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<document id="DF50AEFFA06821F5736065296F1DBEF1" ID-DOI="10.11646/zootaxa.5470.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="12189689" ID-ZooBank="6A4270C2-D3F9-404F-91E7-4A73F2A99AE4" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718891908847" checkinUser="plazi" docAuthor="Harzhauser, Mathias, Landau, Bernard M. &amp; Vermeij, Geerat J." docDate="2024" docId="36508782FFF4DB1A328871BF0B2CFCC9" docLanguage="en" docName="zootaxa.5470.1.1.pdf" docOrigin="Zootaxa 5470 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5470.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Pseudofusus palatinus" docType="treatment" docVersion="2" lastPageNumber="54" masterDocId="CA69FFFAFFC0DB2F321F745D0A6FFFF0" masterDocTitle="The Dolicholatiridae and Fasciolariidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea" masterLastPageNumber="92" masterPageNumber="1" pageNumber="53" updateTime="1718893469852" updateUser="ExternalLinkService">
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<mods:title id="96D12BA61550C149FC62A6FE10C24FB7">The Dolicholatiridae and Fasciolariidae (Gastropoda, Buccinoidea) of the Miocene Paratethys Sea</mods:title>
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<mods:affiliation id="D0E4BDE0CE7603B4518F16919652DB3F">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, Netherlands; Instituto Dom Luiz da Universidade de Lisboa, Campo Grande, 1749 - 016 Lisboa, Portugal; and International Health Centres, Av. Infante de Henrique 7, Areias São João, P- 8200 Albufeira, Portugal. bernardmlandau @ gmail. com https: // orcid. org / 0000 - 0002 - 7768 - 8494</mods:affiliation>
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<mods:affiliation id="AF43875FB92A0C3EF7E82EDADC54804B">Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at https: // orcid. org / 0000 - 0002 - 4471 - 6655 &amp; Department of Geology, University of California at Davis, One Shields Avenue, Davis Ca 95616, USA. gjvermeij @ ucdavis. edu; https: // orcid. org / 0000 - 0002 - 6450 - 5687 &amp; Natural History Museum Vienna, Burgring 7, 1010 Vienna, Austria. mathias. harzhauser @ nhm-wien. ac. at https: // orcid. org / 0000 - 0002 - 4471 - 6655</mods:affiliation>
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<heading id="E50E81F8FFF4DB1B328871BF0804FA0C" bold="true" box="[151,619,1506,1533]" fontSize="11" level="2" pageId="52" pageNumber="53" reason="4">
<taxonomicName id="79F94D17FFF4DB1B328871BF0804FA0C" authority="(Strausz, 1954)" baseAuthorityName="Strausz" baseAuthorityYear="1954" box="[151,619,1506,1533]" class="Gastropoda" family="Fasciolariidae" genus="Pseudofusus" kingdom="Animalia" order="Neogastropoda" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="palatinus">
<emphasis id="8C8DEA86FFF4DB1B328871BF0804FA0C" bold="true" box="[151,619,1506,1533]" pageId="52" pageNumber="53">
<emphasis id="8C8DEA86FFF4DB1B328871BF0BC7FA0D" bold="true" box="[151,424,1506,1533]" italics="true" pageId="52" pageNumber="53">Pseudofusus palatinus</emphasis>
(
<bibRefCitation id="DA684B65FFF4DB1B33A771BF080CFA0C" author="Strausz, L." box="[440,611,1506,1532]" pageId="52" pageNumber="53" pagination="1 - 150" refId="ref78528" refString="Strausz, L. (1954) Varpalotai felso-mediterran csigak. Geologica Hungarica, 25, 1 - 150. [https: // epa. oszk. hu / 02900 / 02989 / 00022 / pdf / EPA 02989 _ geologica _ hungarica _ ser _ paleo _ 25 _ 1954. pdf]" type="journal article" year="1954">Strausz, 1954</bibRefCitation>
)
