</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus box="[450,504,554,580]" pageId="8" pageNumber="71">Type</typeStatus>
species:
<taxonomicName authority="Knoch, 1801" authorityName="Knoch" authorityYear="1801" box="[598,941,554,580]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinoida" pageId="8" pageNumber="71" phylum="Mollusca" rank="species" species="foveolata">
<emphasis box="[598,787,554,580]" italics="true" pageId="8" pageNumber="71">Chlamys foveolata</emphasis>
<bibRefCitation author="Knoch" box="[793,941,554,580]" firstAuthor="Knoch" pageId="8" pageNumber="71" refId="ref10144" refString="Knoch AW (1801) Neue Beytrage zur Insectenkunde, vol. 1. Schwinkert, Leipzig, 208 pp." type="book" year="1801">Knoch, 1801</bibRefCitation>
</taxonomicName>
, by subsequent designation of
<bibRefCitation author="Navajas" box="[344,554,589,615]" firstAuthor="Navajas" pageId="8" pageNumber="71" pagination="243 - 245" refId="ref10538" refString="Navajas E (1946) Os genotipos da Fam. Fulcidacidae (Col. Chrysomelidae). Papeis avulsos do Departomento de Zoologia 7: 243 - 245" type="journal article" year="1946">Navajas, 1946: 244</bibRefCitation>
[preoccupied by
<taxonomicName authority="Bolten, 1798" authorityName="Bolten" authorityYear="1798" box="[740,981,589,615]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinoida" pageId="8" pageNumber="71" phylum="Mollusca" rank="genus">
<emphasis box="[740,831,589,615]" italics="true" pageId="8" pageNumber="71">Chlamys</emphasis>
Bolten, 1798
</taxonomicName>
, Mollusca].
</treatmentCitationGroup>
<treatmentCitationGroup box="[140,888,624,651]" pageId="8" pageNumber="71">
<treatmentCitation box="[140,516,624,651]" pageId="8" pageNumber="71">
<taxonomicName authority="Rafinesque, 1815: 116" authorityName="Rafinesque" authorityPageNumber="116" authorityYear="1815" box="[140,516,624,651]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis box="[140,253,624,650]" italics="true" pageId="8" pageNumber="71">Chlamisus</emphasis>
<bibRefCitation author="Rafinesque" box="[260,516,624,651]" firstAuthor="Rafinesque" pageId="8" pageNumber="71" refId="ref10566" refString="Rafinesque CS (1815) Analyse de la nature ou tableau de l'univers et des corps organises. Palermo, 224 pp." type="book" year="1815">Rafinesque, 1815: 116</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(new name for
<taxonomicName authority="Knoch" authorityName="Knoch" authorityYear="1801" box="[695,874,624,651]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinoida" pageId="8" pageNumber="71" phylum="Mollusca" rank="genus">
<emphasis box="[695,788,624,650]" italics="true" pageId="8" pageNumber="71">Chlamys</emphasis>
Knoch
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="71" type="diagnosis">
<paragraph blockId="8.[140,1108,694,1672]" pageId="8" pageNumber="71">
<emphasis bold="true" box="[140,265,694,720]" pageId="8" pageNumber="71">
<emphasis box="[140,260,694,720]" inLineHeading="true" pageId="8" pageNumber="71" reason="1">Diagnosis</emphasis>
.
