223 lines
31 KiB
XML
223 lines
31 KiB
XML
<document ID-CLB-Dataset="6712" ID-DOI="http://dx.doi.org/10.3897/dez.65.21000" ID-GBIF-Dataset="03cce405-5887-4474-b040-35de18e8d020" ID-Pensoft-Pub="1860-1324-1-1" ID-Pensoft-UUID="77D7AEC9294554E5BBDCABA37901022A" ID-Zenodo-Dep="1166397" ID-ZooBank="73DEE0F32BB04A21B4455E168FE50F54" ModsDocID="1860-1324-65-1-1" checkinTime="1621372489409" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2018" docId="3B420E241A1BA514BF5825E6DFC8C940" docLanguage="en" docName="DeutEntomolZeit 65(1): 1-63" docOrigin="Deutsche Entomologische Zeitschrift 65 (1)" docPubDate="2018-01-18" docSource="http://dx.doi.org/10.3897/dez.65.21000" docTitle="Mecyclothorax manautei Liebherr 2018, sp. n." docType="treatment" docUuid="0A0B84B4-7BC4-4925-B2DE-6AAF3582A1BF" docUuidSource="ZooBank" docVersion="5" id="77D7AEC9294554E5BBDCABA37901022A" lastPageNumber="34" masterDocId="77D7AEC9294554E5BBDCABA37901022A" masterDocTitle="Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel" masterLastPageNumber="63" masterPageNumber="1" pageNumber="33" updateTime="1732929906348" updateUser="ExternalLinkService">
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<mods:titleInfo id="9365B538289139D3F7E29DBA8EE5F72A">
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<mods:title id="5178581D3BB19E938F44DBFBB71B11AD">Cladistic classification of Mecyclothorax Sharp (Coleoptera, Carabidae, Moriomorphini) and taxonomic revision of the New Caledonian subgenus Phacothorax Jeannel</mods:title>
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<mods:roleTerm id="E2EE99E62EC8268FE95A67068E6F21A1">Author</mods:roleTerm>
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<mods:namePart id="B5E27F7DC9DD80059517B777623DD321">Liebherr, James K.</mods:namePart>
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<mods:nameIdentifier id="16E7C6AE26688B34288EB0E3028C9CB5" type="ORCID">https://orcid.org/0000-0001-9831-884X</mods:nameIdentifier>
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<mods:affiliation id="466ACC7ABF114339A6DCEE3514195407">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:nameIdentifier id="16D82165DEF0FAA2F5A951B9A5BAB1C7" type="email">jkl5@cornell.edu</mods:nameIdentifier>
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<mods:typeOfResource id="F7811DA404571443F07B883C89F9BE87">text</mods:typeOfResource>
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<mods:title id="F229792C61A1E8A42262AF85A7C845FE">Deutsche Entomologische Zeitschrift</mods:title>
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<mods:part id="96F247CE9175A45B9CE8C375CAC11DCA">
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<mods:date id="DE63D44F2CBADD340D24FB4DC68ABDE8">2018</mods:date>
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<mods:number id="1A35E30922EC6F546C282917CFAA73DB">2018-01-18</mods:number>
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<mods:number id="03F4B5F1CD5046DB81196AF08A50AFF6">65</mods:number>
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<mods:number id="85655D232739566E1814A43948FB801A">1</mods:number>
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<mods:url id="EF312FB2148D332AE095DF6B1FC39AB5">http://dx.doi.org/10.3897/dez.65.21000</mods:url>
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<mods:classification id="28CEE41A816C0E7E5EE767FBC7781574">journal article</mods:classification>
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<mods:identifier id="CEA5B4E41670DDCCD08484E23B196D00" type="DOI">http://dx.doi.org/10.3897/dez.65.21000</mods:identifier>
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<treatment id="3B420E241A1BA514BF5825E6DFC8C940" ID-GBIF-Taxon="182223675" LSID="urn:lsid:zoobank.org:act:0A0B84B4-7BC4-4925-B2DE-6AAF3582A1BF" httpUri="http://treatment.plazi.org/id/3B420E241A1BA514BF5825E6DFC8C940" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" scope_family="Carabidae" scope_order="Coleoptera" scope_tribe="Moriomorphini">
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<subSubSection id="1FF7171EAA858C5FCC4D74CDE503BAD5" pageId="32" pageNumber="33" type="nomenclature">
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<paragraph id="0B6EF9E5C873DAD6F726A3CFCECE63FA" pageId="32" pageNumber="33">
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11.