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<figureCitation id="26C22A11FFF4DB1B3288725A0A92F9D1" box="[151,253,1543,1569]" captionStart="FIGURE 21" captionStartId="51.[151,250,1976,2000]" captionTargetBox="[151,1436,1345,1952]" captionTargetId="figure-534@51.[151,1436,1345,1952]" captionTargetPageId="51" captionText="FIGURE 21. A1A2. Pseudofusus balteus nov. sp., NHMW 2023/0370/0001, holotype, Letkés (Hungary). B1B3. Pseudofusus palatinus (Strausz, 1954), neotype, Inv.55.78.1, Bakony Natural History Museum (Rirc, Hungary), Várpalota (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.12189742" httpUri="https://zenodo.org/record/12189742/files/figure.png" pageId="52" pageNumber="53">Figs 21B</figureCitation>
<figureCitation id="26C22A11FFF4DB1B32E272440B4AF9D1" box="[253,293,1543,1575]" captionStart="FIGURE 1" captionStartId="5.[151,250,1655,1679]" captionTargetBox="[232,1343,308,1621]" captionTargetId="figure-85@5.[232,1355,308,1621]" captionTargetPageId="5" captionText="FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width, S: siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.12189691" httpUri="https://zenodo.org/record/12189691/files/figure.png" pageId="52" pageNumber="53">
<subScript id="227D34D1FFF4DB1B32E272440B6AF9D7" attach="left" box="[253,261,1561,1575]" fontSize="6" pageId="52" pageNumber="53">1</subScript>
B
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<figureCitation id="26C22A11FFF4DB1B333972440B41F9D7" box="[294,302,1561,1575]" captionStart="FIGURE 3" captionStartId="9.[151,250,1124,1148]" captionTargetBox="[346,1241,181,1100]" captionTargetId="figure-26@9.[346,1241,181,1100]" captionTargetPageId="9" captionText="FIGURE 3. A1A2. Dulaiania pleurotomoides (Hoernes &amp; Auinger 1890), holotype, NHMW 1855/0043/0041." figureDoi="http://doi.org/10.5281/zenodo.12189695" httpUri="https://zenodo.org/record/12189695/files/figure.png" pageId="52" pageNumber="53">
<subScript id="227D34D1FFF4DB1B333972440B41F9D7" attach="right" box="[294,302,1561,1575]" fontSize="6" pageId="52" pageNumber="53">3</subScript>
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</paragraph>
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<paragraph id="BE463694FFF4DB1B328872110E44F993" blockId="52.[151,1067,1611,1697]" box="[151,1067,1611,1635]" pageId="52" pageNumber="53">
<treatmentCitationGroup id="9EE911BAFFF4DB1B328872110E44F993" box="[151,1067,1611,1635]" pageId="52" pageNumber="53">
<emphasis id="8C8DEA86FFF4DB1B328872110ACBF993" box="[151,164,1612,1635]" italics="true" pageId="52" pageNumber="53">*</emphasis>
<taxonomicName id="79F94D17FFF4DB1B32B572110829F993" ID-CoL="8QFZ2" authority="Harzhauser &amp; Landau &amp; Vermeij, 2024" authorityName="Harzhauser &amp; Landau &amp; Vermeij" authorityYear="2024" box="[170,582,1611,1635]" class="Gastropoda" family="Fasciolariidae" genus="Fusus" kingdom="Animalia" order="Neogastropoda" pageId="52" pageNumber="53" phylum="Mollusca" rank="variety" species="clavatus" status="var. nov." subGenus="Streptochetus" variety="palatinus">
<emphasis id="8C8DEA86FFF4DB1B32B572110A88F993" box="[170,231,1612,1635]" italics="true" pageId="52" pageNumber="53">Fusus</emphasis>
(
<emphasis id="8C8DEA86FFF4DB1B32EA72160B11F993" box="[245,382,1611,1635]" italics="true" pageId="52" pageNumber="53">Streptochetus</emphasis>
)
<emphasis id="8C8DEA86FFF4DB1B339272160829F993" box="[397,582,1611,1635]" italics="true" pageId="52" pageNumber="53">clavatus palatinus</emphasis>
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<treatmentCitation id="3F581085FFF4DB1B30A372160907F993" author="Strausz, L." box="[700,872,1611,1635]" page="31" pageId="52" pageNumber="53" year="1954">
<bibRefCitation id="DA684B65FFF4DB1B30A372160907F993" author="Strausz, L." box="[700,872,1611,1635]" pageId="52" pageNumber="53" pagination="1 - 150" refId="ref78528" refString="Strausz, L. (1954) Varpalotai felso-mediterran csigak. Geologica Hungarica, 25, 1 - 150. [https: // epa. oszk. hu / 02900 / 02989 / 00022 / pdf / EPA 02989 _ geologica _ hungarica _ ser _ paleo _ 25 _ 1954. pdf]" type="journal article" year="1954">Strausz 1954: 31</bibRefCitation>
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, pl. 4, figs 100ab.