</emphasis>
Length
<quantity box="[358,525,695,721]" metricMagnitude="-3" metricUnit="m" metricValue="5.404999999999999" metricValueMax="7.72" metricValueMin="3.09" pageId="8" pageNumber="71" unit="mm" value="5.404999999999999" valueMax="7.72" valueMin="3.09">3.09-7.72 mm</quantity>
, width
<quantity box="[608,774,694,721]" metricMagnitude="-3" metricUnit="m" metricValue="3.63" metricValueMax="5.36" metricValueMin="1.9" pageId="8" pageNumber="71" unit="mm" value="3.63" valueMax="5.36" valueMin="1.9">1.90-5.36 mm</quantity>
. General body shape cylindrical. Body usually not metallic in color. Frons in canthus of eye usually without yellow spots (face may be entirely or partly yellow, with yellow area extending into canthus, but elytra do not have velvety spots). Frons glabrous or (rarely) covered with dense hairs. Pronotum and elytra usually glabrous or (rarely) covered with dense short hairs. Antenna serrate beyond 3
<superScript attach="left" box="[361,375,870,884]" fontSize="6" pageId="8" pageNumber="71">rd</superScript>
or 4
<superScript attach="left" box="[431,445,870,884]" fontSize="6" pageId="8" pageNumber="71">th</superScript>
antennomeres, 2
<superScript attach="left" box="[636,654,870,884]" fontSize="6" pageId="8" pageNumber="71">nd</superScript>
antennomere slightly widened, globose, 5
<superScript attach="left" box="[154,168,905,919]" fontSize="6" pageId="8" pageNumber="71">th</superScript>
antennomere nearly as large as 6
<superScript attach="none" box="[533,547,905,919]" fontSize="6" pageId="8" pageNumber="71">th</superScript>
. Pronotum medially elevated, with various bumps and short carinae. Pronotal base opposite mesoscutellum (posterior pronotal lobe) with well differentiated notch. Prosternum posteriorly narrowed, posteriorly much narrower than anterior margin. Anterior margin of metasternum concave. Mesoscutellum short, transverse. Metascutellum not exposed. Sutural serration of elytra usually incomplete (suture entire immediately following mesoscutellum). Elytral tubercules well developed. Males without spines or spinulae on first ventrite. Tibiae slightly curved, slightly flattened, with sharp dorsal edge. Fore- and midtibial apices with spine. Tarsal claw appendiculate. Male ejaculatory guide (part of internal sac of aedeagus) symmetrical, without sheath. Apex of spermathecal duct as wide as rest of pump.
</paragraph>
</subSubSection>
<subSubSection box="[185,835,1257,1285]" pageId="8" pageNumber="71" type="distribution">
<paragraph blockId="8.[140,1108,694,1672]" box="[185,835,1257,1285]" pageId="8" pageNumber="71">
<emphasis bold="true" box="[185,344,1257,1283]" pageId="8" pageNumber="71">
<emphasis box="[185,339,1257,1283]" inLineHeading="true" pageId="8" pageNumber="71" reason="1">Distribution</emphasis>
.
</emphasis>
Cosmopolitan (
<bibRefCitation author="Monros" box="[528,687,1258,1285]" firstAuthor="Monros" pageId="8" pageNumber="71" pagination="489 - 672" refId="ref10435" refString="Monros F (1952) Revision de las especies argentinas de Chlamisinae (Col., Chrysomelidae) (5 a contribucion al conocimiento de Chlamisinae). Acta Zoologica Lilloana 10: 489 - 672." type="journal article" year="1952">Monrós 1952</bibRefCitation>
;
<bibRefCitation author="Reid" box="[699,822,1258,1284]" firstAuthor="Reid" pageId="8" pageNumber="71" pagination="315 - 323" refId="ref10589" refString="Reid CAM (1991) The Australian species of Chlamisini (Coleoptera: Chrysomelidae). Journal of the Australian Entomological Society 30: 315 - 323." type="journal article" year="1991">Reid 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="72" pageId="8" pageNumber="71" type="discussion">
<paragraph blockId="8.[140,1108,694,1672]" pageId="8" pageNumber="71">
<emphasis bold="true" box="[185,298,1293,1319]" pageId="8" pageNumber="71">
<emphasis box="[185,293,1293,1319]" inLineHeading="true" pageId="8" pageNumber="71" reason="1">Remarks</emphasis>
.