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<taxonomicName id="C1CC8C4ED5540F43AF4F6D11CCF72508" ID-CoL="8MBJS" LSID="http://zoobank.org/0A0B84B4-7BC4-4925-B2DE-6AAF3582A1BF" authority="Liebherr" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax manautei" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="manautei" status="sp. n.">Mecyclothorax manautei Liebherr</taxonomicName>
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<taxonomicNameLabel id="0482F93BD62659B5666C87819DB11BD6" pageId="32" pageNumber="33">sp. n.</taxonomicNameLabel>
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<figureCitation id="4D36ECAA32E3A23023244A7326C95A07" captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="32" pageNumber="33">Figures 17I-J</figureCitation>
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<figureCitation id="960DC77B84E029806DB985177649DDC8" captionStart="Figure 18" captionStartId="F19" captionText="Figure 18. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. manautei; F, M. paniensis; G, M. mouensis." figureDoi="10.3897/dez.65.21000.figure18" httpUri="https://binary.pensoft.net/fig/179974" pageId="32" pageNumber="33">, 18E</figureCitation>
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<figureCitation id="F02A2CB00A265CF3E064B48BED80B002" captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="32" pageNumber="33">, 20A</figureCitation>
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<figureCitation id="99DC3E6781B1D89178257049522B9516" captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="32" pageNumber="33">, 23A</figureCitation>
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<figureCitation id="134A600897C6E7A25679D05012856AE9" captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="32" pageNumber="33">, 24A</figureCitation>
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<figureCitation id="D3C8B89EC1A5593E35C1BD10BE970E64" captionStart="Figure 25" captionStartId="F26" captionText="Figure 25. Geographical distributions of Mecyclothorax (Phacothorax) spp. (asterisked locality attributed to M. mouensis based on non-type female specimen)." figureDoi="10.3897/dez.65.21000.figure25" httpUri="https://binary.pensoft.net/fig/179982" pageId="32" pageNumber="33">, 25</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="51F4A55FF8EEC1052F6C0FC0A9280816" pageId="32" pageNumber="33" type="diagnosis">
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<paragraph id="9CA783AFCD236555833AD512CCF0BD02" pageId="32" pageNumber="33">Diagnosis.</paragraph>
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<paragraph id="9743192BD0292D9AE71C976144F3C5CE" pageId="32" pageNumber="33">
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This species can be diagnosed by small body size-standardized body length 3.5-3.9 mm-and the transversely ovoid pronotum with evenly convex lateral margins basally, i.e., hind angles lacking (Fig.
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<figureCitation id="0807677C78A3DC8F879B50D4E1235BBB" captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="32" pageNumber="33">20A</figureCitation>
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). The elytral striae are much more reduced than in the prior two species-
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<taxonomicName id="2B670AD28490E55607D82D6B357FA136" lsidName="M. laterovatulus" pageId="32" pageNumber="33" rank="species" species="laterovatulus">
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<emphasis id="0B61E90C3390C39248C08910A2A98044" italics="true" pageId="32" pageNumber="33">M. laterovatulus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="17FED5C9345AE39F1AAD85590C6551CE" lsidName="M. najtae" pageId="32" pageNumber="33" rank="species" species="najtae">
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<emphasis id="8D6A206614A50FAA757EAB670D436B25" italics="true" pageId="32" pageNumber="33">M. najtae</emphasis>
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</taxonomicName>
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-with the sutural stria obsolete at midlength, and only striae 2-3 evident there, though very shallow. The eyes broadly cover the ocular lobe, ocular lobe ratio 0.88-0.89, and are very convex, i.e.
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<normalizedToken id="C156171CB37CCA98D1C06D87A497DF80" originalValue="“popeyed”">"popeyed"</normalizedToken>
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, with EyL/EyD = 2.2-2.4. Chaetotaxy +/+//+/-//+/2/+/+.