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</paragraph>
<paragraph id="BE463694FFF4DB1B328872360E76F972" blockId="52.[151,1067,1611,1697]" box="[151,1049,1642,1666]" pageId="52" pageNumber="53">
<treatmentCitationGroup id="9EE911BAFFF4DB1B328872360E76F972" box="[151,1049,1642,1666]" pageId="52" pageNumber="53">
<taxonomicName id="79F94D17FFF4DB1B3288723608F4F972" ID-CoL="8QFZ2" authority="Strausz, 1954" authorityName="Strausz" authorityYear="1954" box="[151,667,1642,1666]" class="Gastropoda" family="Fasciolariidae" genus="Fusus" kingdom="Animalia" order="Neogastropoda" pageId="52" pageNumber="53" phylum="Mollusca" rank="subSpecies" species="lamellosus" subGenus="Aptyxis" subSpecies="palatinus">
<emphasis id="8C8DEA86FFF4DB1B328872360ABBF972" box="[151,212,1643,1666]" italics="true" pageId="52" pageNumber="53">Fusus</emphasis>
(
<emphasis id="8C8DEA86FFF4DB1B32FD72370B42F971" box="[226,301,1642,1665]" italics="true" pageId="52" pageNumber="53">Aptyxis</emphasis>
)
<emphasis id="8C8DEA86FFF4DB1B332372370862F972" box="[316,525,1642,1666]" italics="true" pageId="52" pageNumber="53">lamellosus palatinus</emphasis>
<bibRefCitation id="DA684B65FFF4DB1B300C723708F4F972" author="Strausz, L." box="[531,667,1642,1666]" pageId="52" pageNumber="53" pagination="1 - 150" refId="ref78528" refString="Strausz, L. (1954) Varpalotai felso-mediterran csigak. Geologica Hungarica, 25, 1 - 150. [https: // epa. oszk. hu / 02900 / 02989 / 00022 / pdf / EPA 02989 _ geologica _ hungarica _ ser _ paleo _ 25 _ 1954. pdf]" type="journal article" year="1954">Strausz, 1954</bibRefCitation>
</taxonomicName>
<treatmentCitation id="3F581085FFF4DB1B30A972370900F972" author="Strausz, L." box="[694,879,1642,1666]" page="345" pageId="52" pageNumber="53" year="1966">
<bibRefCitation id="DA684B65FFF4DB1B30A972370900F972" author="Strausz, L." box="[694,879,1642,1666]" pageId="52" pageNumber="53" refId="ref78664" refString="Strausz, L. (1966) Die miozan-mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp." type="book" year="1966">Strausz 1966: 345</bibRefCitation>
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, pl. 27, figs 12.
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</paragraph>
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<taxonomicName id="79F94D17FFF4DB1B328872D40BF3F951" ID-CoL="8QGH9" authority="(Strausz)" baseAuthorityName="Strausz" baseAuthorityYear="1954" box="[151,412,1673,1697]" class="Gastropoda" family="Fasciolariidae" genus="Aptyxis" kingdom="Animalia" order="Neogastropoda" pageId="52" pageNumber="53" phylum="Mollusca" rank="species" species="palatina">
<emphasis id="8C8DEA86FFF4DB1B328872D40B53F951" box="[151,316,1673,1697]" italics="true" pageId="52" pageNumber="53">Aptyxis palatina</emphasis>
(Strausz)
</taxonomicName>
<treatmentCitation id="3F581085FFF4DB1B33A872D4080AF950" author="Kovacs, Z." box="[439,613,1672,1697]" page="95" pageId="52" pageNumber="53" year="2022">
<bibRefCitation id="DA684B65FFF4DB1B33A872D4080AF950" author="Kovacs, Z." box="[439,613,1672,1697]" pageId="52" pageNumber="53" pagination="65 - 99" refId="ref71724" refString="Kovacs, Z. (2022) Middle Miocene Buccinoidea (Neogastropoda) assemblage from the Faget Basin (Romania) in the collection of the Hungarian Natural History Museum, Budapest. Fragmenta Palaeontologica Hungarica, 37, 65 - 99. https: // doi. org / 10.17111 / FragmPalHung. 2021.37.65" type="journal article" year="2022">Kovács 2022: 95</bibRefCitation>
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, figs 98100.