</emphasis>
<taxonomicName authorityName="Rafinesque" authorityYear="1815" box="[304,417,1293,1319]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis box="[304,417,1293,1319]" italics="true" pageId="8" pageNumber="71">Chlamisus</emphasis>
</taxonomicName>
is close to
<taxonomicName authorityName="Lacordaire" authorityYear="1848" box="[537,610,1294,1319]" class="Insecta" family="Chrysomelidae" genus="Exema" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis box="[537,610,1294,1319]" italics="true" pageId="8" pageNumber="71">Exema</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Karren" authorityYear="1972" box="[665,811,1293,1319]" class="Insecta" family="Chrysomelidae" genus="Neochlamisus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis box="[665,811,1293,1319]" italics="true" pageId="8" pageNumber="71">Neochlamisus</emphasis>
</taxonomicName>
. From
<taxonomicName authorityName="Karren" authorityYear="1972" box="[893,1039,1293,1319]" class="Insecta" family="Chrysomelidae" genus="Neochlamisus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis box="[893,1039,1293,1319]" italics="true" pageId="8" pageNumber="71">Neochlamisus</emphasis>
</taxonomicName>
it can be separated by the following characters: body usually not metallic in color; frons in canthus of eye usually without yellow spots (face may be all or partly yellow, with yellow area extending into canthus, but elytra do not have velvety spots); and male ejaculatory guide symmetrical, without sheath. From
<taxonomicName authorityName="Lacordaire" authorityYear="1848" box="[690,763,1435,1460]" class="Insecta" family="Chrysomelidae" genus="Exema" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis box="[690,763,1435,1460]" italics="true" pageId="8" pageNumber="71">Exema</emphasis>
</taxonomicName>
it can be separated by the following characters: males without spines or spinulae on first ventrite; 5
<superScript attach="left" box="[937,951,1468,1482]" fontSize="6" pageId="8" pageNumber="71">th</superScript>
antennomere nearly as large as 6
<superScript attach="none" box="[355,369,1503,1517]" fontSize="6" pageId="8" pageNumber="71">th</superScript>
; sutural serration of elytra usually incomplete (suture entire immediately following mesoscutellum); and prosternum posteriorly pointed (narrowed), posteriorly much narrower than anterior margin.
</paragraph>
<paragraph blockId="8.[140,1108,694,1672]" lastBlockId="9.[140,1108,166,1320]" lastPageId="9" lastPageNumber="72" pageId="8" pageNumber="71">
No Australasian species were examined for this study, but according to
<bibRefCitation author="Reid" box="[971,1108,1610,1636]" firstAuthor="Reid" pageId="8" pageNumber="71" pagination="315 - 323" refId="ref10589" refString="Reid CAM (1991) The Australian species of Chlamisini (Coleoptera: Chrysomelidae). Journal of the Australian Entomological Society 30: 315 - 323." type="journal article" year="1991">Reid (1991)</bibRefCitation>
native Australasian species of
<taxonomicName authorityName="Rafinesque" authorityYear="1815" box="[483,596,1645,1671]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="71" phylum="Arthropoda" rank="genus">
<emphasis box="[483,596,1645,1671]" italics="true" pageId="8" pageNumber="71">Chlamisus</emphasis>
</taxonomicName>
do not fit into any current definition given for either American (
<bibRefCitation author="Karren" box="[380,530,167,193]" firstAuthor="Karren" pageId="9" pageNumber="72" pagination="875 - 988" refId="ref10110" refString="Karren JB (1972) A revision of the subfamily Chlamisinae of America north of Mexico (Coleoptera: Chrysomelidae). The University of Kansas Science Bulletin 49: 875 - 988." type="journal article" year="1972">Karren 1972</bibRefCitation>
;
<bibRefCitation author="Monros" box="[544,702,166,193]" firstAuthor="Monros" pageId="9" pageNumber="72" pagination="451 - 463" refId="ref10402" refString="Monros F (1951) Notes on Chrysomelid beetles of the subfamily Chlamisinae, with descriptions of new species. Proceedings of the United States National Museum 101: 451 - 463." type="journal article" year="1951">Monrós 1951</bibRefCitation>
) or Oriental (
<bibRefCitation author="Gressitt and Kimoto" firstAuthor="Gressitt" pageId="9" pageNumber="72" pagination="1 - 299" refId="ref9895" refString="Gressitt JL, Kimoto S (1961) The Chrysomelidae (Coleopt.) of China and Korea. Part 1. Pacific Insects Monographs 1 A: 1 - 299." type="journal article" year="1961">Gressitt and Kimoto, 1961</bibRefCitation>
) species.