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</paragraph>
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<caption id="E8204351CBFC298BBA741882F81B29A4" ID-Zenodo-Dep="3488467" doi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="32" pageNumber="33" start="Figure 20" startId="F21">
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<paragraph id="36A54187014F399143DE16D1B98F6D89" pageId="32" pageNumber="33">
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<emphasis id="8A474C72F0E65A5D0C9A9F5B1339EBB4" bold="true" pageId="32" pageNumber="33">Figure 20.</emphasis>
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New Caledonian
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<taxonomicName id="4636BEEBAF909A4055DB74B0AB2A744A" authority="" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax (Phacothorax)" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="subGenus" subGenus="Phacothorax">Mecyclothorax (Phacothorax)</taxonomicName>
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spp., dorsal view:
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<emphasis id="2A440BCE8EB8E9037BF6C522616A8B37" bold="true" pageId="32" pageNumber="33">A</emphasis>
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,
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<taxonomicName id="2D42875A20B724E6BEA28D1A9B4E5C08" lsidName="M. manautei" pageId="32" pageNumber="33" rank="species" species="manautei">
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<emphasis id="08FDFEAF94F7D87EDBCFA5300FF96A9F" italics="true" pageId="32" pageNumber="33">M. manautei</emphasis>
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</taxonomicName>
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;
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<emphasis id="CD6FFAD65928AB52F5CA2356485D31AB" bold="true" pageId="32" pageNumber="33">B</emphasis>
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,
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<taxonomicName id="1552C1A480B0FE03F2CD3AA67C698305" lsidName="M. paniensis" pageId="32" pageNumber="33" rank="species" species="paniensis">
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<emphasis id="B79E78ED090584FD76B61EDFD27E2D9F" italics="true" pageId="32" pageNumber="33">M. paniensis</emphasis>
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</taxonomicName>
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;
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<emphasis id="428363A0700F01E35970C2555320CAA4" bold="true" pageId="32" pageNumber="33">C</emphasis>
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,
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<taxonomicName id="4668058286EE212B38BEDC82B9A53E44" lsidName="M. mouensis" pageId="32" pageNumber="33" rank="species" species="mouensis">
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<emphasis id="0077EB25087493596E005671ADAA5319" italics="true" pageId="32" pageNumber="33">M. mouensis</emphasis>
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</taxonomicName>
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;
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<emphasis id="0A71F557AA1820D83FC446D145F96FB7" bold="true" pageId="32" pageNumber="33">D</emphasis>
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,
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<taxonomicName id="C59A70B20E00C9A36D2DF9713E0CE72C" lsidName="M. kanak" pageId="32" pageNumber="33" rank="species" species="kanak">
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<emphasis id="B4B8045E6EDE2D0A5318D99D47E6AED7" italics="true" pageId="32" pageNumber="33">M. kanak</emphasis>
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</taxonomicName>
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(Mt. Humboldt, 580 m);
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<emphasis id="57830B82FA177AF7DB07E7461D42D45D" bold="true" pageId="32" pageNumber="33">E</emphasis>
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,
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<taxonomicName id="F21784B112D2A25BEFD0CF237568C0F4" lsidName="M. kanak" pageId="32" pageNumber="33" rank="species" species="kanak">
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<emphasis id="294960FA093C43020D5AB34E30B20937" italics="true" pageId="32" pageNumber="33">M. kanak</emphasis>
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</taxonomicName>
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(Mt. Humboldt, 630 m);
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<emphasis id="0AF86E15FBF2F0CEEFA7638D8FA6F2A8" bold="true" pageId="32" pageNumber="33">F</emphasis>
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,
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<taxonomicName id="73BDFA26F776022DDC51CDF5A0A3D323" lsidName="M. kanak" pageId="32" pageNumber="33" rank="species" species="kanak">
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<emphasis id="137E93034E06D551CFBD564B75E2C4A3" italics="true" pageId="32" pageNumber="33">M. kanak</emphasis>
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</taxonomicName>
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(Mt. Dzumac);
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<emphasis id="1EFC51707C5792265A86C9F61867D4E8" bold="true" pageId="32" pageNumber="33">G</emphasis>