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</paragraph>
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<emphasis id="8C8DEA86FFF4DB1B328872920B2AF919" bold="true" box="[151,325,1743,1769]" pageId="52" pageNumber="53">Type material.</emphasis>
<materialsCitation id="0E913CC9FFF4DB1B3352729209DEF8FD" collectionCode="MD" country="Hungary" county="Varpalota" location="Bakony Natural History Museum" municipality="Rirc" pageId="52" pageNumber="53" specimenCount="1" typeStatus="neotype">
<typeStatus id="61428836FFF4DB1B335272920BC3F919" box="[333,428,1743,1769]" pageId="52" pageNumber="53" type="neotype">Neotype</typeStatus>
(designated by
<bibRefCitation id="DA684B65FFF4DB1B30407292094EF91A" author="Kovacs, Z." box="[607,801,1743,1770]" pageId="52" pageNumber="53" pagination="65 - 99" refId="ref71724" refString="Kovacs, Z. (2022) Middle Miocene Buccinoidea (Neogastropoda) assemblage from the Faget Basin (Romania) in the collection of the Hungarian Natural History Museum, Budapest. Fragmenta Palaeontologica Hungarica, 37, 65 - 99. https: // doi. org / 10.17111 / FragmPalHung. 2021.37.65" type="journal article" year="2022">Kovács 2022: 95</bibRefCitation>
), Inv.55.78.1, SL:
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,
<collectionCode id="D8E8AE51FFF4DB1B364F72920EEEF919" box="[1104,1153,1743,1769]" country="Germany" name="Museum Donaueschingen" pageId="52" pageNumber="53">MD</collectionCode>
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,
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(
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<collectingCountry id="C6EE7604FFF4DB1B33BC72AE0866F8FD" box="[419,521,1779,1805]" name="Hungary" pageId="52" pageNumber="53">Hungary</collectingCountry>
),
<collectingCounty id="57274E18FFF4DB1B300372AE08E5F8FD" box="[540,650,1779,1805]" pageId="52" pageNumber="53">Várpalota</collectingCounty>
(
<collectingCountry id="C6EE7604FFF4DB1B308572AE096FF8FD" box="[666,768,1779,1805]" name="Hungary" pageId="52" pageNumber="53">Hungary</collectingCountry>
),
<figureCitation id="26C22A11FFF4DB1B310B72AE0915F8FD" box="[788,890,1779,1805]" captionStart="FIGURE 21" captionStartId="51.[151,250,1976,2000]" captionTargetBox="[151,1436,1345,1952]" captionTargetId="figure-534@51.[151,1436,1345,1952]" captionTargetPageId="51" captionText="FIGURE 21. A1A2. Pseudofusus balteus nov. sp., NHMW 2023/0370/0001, holotype, Letkés (Hungary). B1B3. Pseudofusus palatinus (Strausz, 1954), neotype, Inv.55.78.1, Bakony Natural History Museum (Rirc, Hungary), Várpalota (Hungary)." figureDoi="http://doi.org/10.5281/zenodo.12189742" httpUri="https://zenodo.org/record/12189742/files/figure.png" pageId="52" pageNumber="53">Figs 21B</figureCitation>
<figureCitation id="26C22A11FFF4DB1B3166735809CDF8FD" box="[889,930,1779,1811]" captionStart="FIGURE 1" captionStartId="5.[151,250,1655,1679]" captionTargetBox="[232,1343,308,1621]" captionTargetId="figure-85@5.[232,1355,308,1621]" captionTargetPageId="5" captionText="FIGURE 1. Measurements for shell morphology; AA: apical angle, SL: shell length, MD: maximum diameter, LWH: last whorl height, AH: aperture height, AL: aperture length, AW: aperture width, S: siphonal canal." figureDoi="http://doi.org/10.5281/zenodo.12189691" httpUri="https://zenodo.org/record/12189691/files/figure.png" pageId="52" pageNumber="53">
<subScript id="227D34D1FFF4DB1B3166735809EEF8E3" attach="left" box="[889,897,1797,1811]" fontSize="6" pageId="52" pageNumber="53">1</subScript>
B
</figureCitation>
<figureCitation id="26C22A11FFF4DB1B31BD735809C5F8E3" box="[930,938,1797,1811]" captionStart="FIGURE 3" captionStartId="9.[151,250,1124,1148]" captionTargetBox="[346,1241,181,1100]" captionTargetId="figure-26@9.[346,1241,181,1100]" captionTargetPageId="9" captionText="FIGURE 3. A1A2. Dulaiania pleurotomoides (Hoernes &amp; Auinger 1890), holotype, NHMW 1855/0043/0041." figureDoi="http://doi.org/10.5281/zenodo.12189695" httpUri="https://zenodo.org/record/12189695/files/figure.png" pageId="52" pageNumber="53">
<subScript id="227D34D1FFF4DB1B31BD735809C5F8E3" attach="left" box="[930,938,1797,1811]" fontSize="6" pageId="52" pageNumber="53">3</subScript>
</figureCitation>
.
</materialsCitation>
</paragraph>
<paragraph id="BE463694FFF4DB1A32D8734A09CBFEED" blockId="52.[151,1437,1743,2021]" lastBlockId="53.[151,1437,151,825]" lastPageId="53" lastPageNumber="54" pageId="52" pageNumber="53">
<emphasis id="8C8DEA86FFF4DB1B32D8734A0BD1F8C1" bold="true" box="[199,446,1815,1841]" pageId="52" pageNumber="53">Revised description.</emphasis>
Medium-sized, slender fusiform shell of eight teleoconch whorls; apical angle 33°. Protoconch paucispiral of two whorls. Last protoconch whorl weakly convex with orthocline axial riblets close to transition to teleoconch. Early teleoconch whorls moderately convex with deeply incised suture. Later whorls strongly convex with weak subsutural cord, periphery placed mid-whorl, suture undulating. Broad, rounded, widely spaced axial ribs crossed by prominent spiral cords (six on penultimate whorl). Secondary spiral cords subobsolete. Last whorl attaining 60% of total height, narrow subsutural concavity with delicately scabrose growth lines, evenly convex periphery, rapidly constricting at base; base bearing numerous prominent spiral cords; fasciole indistinct. Aperture moderately wide, pyriform. Columellar callus forming thin, narrow rim. Columella broadly and weakly excavated, smooth. Anal canal indistinct. Outer lip with weakly thickened terminal varix with about eight long lirae extending deep within aperture. Siphonal canal long, narrow, straight.
</paragraph>
</subSubSection>
<subSubSection id="F6E3651FFFF5DB1A32D8757B09ABFD01" pageId="53" pageNumber="54" type="discussion">
<paragraph id="BE463694FFF5DB1A32D8757B087CFE21" blockId="53.[151,1437,151,825]" pageId="53" pageNumber="54">
<emphasis id="8C8DEA86FFF5DB1A32D8757B0B23FEB0" bold="true" box="[199,332,294,320]" pageId="53" pageNumber="54">Discussion.</emphasis>
This species was placed in the monotypic extant
<taxonomicName id="79F94D17FFF5DB1A317E757A0E32FEB1" authority="Troschel, 1868" authorityName="Troschel" authorityYear="1868" box="[865,1117,295,321]" class="Gastropoda" family="Fasciolariidae" genus="Aptyxis" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="8C8DEA86FFF5DB1A317E757A09DCFEB1" box="[865,947,295,321]" italics="true" pageId="53" pageNumber="54">Aptyxis</emphasis>
Troschel, 1868
</taxonomicName>
by
<bibRefCitation id="DA684B65FFF5DB1A369D757A0F49FEB1" author="Strausz, L." box="[1154,1318,295,321]" pageId="53" pageNumber="54" refId="ref78664" refString="Strausz, L. (1966) Die miozan-mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp." type="book" year="1966">Strausz (1966)</bibRefCitation>
,
<bibRefCitation id="DA684B65FFF5DB1A3730757A0B6AFE95" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." pageId="53" pageNumber="54" pagination="3 - 576" refId="ref72587" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the Middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576." type="journal article" year="2013">
Landau
<emphasis id="8C8DEA86FFF5DB1A3797757A0ADBFE95" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2013)
</bibRefCitation>
and
<bibRefCitation id="DA684B65FFF5DB1A332675160B8EFE95" author="Kovacs, Z." box="[313,481,331,357]" pageId="53" pageNumber="54" pagination="65 - 99" refId="ref71724" refString="Kovacs, Z. (2022) Middle Miocene Buccinoidea (Neogastropoda) assemblage from the Faget Basin (Romania) in the collection of the Hungarian Natural History Museum, Budapest. Fragmenta Palaeontologica Hungarica, 37, 65 - 99. https: // doi. org / 10.17111 / FragmPalHung. 2021.37.65" type="journal article" year="2022">Kovács (2022)</bibRefCitation>
. The extant
<typeStatus id="61428836FFF5DB1A3079751608F9FE95" box="[614,662,331,357]" pageId="53" pageNumber="54">type</typeStatus>
species
<taxonomicName id="79F94D17FFF5DB1A30EC75160EE0FE95" authority="(Linnaeus, 1758)" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[755,1167,330,357]" class="Gastropoda" family="Fasciolariidae" genus="Aptyxis" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="syracusana">
<emphasis id="8C8DEA86FFF5DB1A30EC751609A4FE94" box="[755,971,331,357]" italics="true" pageId="53" pageNumber="54">Aptyxis syracusana</emphasis>
(
<bibRefCitation id="DA684B65FFF5DB1A31C775160EE9FE95" author="Linnaeus, C." box="[984,1158,330,357]" pageId="53" pageNumber="54" refId="ref72750" refString="Linnaeus, C. (1758) Systema naturae per regna tria naturae. Editio decima, reformata. Tomus 1. Laurentii Salvii, Holmiae, 823 pp. [https: // www. biodiversitylibrary. org / item / 10277 # page / 3 / mode / 1 up]" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
)
</taxonomicName>
from the Mediterranean Sea is characterized by a marked shoulder with broad subsutural ramp on which the axial ribs weaken distinctly (
<bibRefCitation id="DA684B65FFF5DB1A32BE75CE0B47FE5D" author="Russo, P." box="[161,296,402,429]" pageId="53" pageNumber="54" pagination="79 - 86" refId="ref76619" refString="Russo, P. (2015) On the systematic position of Murex syracusanus Linnaeus, 1758 (Gastropoda, Fasciolariidae) with revaluation of the genus Aptyxis. Bollettino Malacologico, 51, 79 - 86." type="journal article" year="2015">Russo 2015</bibRefCitation>
). This morphology is quite dissimilar with the Paratethyan species, which fits well within
<taxonomicName id="79F94D17FFF5DB1A370F75CE0FF3FE5D" authorityName="Monterosato" authorityYear="1884" box="[1296,1436,403,429]" class="Gastropoda" family="Fasciolariidae" genus="Pseudofusus" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="8C8DEA86FFF5DB1A370F75CE0FF3FE5D" box="[1296,1436,403,429]" italics="true" pageId="53" pageNumber="54">Pseudofusus</emphasis>
</taxonomicName>
as defined by
<bibRefCitation id="DA684B65FFF5DB1A332C75EA0860FE21" author="Fassio, G. &amp; Russo, P. &amp; Bonomolo, G. &amp; Fedosov, A. E. &amp; Modica, M. V. &amp; Nocella, E. &amp; Oliverio, M." box="[307,527,439,465]" pageId="53" pageNumber="54" pagination="623 - 636" refId="ref66732" refString="Fassio, G., Russo, P., Bonomolo, G., Fedosov, A. E., Modica, M. V., Nocella, E. &amp; Oliverio, M. (2022) A molecular framework for the systematics of the Mediterranean spindle-shells (Gastropoda, Neogastropoda, Fasciolariidae, Fusininae). Mediterranean Marine Science, 23, 623 - 636. https: // doi. org / 10.12681 / mms. 29935" type="journal article" year="2022">
Fassio
<emphasis id="8C8DEA86FFF5DB1A339D75EA0BD5FE21" box="[386,442,439,465]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
(2022)
</bibRefCitation>
.
</paragraph>
<paragraph id="BE463694FFF5DB1A32D875860921FD3D" blockId="53.[151,1437,151,825]" pageId="53" pageNumber="54">
<taxonomicName id="79F94D17FFF5DB1A32D875860940FE05" authority="sensu Landau et al. (2013)" authorityName="Landau" authorityYear="2013" box="[199,815,475,501]" class="Gastropoda" family="Fasciolariidae" genus="Pseudofusus" isUncertain="true" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="palatinus">
<emphasis id="8C8DEA86FFF5DB1A32D875860B3CFE05" box="[199,339,475,501]" italics="true" pageId="53" pageNumber="54">Pseudofusus</emphasis>
aff.
<emphasis id="8C8DEA86FFF5DB1A339575860856FE04" box="[394,569,475,501]" italics="true" pageId="53" pageNumber="54">palatinus sensu</emphasis>
<bibRefCitation id="DA684B65FFF5DB1A305D75860940FE05" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[578,815,475,501]" pageId="53" pageNumber="54" pagination="3 - 576" refId="ref72587" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the Middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576." type="journal article" year="2013">
Landau
<emphasis id="8C8DEA86FFF5DB1A3080758608BDFE05" box="[671,722,475,501]" italics="true" pageId="53" pageNumber="54">et al</emphasis>
. (2013)
</bibRefCitation>
</taxonomicName>
, from the Serravallian of the
<collectingRegion id="7C3DF876FFF5DB1A369D75860E86FE05" box="[1154,1257,475,501]" country="Turkey" name="Karaman" pageId="53" pageNumber="54">Karaman</collectingRegion>
Basin (
<collectingCountry id="C6EE7604FFF5DB1A375E75860FFAFE05" box="[1345,1429,475,501]" name="Turkey" pageId="53" pageNumber="54">Turkey</collectingCountry>
) is a closely related species but differs in having a lower spire (see
<bibRefCitation id="DA684B65FFF5DB1A318575A20E18FDE9" author="Landau, B. M. &amp; Harzhauser, M. &amp; Yslamodlu, Y. &amp; Marques da Silva, C." box="[922,1143,511,537]" pageId="53" pageNumber="54" pagination="3 - 576" refId="ref72587" refString="Landau, B. M., Harzhauser, M., Yslamodlu, Y. &amp; Marques da Silva, C. (2013) Systematics and palaeobiogeography of the gastropods of the Middle Miocene (Serravallian) Karaman Basin of Turkey. Cainozoic Research, 11 - 13, 3 - 576." type="journal article" year="2013">
Landau
<emphasis id="8C8DEA86FFF5DB1A31E875A20E5DFDE9" box="[1015,1074,511,537]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2013
</bibRefCitation>
: pl. 31, fig. 6, pl. 67, fig. 8).
<taxonomicName id="79F94D17FFF5DB1A32A1767E081EFDCD" authority="(Peyrot, 1938)" baseAuthorityName="Peyrot" baseAuthorityYear="1938" box="[190,625,547,573]" class="Gastropoda" family="Fasciolariidae" genus="Pseudofusus" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="ligerianus">
<emphasis id="8C8DEA86FFF5DB1A32A1767E0BABFDCD" box="[190,452,547,573]" italics="true" pageId="53" pageNumber="54">Pseudofusus ligerianus</emphasis>
(
<bibRefCitation id="DA684B65FFF5DB1A33CC767E0806FDCD" author="Peyrot, A." box="[467,617,547,573]" pageId="53" pageNumber="54" pagination="5 - 361" refId="ref75227" refString="Peyrot, A. (1938) Les Mollusques Testaces univalves des depots helvetiens du Bassin Ligerien. Catalogue critique, descriptif et illustre. Actes de la Societe Linneenne de Bordeaux, 89, 5 - 361. [https: // www. biodiversitylibrary. org / item / 247820 # page / 12 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 147213" type="journal article" year="1938">Peyrot, 1938</bibRefCitation>
)
</taxonomicName>
, from the Middle Miocene of the Northeastern Atlantic, has shouldered whorls, bears a higher number of spiral cords and has comparatively weaker axial ribs (see
<bibRefCitation id="DA684B65FFF5DB1A3684761A0F48FD91" author="Peyrot, A." box="[1179,1319,583,609]" pageId="53" pageNumber="54" pagination="5 - 361" refId="ref75227" refString="Peyrot, A. (1938) Les Mollusques Testaces univalves des depots helvetiens du Bassin Ligerien. Catalogue critique, descriptif et illustre. Actes de la Societe Linneenne de Bordeaux, 89, 5 - 361. [https: // www. biodiversitylibrary. org / item / 247820 # page / 12 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 147213" type="journal article" year="1938">Peyrot 1938</bibRefCitation>
: pl. 4, figs 13, 14; Glibert 1952: pl. 11, fig. 3).