<bibRefCitation author="Reid" box="[310,449,202,228]" firstAuthor="Reid" pageId="9" pageNumber="72" pagination="315 - 323" refId="ref10589" refString="Reid CAM (1991) The Australian species of Chlamisini (Coleoptera: Chrysomelidae). Journal of the Australian Entomological Society 30: 315 - 323." type="journal article" year="1991">Reid (1991)</bibRefCitation>
also recognized several characteristics distinguishing Oriental (South-East Asian) and American species.
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" pageId="9" pageNumber="72">
Comparison of the
<typeStatus box="[394,487,273,299]" pageId="9" pageNumber="72">paratype</typeStatus>
of
<taxonomicName authority="Medvedev, 1993" authorityName="Medvedev" authorityYear="1993" box="[519,877,272,298]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="rousei">
<emphasis box="[519,692,272,298]" italics="true" pageId="9" pageNumber="72">Chlamisus rousei</emphasis>
Medvedev, 1993
</taxonomicName>
described from
<collectingCountry name="Puerto Rico" pageId="9" pageNumber="72">Puerto Rico</collectingCountry>
with specimens of
<taxonomicName authority="Suffrian, 1866" authorityName="Suffrian" authorityYear="1866" box="[430,811,307,333]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="straminea">
<emphasis box="[430,646,307,333]" italics="true" pageId="9" pageNumber="72">Chlamisus straminea</emphasis>
Suffrian, 1866
</taxonomicName>
from
<collectingCountry box="[878,1009,307,334]" name="Puerto Rico" pageId="9" pageNumber="72">Puerto Rico</collectingCountry>
and
<collectingCountry name="Virgin Islands" pageId="9" pageNumber="72">Virgin Islands</collectingCountry>
revealed that they are conspecific, therefore we here synonymize these names.
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" pageId="9" pageNumber="72">
There are approximately 400 species of
<taxonomicName authorityName="Rafinesque" authorityYear="1815" box="[652,765,378,404]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="genus">
<emphasis box="[652,765,378,404]" italics="true" pageId="9" pageNumber="72">Chlamisus</emphasis>
</taxonomicName>
in the world (
<bibRefCitation author="Monros" box="[940,1102,378,405]" firstAuthor="Monros" pageId="9" pageNumber="72" pagination="489 - 672" refId="ref10435" refString="Monros F (1952) Revision de las especies argentinas de Chlamisinae (Col., Chrysomelidae) (5 a contribucion al conocimiento de Chlamisinae). Acta Zoologica Lilloana 10: 489 - 672." type="journal article" year="1952">Monrós 1952</bibRefCitation>
;
<bibRefCitation author="Reid" box="[140,261,413,439]" firstAuthor="Reid" pageId="9" pageNumber="72" pagination="315 - 323" refId="ref10589" refString="Reid CAM (1991) The Australian species of Chlamisini (Coleoptera: Chrysomelidae). Journal of the Australian Entomological Society 30: 315 - 323." type="journal article" year="1991">Reid 1991</bibRefCitation>
); approximately 100 Neotropical, 45 Palearctic, 18 Nearctic, 22 Oriental, 6 Afrotropical, and 2 Australasian (naturally occurring).