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,
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<taxonomicName id="A1D2EEA03C141BFD54B4AFF5FFC447DD" lsidName="M. kanak" pageId="32" pageNumber="33" rank="species" species="kanak">
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<emphasis id="6CDF9B0A1DBBC038E0C5B5B4D5A85BC9" italics="true" pageId="32" pageNumber="33">M. kanak</emphasis>
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</taxonomicName>
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(Col de
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<normalizedToken id="098F63EEBDB2C7A9EEC90C4102FE49E3" originalValue="Yaté">Yate</normalizedToken>
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);
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<emphasis id="3141D5D7D188120093917FA52BA60FBD" bold="true" pageId="32" pageNumber="33">H</emphasis>
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,
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<taxonomicName id="BCA490C8CEE3FB82DAAB4D51E616A1EB" lsidName="M. picdupinsensis" pageId="32" pageNumber="33" rank="species" species="picdupinsensis">
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<emphasis id="9195BF4A13462741D7005328B47D884A" italics="true" pageId="32" pageNumber="33">M. picdupinsensis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="EEC8FD1761E9B6CF2BECECF9E0E27D3E" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="description">
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<paragraph id="21500E1EFA3B282F44661DC7B07F46BF" pageId="32" pageNumber="33">Description</paragraph>
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<paragraph id="F118762EAA3DA0D4D56C8B97872AEBA9" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
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(n = 4). Head capsule quadrate, eyes very convex, popeyed, ocular lobe meeting gena at obtuse angle close to eye posterior margin; 14 ommatidia along horizontal diameter of eye; ocular ratio 1.48-1.52, ocular lobe ratio 0.88-0.89, EyL/EyD = 2.2-2.4; frontal grooves deep, slightly convergent to deepest portion just posterad clypeus, deeply extended onto clypeus; mandibles elongate, mandibular ratio 1.92; ligular anterior margin rounded to ligular seta, concave between setae, the two setae separated by one to two setal diameters; paraglossae thin, extended twice as far beyond ligular margin as their basal length to margin; antennae elongate, antennomere 9 length 2.25
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<normalizedToken id="D5BA61C0B95D7165F9DB6CE803FCC50D" originalValue="×">x</normalizedToken>
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maximal breadth; antennomere 3 glabrous except for apical ring of setae. Pronotum transversely ovoid, lateral margin evenly convex anterad base, hind angles absent though
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<pageBreakToken id="CF11B58C7943CE504B703F25B8F904C9" pageId="33" pageNumber="34" start="start">suggested</pageBreakToken>
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by presence of short peduncular collar at base of pronotum (Fig.
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<figureCitation id="85E77BA75A993B318EDEECB708162C6C" captionStart="Figure 20" captionStartId="F21" captionText="Figure 20. New Caledonian Mecyclothorax (Phacothorax) spp., dorsal view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Humboldt, 580 m); E, M. kanak (Mt. Humboldt, 630 m); F, M. kanak (Mt. Dzumac); G, M. kanak (Col de Yate); H, M. picdupinsensis." figureDoi="10.3897/dez.65.21000.figure20" httpUri="https://binary.pensoft.net/fig/179976" pageId="33" pageNumber="34">20A</figureCitation>
|
||
), MPW/BPW = 2.46-2.55, MPW/PL = 1.29-1.31; front angles slightly protruded, obtuse-rounded, APW/BPW = 1.39-1.48; median base unmargined basally, trapezoidally depressed relative to disc, the median longitudinal impression divided basally into two transverse impressions that isolate disc from base, each transverse basal impression terminated laterally in pitlike laterobasal depression; median longitudinal impression finely incised on disc, terminated anteriorly just anterad anterior transverse impression, a lenticular fovea at posterior juncture with basal transverse impressions; anterior transverse impression shallow, broad, but traceable to front angles; proepisternum separated from prosternum by a distinct groove both anteriorly and ventrally; smooth anteapical impression well developed laterally, continuous though shallower ventrally; prosternal process deeply and broadly depressed between procoxae, that depression extended 2/3 distance toward anterior prothoracic margin, the depression broadest at its anterior terminus. Elytra broadly ovoid, humeri sloped posteriorly laterad scutellum, humeral angle indicated only by slight change in curvature at juncture of narrow basal elytral groove and broader lateral marginal depression; MEW/EL = 0.79-0.94; all striae reduced on disc, stria 3 and 6 more evident, but intervening intervals at most slightly convex; only stria 8 evident apically, stria 2 present as very broad and shallow depression; elytra appressed and conjoined apically, the suture upraised at apex. Pterothoracic mesepisternal anterior furrow with 5 punctures in a curved line; mesosternal-mesepisternal suture complete (as in Fig.