<taxonomicName id="79F94D17FFF5DB1A302F763609BCFD75" authority="(Millet, 1864)" baseAuthorityName="Millet" baseAuthorityYear="1864" box="[560,979,619,645]" class="Gastropoda" family="Fasciolariidae" genus="Pseudofusus" kingdom="Animalia" order="Neogastropoda" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="omphalis">
<emphasis id="8C8DEA86FFF5DB1A302F76360944FD75" box="[560,811,619,645]" italics="true" pageId="53" pageNumber="54">Pseudofusus omphalis</emphasis>
(
<bibRefCitation id="DA684B65FFF5DB1A3124763609A4FD75" author="Millet de la Turtaudiere, P. - A." box="[827,971,619,645]" pageId="53" pageNumber="54" refId="ref73444" refString="Millet de la Turtaudiere, P. - A. (1864) Indicateur de Maine-et-Loire ou indications par commune de ce que chacune d'elles renferme, 1. Cosnier et Lachese, Angers, 754 pp." type="book" year="1864">Millet, 1864</bibRefCitation>
)
</taxonomicName>
, from the Late Miocene and Pliocene of the Northeastern Atlantic, differs in its stockier outline, lower spire and shorter siphonal canal (
<bibRefCitation id="DA684B65FFF5DB1A36A376D20ABCFD3D" author="Van Dingenen, F. &amp; Ceulemans, L. &amp; Landau, B. M." pageId="53" pageNumber="54" pagination="23 - 61" refId="ref79399" refString="Van Dingenen, F., Ceulemans, L. &amp; Landau, B. M. (2017) The lower Pliocene gastropods of Le Pigeon Blanc (Loire-Atlantique, Northwest France), 4. Neogastropoda (in part). Cainozoic Research, 17, 23 - 61." type="journal article" year="2017">
<collectingRegion id="7C3DF876FFF5DB1A36A376D20E85FD59" box="[1212,1258,655,681]" country="Turkey" name="Van" pageId="53" pageNumber="54">Van</collectingRegion>
Dingenen
<emphasis id="8C8DEA86FFF5DB1A377B76D20FF2FD59" box="[1380,1437,655,681]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2017
</bibRefCitation>
: pl. 2 fig. 4;
<bibRefCitation id="DA684B65FFF5DB1A337E76EE0858FD3D" author="Landau, B. M. &amp; Ceulemans, L. &amp; Van Dingenen, F." box="[353,567,691,717]" pageId="53" pageNumber="54" pagination="135 - 215" refId="ref72456" refString="Landau, B. M., Ceulemans, L. &amp; Van Dingenen, F. (2019) The upper Miocene gastropods of northwestern France, 4. Neogastropoda. Cainozoic Research, 19, 135 - 215." type="journal article" year="2019">
Landau
<emphasis id="8C8DEA86FFF5DB1A33A376EE0B9BFD3D" box="[444,500,691,717]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
2019
</bibRefCitation>
: p. 172, pl. 35, figs 18).
</paragraph>
<paragraph id="BE463694FFF5DB1A32D8768B09ABFD01" blockId="53.[151,1437,151,825]" box="[199,964,726,753]" pageId="53" pageNumber="54">
<emphasis id="8C8DEA86FFF5DB1A32D8768B0BC7FD00" bold="true" box="[199,424,726,752]" pageId="53" pageNumber="54">Paleoenvironment.</emphasis>
Shallow marine, inner neritic (own data, M.H.).
</paragraph>
</subSubSection>
<subSubSection id="F6E3651FFFF5DB1A32D876A70B2CFCC9" pageId="53" pageNumber="54" type="distribution">
<paragraph id="BE463694FFF5DB1A32D876A70B2CFCC9" blockId="53.[151,1437,151,825]" pageId="53" pageNumber="54">
<emphasis id="8C8DEA86FFF5DB1A32D876A7081AFCE5" bold="true" box="[199,629,762,789]" pageId="53" pageNumber="54">Distribution in Central Paratethys.</emphasis>
Badenian (Middle Miocene):
<emphasis id="8C8DEA86FFF5DB1A31C876A60EF2FCE4" box="[983,1181,763,788]" italics="true" pageId="53" pageNumber="54">Pannonian Basin</emphasis>
: Várpalota (
<collectingCountry id="C6EE7604FFF5DB1A372E76A60FFAFCE5" box="[1329,1429,763,789]" name="Hungary" pageId="53" pageNumber="54">Hungary</collectingCountry>
) (
<bibRefCitation id="DA684B65FFF5DB1A328077420B59FCC9" author="Strausz, L." box="[159,310,799,825]" pageId="53" pageNumber="54" refId="ref78664" refString="Strausz, L. (1966) Die miozan-mediterranen Gastropoden Ungarns. Akademiai Kiado, Budapest, 693 pp." type="book" year="1966">Strausz 1966</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>