</paragraph>
</subSubSection>
<subSubSection pageId="9" pageNumber="72" type="materials_examined">
<paragraph blockId="9.[140,1108,166,1320]" box="[185,416,483,509]" pageId="9" pageNumber="72">
<emphasis bold="true" box="[185,416,483,509]" pageId="9" pageNumber="72">
<emphasis box="[185,286,483,509]" inLineHeading="true" pageId="9" pageNumber="72" reason="1">Material</emphasis>
examined.
</emphasis>
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[185,520,519,546]" pageId="9" pageNumber="72">
<taxonomicName authority="(Knoch)" baseAuthorityName="Knoch" baseAuthorityYear="1801" box="[185,516,519,546]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis box="[185,409,519,545]" italics="true" pageId="9" pageNumber="72">Chlamisus foveolatus</emphasis>
(Knoch)
</taxonomicName>
:
</paragraph>
<materialsCitation ID-GBIF-Occurrence="2612320303" pageId="9" pageNumber="72">
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1108,553,581]" pageId="9" pageNumber="72">
1) a. [
<collectingCountry box="[295,355,553,580]" name="United States of America" pageId="9" pageNumber="72">USA</collectingCountry>
] Beltsville,
<collectionCode box="[485,537,554,580]" country="Germany" name="Museum Donaueschingen" pageId="9" pageNumber="72">MD</collectionCode>
,
<collectionCode box="[548,601,554,580]" pageId="9" pageNumber="72">VIII</collectionCode>
. 5/ b.
<taxonomicName authority="(Knoch)" baseAuthorityName="Knoch" baseAuthorityYear="1801" box="[673,999,554,581]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis box="[673,897,554,580]" italics="true" pageId="9" pageNumber="72">Chlamisus foveolatus</emphasis>
(Knoch)
</taxonomicName>
det. Kar-
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1107,588,616]" pageId="9" pageNumber="72">
ren 1971. 2) a. [
<collectingCountry box="[411,471,588,615]" name="United States of America" pageId="9" pageNumber="72">USA</collectingCountry>
]
<collectionCode box="[484,518,589,615]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="9" pageNumber="72">VI</collectionCode>
.10.11, Southern Pines, NC, AH Manee/ b.
<emphasis box="[1015,1107,589,615]" italics="true" pageId="9" pageNumber="72">Chlami-</emphasis>
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,683,624,651]" pageId="9" pageNumber="72">
<emphasis box="[231,261,625,650]" italics="true" pageId="9" pageNumber="72">sus</emphasis>
<taxonomicName authority="(Knoch)" baseAuthorityName="Knoch" baseAuthorityYear="1801" box="[268,475,624,651]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="foveolatus">
<emphasis box="[268,372,624,650]" italics="true" pageId="9" pageNumber="72">foveolatus</emphasis>
(Knoch)
</taxonomicName>
det. Karren 1971.
</paragraph>
</materialsCitation>
<paragraph blockId="9.[140,1108,166,1320]" box="[185,556,659,686]" pageId="9" pageNumber="72">
<taxonomicName authority="(Chevrolat)" baseAuthorityName="Chevrolat" baseAuthorityYear="1835" box="[185,552,659,686]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="maculipes">
<emphasis box="[185,410,659,685]" italics="true" pageId="9" pageNumber="72">Chlamisus maculipes</emphasis>
(Chevrolat)
</taxonomicName>
:
</paragraph>
<materialsCitation ID-GBIF-Occurrence="2612320311" pageId="9" pageNumber="72">
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1107,694,721]" pageId="9" pageNumber="72">
1) a.
<collectingCountry box="[296,383,695,721]" name="Mexico" pageId="9" pageNumber="72">Mexico</collectingCountry>
, Salle/ b. F. Monrós Collection, 1959/ c.
<taxonomicName baseAuthorityName="Chevrolat" baseAuthorityYear="1835" box="[879,1107,695,721]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="maculipes">
<emphasis box="[879,1107,695,721]" italics="true" pageId="9" pageNumber="72">Chlamisus maculipes</emphasis>
</taxonomicName>
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,571,730,757]" pageId="9" pageNumber="72">(Chevr.) F. Monrós det. 1949.</paragraph>
</materialsCitation>
<paragraph blockId="9.[140,1108,166,1320]" box="[185,505,765,791]" pageId="9" pageNumber="72">
<taxonomicName authority="Monros" authorityName="Monros" box="[185,501,765,791]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="obidensis">
<emphasis box="[185,401,765,791]" italics="true" pageId="9" pageNumber="72">Chlamisus obidensis</emphasis>
Monrós
</taxonomicName>
:
</paragraph>
<materialsCitation ID-GBIF-Occurrence="2612320319" pageId="9" pageNumber="72">
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1107,800,827]" pageId="9" pageNumber="72">
1) a. [
<collectingCountry box="[295,366,800,827]" name="Brazil" pageId="9" pageNumber="72">Brazil</collectingCountry>
] Teffe (Ega), Amazonas, M. de Mathan, IV.78 et I.79/ b. F. Mon-
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,471,835,862]" pageId="9" pageNumber="72">rós Collection, 1959.</paragraph>
</materialsCitation>
<paragraph blockId="9.[140,1108,166,1320]" box="[185,509,871,898]" pageId="9" pageNumber="72">
<taxonomicName authority="(Lefevre)" baseAuthorityName="Lefevre" box="[185,505,871,898]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="pilifrons">
<emphasis box="[185,393,871,897]" italics="true" pageId="9" pageNumber="72">Chlamisus pilifrons</emphasis>
(Lefevre)
</taxonomicName>
:
</paragraph>
<materialsCitation ID-GBIF-Occurrence="2612320313" pageId="9" pageNumber="72">
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1107,905,933]" pageId="9" pageNumber="72">
1) a. [
<collectingCountry box="[295,360,905,932]" name="India" pageId="9" pageNumber="72">India</collectingCountry>
] Rhamnaq (Hindustan)/ b. Ex. Musaeo Lefevre, 1894/ c. F. Mon-
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1015,941,968]" pageId="9" pageNumber="72">
rós Collection, 1959/ d.
<taxonomicName authority="Lef. F. Monros" box="[510,891,941,968]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="pilifrons">
<emphasis box="[510,718,941,967]" italics="true" pageId="9" pageNumber="72">Chlamisus pilifrons</emphasis>
Lef. F. Monrós
</taxonomicName>
det. 1956.
</paragraph>
</materialsCitation>
<paragraph blockId="9.[140,1108,166,1320]" box="[185,473,976,1002]" pageId="9" pageNumber="72">
<taxonomicName authority="Chen" authorityName="Chen" box="[185,469,976,1002]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="pubiceps">
<emphasis box="[185,395,976,1002]" italics="true" pageId="9" pageNumber="72">Chlamisus pubiceps</emphasis>
Chen
</taxonomicName>
:
</paragraph>
<materialsCitation ID-GBIF-Occurrence="2612320308" pageId="9" pageNumber="72">
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1107,1011,1038]" pageId="9" pageNumber="72">
1) a.
<collectingCountry box="[288,363,1011,1037]" name="Russia" pageId="9" pageNumber="72">Russia</collectingCountry>
, Far East, Ussury region, Kondratevka,
<date box="[814,959,1012,1038]" pageId="9" pageNumber="72" value="1992-07-28">28.VII.1992</date>
, leg. A. Kon-
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,917,1047,1073]" pageId="9" pageNumber="72">
stantinov/ b.
<taxonomicName authority="Chen" authorityName="Chen" box="[380,663,1047,1073]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="pubiceps">
<emphasis box="[380,590,1047,1073]" italics="true" pageId="9" pageNumber="72">Chlamisus pubiceps</emphasis>
Chen
</taxonomicName>
, det. A. Konstantinov.
</paragraph>
</materialsCitation>
<paragraph blockId="9.[140,1108,166,1320]" box="[185,485,1082,1108]" pageId="9" pageNumber="72">
<taxonomicName authority="Chen" authorityName="Chen" authorityYear="1940" box="[185,480,1082,1108]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="semirufus">
<emphasis box="[185,407,1082,1108]" italics="true" pageId="9" pageNumber="72">Chlamisus semirufus</emphasis>
Chen
</taxonomicName>
:
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,752,1117,1144]" pageId="9" pageNumber="72">
<materialsCitation ID-GBIF-Occurrence="2612320301" box="[231,752,1117,1144]" pageId="9" pageNumber="72">
1) a. nr. Foochow,
<collectingCountry box="[442,515,1117,1143]" name="China" pageId="9" pageNumber="72">China</collectingCountry>
, 1921-4, CR Kellog.
</materialsCitation>
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[185,516,1152,1178]" pageId="9" pageNumber="72">
<taxonomicName authority="Suffrian" authorityName="Suffrian" authorityYear="1866" box="[185,512,1152,1178]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="straminea">
<emphasis box="[185,411,1152,1178]" italics="true" pageId="9" pageNumber="72">Chlamisus straminea</emphasis>
Suffrian
</taxonomicName>
:
</paragraph>
<materialsCitation ID-GBIF-Occurrence="2612320323" pageId="9" pageNumber="72">
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1107,1187,1214]" pageId="9" pageNumber="72">
1) a. [
<collectingCountry box="[293,433,1187,1214]" name="Puerto Rico" pageId="9" pageNumber="72">Puerto Rico</collectingCountry>
] on leaves of
<emphasis box="[582,687,1188,1213]" italics="true" pageId="9" pageNumber="72">Diascorea</emphasis>
,
<collectingRegion box="[700,768,1188,1214]" country="Puerto Rico" name="Ponce" pageId="9" pageNumber="72">Ponce</collectingRegion>
PR, 11.
<collectionCode box="[855,913,1188,1214]" pageId="9" pageNumber="72">VIII</collectionCode>
.33,
<collectionCode box="[960,983,1188,1214]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="72">R</collectionCode>
.G. Oakley,
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1108,1222,1249]" pageId="9" pageNumber="72">
SS 4489. 2) a. St. Thomas Vir. Ids.,
<date box="[626,770,1222,1249]" pageId="9" pageNumber="72" value="1917-06-05">June 5, 1917</date>
(122), Harold Morrison/ b. F.
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,1108,1258,1285]" pageId="9" pageNumber="72">
Monrós Collection, 1959/ c.
<taxonomicName authority="Suffr. F. Monros" box="[565,983,1258,1284]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="straminea">
<emphasis box="[565,791,1258,1284]" italics="true" pageId="9" pageNumber="72">Chlamisus straminea</emphasis>
Suffr. F. Monrós
</taxonomicName>
det. 1954.
</paragraph>
<paragraph blockId="9.[140,1108,166,1320]" box="[231,954,1293,1320]" pageId="9" pageNumber="72">
3) a. From
<collectingCountry box="[357,493,1293,1320]" name="Puerto Rico" pageId="9" pageNumber="72">Puerto Rico</collectingCountry>
/ b.
<typeStatus box="[538,651,1294,1320]" pageId="9" pageNumber="72">Paratypus</typeStatus>
,
<taxonomicName authority="L. Medv." authorityName="L. Medv." box="[662,954,1293,1320]" class="Insecta" family="Chrysomelidae" genus="Chlamisus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="72" phylum="Arthropoda" rank="species" species="rousei">
<emphasis box="[662,844,1293,1319]" italics="true" pageId="9" pageNumber="72">Chlamisus rousei</emphasis>
L. Medv.
</taxonomicName>
</paragraph>
</materialsCitation>
</subSubSection>
</treatment>
</document>