|
||
<figureCitation id="95A90C5DE31AC1A8BF08E23D322ACF29" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Thoracic structures of Mecyclothorax spp. A-B. Lateral view of elytral epipleuron and meso-and metapleurites: A, M. laterorectus; B, M. fleutiauxi. C-D. Metathoracic flight wing: C, M. punctipennis, showing fully developed flight wing folded under elytron; D, M. lissus, stenopterous wing rudiment that extends beyond metanotum, rudimentary wing venation present at base of strap. Abbreviations include: mms, mesepisternal-mesosternal suture; msp, mesepimeron; mst, mesepisternum; mtp, metepimeron; mtt, metepisternum." figureDoi="10.3897/dez.65.21000.figure3" httpUri="https://binary.pensoft.net/fig/179959" pageId="33" pageNumber="34">3A</figureCitation>
|
||
); metepisternum maximum width/lateral length = 1.08; metepisternal-metepimeral suture complete. Abdomen with broad crescent-shaped depression along suture between first and second ventrite, second ventrite only slightly depressed within crescent; suture between second and third ventrites reduced though traceable laterally; ventrites 2-6 with broad, shallow, linear plaques near lateral margin. Microsculpture of frons reduced, surface glossy, indistinct transversely stretched sculpticells evident on vertex; pronotal disc glossy, transverse-line microsculpture visible over portions of surface depending on direction of light reflection, trapezoidal median base with only indistinct indications of transverse microsculpture; elytra iridescent, disc with elongate transverse mesh mixed with transverse lines, elytral apex covered with elongate transverse-mesh microsculpture.
|
||
</paragraph>
|
||
<paragraph id="E69D8000B6F798D30B53D85E85F95A61" pageId="33" pageNumber="34">
|
||
<emphasis id="217F1547B6258E0A69FCC85F2DB7D55E" italics="true" pageId="33" pageNumber="34">Male genitalia</emphasis>
|
||
(n = 1). Antecostal margin of abdominal mediotergite IX broadly angulate, not extended (Fig.
|
||
<figureCitation id="CF8DABCC7C350CBDC4037C040D04D74D" captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="33" pageNumber="34">17J</figureCitation>
|
||
); right paramere narrow basally, evenly narrowed to apex, with 5 setae along ventral margin and two short apical setae (Fig.
|
||
<figureCitation id="812304DBDEA090CFCE0E47E94BB7199B" captionStart="Figure 18" captionStartId="F19" captionText="Figure 18. Paired left (above) and right (below) parameres of Mecyclothorax (Phacothorax) spp. (ectal view): A, M. plurisetosus; B, M. megalovatulus; C, M. octavius; D, M. laterovatulus; E, M. manautei; F, M. paniensis; G, M. mouensis." figureDoi="10.3897/dez.65.21000.figure18" httpUri="https://binary.pensoft.net/fig/179974" pageId="33" pageNumber="34">18E</figureCitation>
|
||
); left paramere broad basally, narrowed to a dorsally twisted apex, parameral apex flexibly articulating with base, two elongate apical setae present; aedeagal median lobe robust, broad dorsoventrally, with apex distinctly down-curved relative to the expanded shaft and flattened into an apical face (Fig.
|
||
<figureCitation id="9DCF9790B2C0007B2E5543E5E8C49B58" captionStart="Figure 17" captionStartId="F18" captionText="Figure 17. Male aedeagal median lobe and parameres, and ring sclerite-mediotergite plus antecostal margin, tergite IX-of Mecyclothorax (Phacothorax) spp.: A-B, M. plurisetosus, right view, dorsal view in situ (Aoupinie); C-D, M. megalovatulus, right view, dorsal view of righ sclerite (Mt. Panie); E-F, M. octavius, right view, dorsal view in situ (Me Maoya); G-H, M. laterovatulus, right view, dorsal view in situ (Aoupinie); I-J, M. manautei, right view, dorsal view in situ (Mt. Humboldt, 1400 m); K-L, M. paniensis, right view, dorsal view in situ (Mt. Panie); M-N, M. mouensis, right view, dorsal view in situ (Mt. Mou). See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure17" httpUri="https://binary.pensoft.net/fig/179973" pageId="33" pageNumber="34">17I</figureCitation>
|
||
); male aedeagal internal sac with short flagellum and flagellar sheath present, a dorsal plate dorsad and basad flagellar complex, and a large dorsoapical field of macrospicules.
|
||
</paragraph>
|
||
<paragraph id="2D0D9CA24AFB341ABFF56072FB0067AD" pageId="33" pageNumber="34">
|
||
<emphasis id="DB257554C8E7A378BC22EB439784969A" italics="true" pageId="33" pageNumber="34">Female reproductive tract</emphasis>
|
||
(n = 1). Bursa copulatrix length slightly greater than circumference, its surface thin, translucent, only slightly wrinkled (Fig.
|
||
<figureCitation id="B9D9B861985FC974522C7065AFE10C6F" captionStart="Figure 23" captionStartId="F24" captionText="Figure 23. Female reproductive tract, gonocoxites and associated laterotergites, Mecyclothorax (Phacothorax) spp: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak (Mt. Dzumac; common oviduct artificially darkened to show configuration); E, M. kanak (Pic du Grand Kaori); F, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure23" httpUri="https://binary.pensoft.net/fig/179980" pageId="33" pageNumber="34">23A</figureCitation>
|
||
); spermathecal duct entering near bursa-common oviduct juncture with duct oriented toward right side of bursa, duct length about 1.25
|
||
<normalizedToken id="D6FB6243FDD01BD7395E8C20CA91F5A0" originalValue="×">x</normalizedToken>
|
||
length of spermathecal reservoir; laminar helminthoid sclerite with rounded apex present at base of spermathecal duct; spermatheca fusiform on narrow duct, spermathecal gland duct entering at base of spermathecal reservoir; ligular apophysis present near base of common oviduct; basal gonocoxite 1 with apical fringe of two setae laterally-a smaller seta may be present medially-and several small setae near apex of medial margin (Fig.
|
||
<figureCitation id="C0D6FEE525D45024BF4568A49C94D1C2" captionStart="Figure 24" captionStartId="F25" captionText="Figure 24. Left female gonocoxa of Mecyclothorax (Phacothorax) spp., ventral view: A, M. manautei; B, M. paniensis; C, M. mouensis; D, M. kanak; E, M. picdupinsensis. See Table 2 for abbreviations." figureDoi="10.3897/dez.65.21000.figure24" httpUri="https://binary.pensoft.net/fig/179981" pageId="33" pageNumber="34">24A</figureCitation>
|
||
); gonocoxite 2 moderately broad basally, basal width 0.6
|
||
<normalizedToken id="CE3B90A76ABFEE206E96C6D7E5BCD836" originalValue="×">x</normalizedToken>
|
||
medial length; two gracile lateral ensiform setae of moderate length present.
|
||
</paragraph>
|
||
<paragraph id="DE130298285780BC477ED11872359ADA" pageId="33" pageNumber="34">
|
||
<emphasis id="7A9EBDD2A622C3E9896B9AFEB2763AC6" italics="true" pageId="33" pageNumber="34">Types</emphasis>
|
||
- Holotype male (MNHN): NEW CALEDONIA 11138 / 21°53
|
||
<normalizedToken id="49759526F267899B57C93EAC242DE84D" originalValue="’SX166°24’">
|
||
'SX
|
||
<geoCoordinate id="811C5A83EDC6C014D946C658E8041406" degrees="166" direction="east" minutes="24" orientation="longitude" precision="925" value="166.4">166°24'E</geoCoordinate>
|
||
</normalizedToken>
|
||
.1400m. / Mt Humboldt, moss forest. / 6-7Nov2002. Monteith & / Burwell.pyreth,trees&logs // QUEENSLAND / MUSEUM LOAN / DATE: Nov. 2003 / No. LEN-1686 (green label) //
|
||
<emphasis id="32938211B31E86FB6127DCD41DED4AAE" italics="true" pageId="33" pageNumber="34">
|
||
New Caledonia
|
||
<taxonomicName id="63D70B859601241C45ED2E4920C1C464" authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
|
||
</emphasis>
|
||
revision / measured specimen 1 / J.K.
|
||
<bibRefCitation id="95438A34A38F8697EDF2FF0B18D55380" DOI="https://doi.org/10.3897/dez.63.10241" author="Liebherr, JK" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="51" pageNumber="52" pagination="211 - 270" refId="B61" refString="Liebherr, JK, 2016. Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia. Deutsche Entomologische Zeitschrift 63: 211 - 270, DOI: https://doi.org/10.3897/dez.63.10241" title="Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia." url="https://doi.org/10.3897/dez.63.10241" volume="63" year="2016">Liebherr 2016</bibRefCitation>
|
||
// HOLOTYPE /
|
||
<taxonomicName id="8BD7B17765B4F758B7878751DC7674F9" authorityName="Sharp" authorityYear="1903" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
|
||
/
|
||
<taxonomicName id="56A43A8F7782F2C087437258F84B2FFE" authorityName="Liebherr" authorityYear="2018" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax manautei" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="manautei">Mecyclothorax manautei</taxonomicName>
|
||
/ J.K.Liebherr 2017 (black-bordered red label).
|
||
</paragraph>
|
||
<paragraph id="C317C0F164E626136D307F6CE22534BE" pageId="33" pageNumber="34">
|
||
Paratypes (3 specimens). NEW CALEDONIA: Mt. Humboldt, moss forest, 1400 m el.,
|
||
<geoCoordinate id="1F69882C04D860834C6D67A63D65CED5" degrees="21" direction="south" minutes="53" orientation="latitude" precision="925" value="-21.883333">21°53'S</geoCoordinate>
|
||
<geoCoordinate id="85091407ABA3021BA5E786B9BD583AF8" degrees="166" direction="east" minutes="24" orientation="longitude" precision="925" value="166.4">166°24'E</geoCoordinate>
|
||
, pyrethrum trees & logs, 06-07-xi-2002, lot 11138, Monteith/Burwell (QMB, 2), beyond summit, 1500 m el.,
|
||
<geoCoordinate id="4ED26E8EC9E2754B4162377FB97E7867" degrees="21" direction="south" minutes="53" orientation="latitude" precision="925" value="-21.883333">21°53'S</geoCoordinate>
|
||
<geoCoordinate id="778032D255FA40DC81B4226CB6949D64" degrees="166" direction="east" minutes="25" orientation="longitude" precision="925" value="166.41667">166°25'E</geoCoordinate>
|
||
, 07-xi-2002, lot 11122, Burwell (QMB, 1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DC0584F220102AF4BF77566FB4153514" pageId="33" pageNumber="34" type="etymology">
|
||
<paragraph id="ABD4BFFE5084681ED8C2E909DE0DD66A" pageId="33" pageNumber="34">Etymology.</paragraph>
|
||
<paragraph id="A0B8B705B50DAB38319AB25172A1E22A" pageId="33" pageNumber="34">
|
||
This species epithet is a patronym honoring Joseph
|
||
<normalizedToken id="53B9F3203F613BDD7DF101D40D76FE6F" originalValue="Manauté">Manaute</normalizedToken>
|
||
, Directeur du Parc Provincial de la
|
||
<normalizedToken id="37F95CADF11C64010DD5F50447F367F6" originalValue="Rivière">Riviere</normalizedToken>
|
||
Bleue chez Province Sud, who provided helicopter support for the Queensland Museum expedition to Mt. Humboldt, allowing Geoff Monteith and Chris Burwell to collect the type series of this species as well as other interesting and important taxa (
|
||
<bibRefCitation id="4FAAB2364EB0F8B39B1A0EF84B58E51C" author="Reid, CAM" journalOrPublisher="Journal of Evolutionary Biology" pageId="52" pageNumber="53" refId="B90" refString="Reid, CAM, Smith, KI, 2004. A new genus and first record of Chrysomelinae from New Caledonia (Coleoptera: Chrysomelidae Memoirs of the Queensland Museum 49: 705-711." title="A new genus and first record of Chrysomelinae from New Caledonia (Coleoptera: Chrysomelidae Memoirs of the Queensland Museum 49: 705 - 711." year="2004">Reid and Smith 2004</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="06F72211BE7847E7799E2E4E8AC74F7F" pageId="33" pageNumber="34" type="distribution">
|
||
<paragraph id="0FB8BB3185B86C01E2DB8BDECC3457A0" pageId="33" pageNumber="34">Distribution and habitat.</paragraph>
|
||
<paragraph id="F919A12358724FE99AA2DB20D922AB25" pageId="33" pageNumber="34">
|
||
The species is known only from elevations 1400-1500 on Mt. Humboldt (Fig.
|
||
<figureCitation id="731F79314BE253F5DD06C0B40CD09185" captionStart="Figure 25" captionStartId="F26" captionText="Figure 25. Geographical distributions of Mecyclothorax (Phacothorax) spp. (asterisked locality attributed to M. mouensis based on non-type female specimen)." figureDoi="10.3897/dez.65.21000.figure25" httpUri="https://binary.pensoft.net/fig/179982" pageId="33" pageNumber="34">25</figureCitation>
|
||
). The four specimens were all collected via application of pyrethrin spray to tree trunks and downed logs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |