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<document id="99587AD007D1E59AD393D03F4852BB9F" ID-DOI="10.5281/zenodo.5331202" ID-ISSN="0374-1036" ID-Zenodo-Dep="5331202" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630169194360" checkinUser="diego" docAuthor="Švec, Zdeněk" docDate="2012" docId="297B87BCA27CFFCAFED7665DFCF8AA8C" docLanguage="en" docName="ActaEntMusNatPra.2012.52.2.411-424.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Agathidium (Agathidium) fikaceki Švec 2012, sp. nov." docType="treatment" docVersion="3" lastPageNumber="414" masterDocId="D542FFC4A27DFFC9FFB9651CFFE7A90F" masterDocTitle="New Leiodinae (Coleoptera: Leiodidae) from India and Papua New Guinea" masterLastPageNumber="424" masterPageNumber="411" pageNumber="412" updateTime="1699172137447" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1432FAE0A89F1FD5E6E3E41D8207E297" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A53103163AA441F835C90A3F4718C7FB">
<mods:title id="EF592532A21A0E8DFE79D8A6709F8ABA">New Leiodinae (Coleoptera: Leiodidae) from India and Papua New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="FCBE89529F23645617E49245914D455B" type="personal">
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<mods:namePart id="21A6D71C7AB96E9395A6D9647DF49E44">Švec, Zdeněk</mods:namePart>
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<mods:typeOfResource id="13B521B335F1575B9CAF97CD41676E25">text</mods:typeOfResource>
<mods:relatedItem id="DE17D4F464A0CF1BAFCBFAF1FEA1EA5C" type="host">
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<mods:title id="984309D772463E8111EBA630B79D63B7">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="B6FB71E6681A48B22D5509A34898E1B3">
<mods:date id="E9B5D74E3BADA0B4315D5F581EF81458">2012</mods:date>
<mods:detail id="157BE2F8B33395F34B24BF223C5ADD4F" type="pubDate">
<mods:number id="C6B0F6F9BB646278473679CCB92D8F3D">2012-12-17</mods:number>
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<mods:number id="5E47D66B3B43CEE6862A2A3538E6E3CB">52</mods:number>
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<mods:number id="7491E5881B520513D7C9AF77683F8FA6">2</mods:number>
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<mods:classification id="322F5DE773F759A9364F0400D520F2C6">journal article</mods:classification>
<mods:identifier id="9FA3DDE6F97E184A0A296895E06BF84B" type="DOI">10.5281/zenodo.5331202</mods:identifier>
<mods:identifier id="6762AB3FECA352D63A51671777A88A7F" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="5DD83998A2884216BECBF777CDA138C7" type="Zenodo-Dep">5331202</mods:identifier>
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<treatment id="297B87BCA27CFFCAFED7665DFCF8AA8C" LSID="urn:lsid:plazi:treatment:297B87BCA27CFFCAFED7665DFCF8AA8C" httpUri="http://treatment.plazi.org/id/297B87BCA27CFFCAFED7665DFCF8AA8C" lastPageId="3" lastPageNumber="414" pageId="1" pageNumber="412">
<subSubSection id="E9C86521A27CFFC8FED7665DFCA3AA55" box="[366,836,833,858]" pageId="1" pageNumber="412" type="nomenclature">
<paragraph id="A16D36AAA27CFFC8FED7665DFCA3AA55" blockId="1.[366,836,833,858]" box="[366,836,833,858]" pageId="1" pageNumber="412">
<heading id="FA2581C6A27CFFC8FED7665DFCA3AA55" bold="true" box="[366,836,833,858]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="412" reason="3">
<emphasis id="93A6EAB8A27CFFC8FED7665DFCA3AA55" bold="true" box="[366,836,833,858]" pageId="1" pageNumber="412">
<taxonomicName id="66D24D29A27CFFC8FED7665DFD0FAA55" ID-CoL="5TPJD" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[366,744,833,858]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="412" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov." subGenus="Agathidium">
<emphasis id="93A6EAB8A27CFFC8FED7665DFE17AA55" bold="true" box="[366,496,833,858]" italics="true" pageId="1" pageNumber="412">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A27CFFC8FDB9665DFD65AA55" bold="true" box="[512,642,833,858]" italics="true" pageId="1" pageNumber="412">Agathidium</emphasis>
)
<emphasis id="93A6EAB8A27CFFC8FD2B665DFD0FAA55" bold="true" box="[658,744,833,858]" italics="true" pageId="1" pageNumber="412">fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A27CFFC8FD56665EFCA3AA55" box="[751,836,834,858]" pageId="1" pageNumber="412" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27CFFC8FDB3667CFD4FAA7B" box="[522,680,864,884]" pageId="1" pageNumber="412" type="description">
<paragraph id="A16D36AAA27CFFC8FDB3667CFD4FAA7B" blockId="1.[522,680,864,884]" box="[522,680,864,884]" pageId="1" pageNumber="412">
(
<figureCitation id="39E92A2FA27CFFC8FDA9667CFD85AA7B" box="[528,610,864,884]" captionStart="Figs" captionStartId="2.[152,191,830,849]" captionTargetBox="[180,1097,228,803]" captionTargetId="figure-234@2.[175,1128,220,806]" captionTargetPageId="2" captionText="Figs. 1–5. Genitalia of Agathidium species. 1–2 – aedeagus of Agathidium (Agathidium) fikaceki sp. nov. (1 – lateral view, 2 – apex in dorsal view). 3–5 – A. (Macroceble) cochleariforme sp. nov. (3 – aedeagus, lateral view; 4 – apex of aedeagus, dorsal view; 5 – spermatheca). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331206" httpUri="https://zenodo.org/record/5331206/files/figure.png" pageId="1" pageNumber="412">Figs. 1, 2</figureCitation>
,
<figureCitation id="39E92A2FA27CFFC8FDD2667DFD46AA7B" box="[619,673,865,884]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="1" pageNumber="412">18, 19</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27CFFC8FFC86696FE00AD2C" pageId="1" pageNumber="412" type="materials_examined">
<paragraph id="A16D36AAA27CFFC8FFC86696FF32AAB7" blockId="1.[113,1090,905,1059]" pageId="1" pageNumber="412">
<emphasis id="93A6EAB8A27CFFC8FFC86696FF01AA92" bold="true" box="[113,230,905,925]" pageId="1" pageNumber="412">
<typeStatus id="7E698808A27CFFC8FFC86696FF7AAA92" box="[113,157,906,925]" pageId="1" pageNumber="412">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D9C5763AA27CFFC8FF506696FEF1AA92" box="[233,278,906,925]" name="India" pageId="1" pageNumber="412">India</collectingCountry>
,
<collectingRegion id="6316F848A27CFFC8FEA46696FE9BAA92" box="[285,380,906,925]" country="India" name="Meghalaya" pageId="1" pageNumber="412">Meghalaya</collectingRegion>
,eastern Khasi Hills,
<quantity id="662A9B4FA27CFFC8FD9E6696FDBDAA92" box="[551,602,906,925]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="412" unit="km" value="11.0">11 km</quantity>
SW of Cherrapunjee,Laitkynsew,
<geoCoordinate id="C4E6506DA27CFFC8FCCE6696FC3FAA91" box="[887,984,905,926]" degrees="25" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="412" precision="15" seconds="48" value="25.213335">25º12′48″N</geoCoordinate>
<geoCoordinate id="C4E6506DA27CFFC8FC656696FBDAAA91" box="[988,1085,905,926]" degrees="91" direction="east" minutes="39" orientation="longitude" pageId="1" pageNumber="412" precision="15" seconds="48" value="91.66334">91º39′48″E</geoCoordinate>
,
<quantity id="662A9B4FA27CFFC8FFC866B9FF4FAAB7" box="[113,168,933,952]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="1" pageNumber="412" unit="m" value="735.0">735 m</quantity>
a.s.l.
</paragraph>
<paragraph id="A16D36AAA27CFFC8FFC866DCFE00AD2C" blockId="1.[113,1090,905,1059]" pageId="1" pageNumber="412">
<emphasis id="93A6EAB8A27CFFC8FFC866DCFF12AADC" bold="true" box="[113,245,959,979]" pageId="1" pageNumber="412">Type material.</emphasis>
<materialsCitation id="11BA3CF7A27CFFC8FF4066DCFC44AD07" collectingDate="2008-04-25" collectionCode="NMPC" collectorName="E Khasi Hills" country="India" elevation="735" latitude="25.213335" location="Meghalaya state" longLatPrecision="21" longitude="91.66334" pageId="1" pageNumber="412" specimenCount="1" stateProvince="Meghalaya" typeStatus="holotype">
<typeStatus id="7E698808A27CFFC8FF4066DCFEAAAADE" box="[249,333,960,979]" pageId="1" pageNumber="412" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="C7C3AE6FA27CFFC8FEC866DCFE54AADC" box="[369,435,960,979]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="412">NMPC</collectionCode>
): ‘
<collectingCountry id="D9C5763AA27CFFC8FE7A66DCFDE1AADC" box="[451,518,960,979]" name="India" pageId="1" pageNumber="412">INDIA</collectingCountry>
:
<collectingRegion id="6316F848A27CFFC8FDB766DCFD7FAADC" box="[526,664,960,979]" country="India" name="Meghalaya" pageId="1" pageNumber="412">Meghalaya state</collectingRegion>
(7+9) /
<collectorName id="0C27537CA27CFFC8FD6266DCFCA8AADC" box="[731,847,960,979]" pageId="1" pageNumber="412">E Khasi Hills</collectorName>
,
<quantity id="662A9B4FA27CFFC8FCEF66DCFC6DAADC" box="[854,906,960,979]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="1" pageNumber="412" unit="km" value="11.0">11 km</quantity>
SW Cherra- / punjee, Laitkynsew
<date id="D56C106AA27CFFC8FF6466C6FED0AAE2" box="[221,311,986,1005]" pageId="1" pageNumber="412" value="2008-04-25">
<collectingDate id="C528E982A27CFFC8FF6466C6FED0AAE2" box="[221,311,986,1005]" pageId="1" pageNumber="412" value="2008-04-25">25.iv.2008</collectingDate>
</date>
/
<geoCoordinate id="C4E6506DA27CFFC8FEF566C6FE48AAE1" box="[332,431,985,1006]" degrees="25" direction="north" minutes="12" orientation="latitude" pageId="1" pageNumber="412" precision="15" seconds="48" value="25.213335">25º12′48″N</geoCoordinate>
<geoCoordinate id="C4E6506DA27CFFC8FE0E66C6FDFCAAE1" box="[439,539,985,1006]" degrees="91" direction="east" minutes="39" orientation="longitude" pageId="1" pageNumber="412" precision="15" seconds="48" value="91.66334">91º39′48″E</geoCoordinate>
,
<quantity id="662A9B4FA27CFFC8FD9F66C6FDB8AAE1" box="[550,607,986,1006]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="1" pageNumber="412" unit="m" value="735.0">
<elevation id="2AFFD199A27CFFC8FD9F66C6FDB8AAE1" box="[550,607,986,1006]" metricMagnitude="2" metricUnit="m" metricValue="7.35" pageId="1" pageNumber="412" unit="m" value="735.0">735 m</elevation>
</quantity>
/ Fikáček, Podskalská, Šípek lgt. // secondary tropical rainforest / with young trees + bamboo just / below the village, thin layer of / leaf litter (sifting)
</materialsCitation>
’.
<typeStatus id="7E698808A27CFFC8FC1666E9FBE0AD08" box="[943,1031,1013,1032]" pageId="1" pageNumber="412" type="paratype">PARATYPES</typeStatus>
: 3 JJ (
<collectionCode id="C7C3AE6FA27CFFC8FFCE610CFF50AD2C" box="[119,183,1040,1059]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="412">NMPC</collectionCode>
,
<collectionCode id="C7C3AE6FA27CFFC8FF79610CFF10AD2C" box="[192,247,1040,1059]" pageId="1" pageNumber="412">ZSPC</collectionCode>
): same data as the
<typeStatus id="7E698808A27CFFC8FE2F610CFE04AD2C" box="[406,483,1040,1059]" pageId="1" pageNumber="412" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27CFFCBFFC86127FC01AC5E" lastPageId="2" lastPageNumber="413" pageId="1" pageNumber="412" type="description">
<paragraph id="A16D36AAA27CFFC8FFC86127FDF5AD9C" blockId="1.[113,1089,1083,1587]" pageId="1" pageNumber="412">
<emphasis id="93A6EAB8A27CFFC8FFC86127FF1BAD5C" bold="true" box="[113,252,1083,1107]" pageId="1" pageNumber="412">Description.</emphasis>
Body length 2.0 mm. Length of body parts (
<typeStatus id="7E698808A27CFFC8FD766127FCD4AD5C" box="[719,819,1083,1107]" pageId="1" pageNumber="412" type="holotype">holotype</typeStatus>
): head
<quantity id="662A9B4FA27CFFC8FCC56127FC28AD5C" box="[892,975,1083,1107]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="412" unit="mm" value="0.3">0.3 mm</quantity>
, pronotum
<quantity id="662A9B4FA27CFFC8FFC86147FF23AD7C" box="[113,196,1115,1139]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="412" unit="mm" value="0.8">0.8 mm</quantity>
, elytra
<quantity id="662A9B4FA27CFFC8FEA86147FE83AD7C" box="[273,356,1115,1139]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="412" unit="mm" value="0.9">0.9 mm</quantity>
, antenna
<quantity id="662A9B4FA27CFFC8FE7F6147FDFDAD7C" box="[454,538,1115,1139]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="412" unit="mm" value="0.7">0.7 mm</quantity>
. Maximum width of body parts (
<typeStatus id="7E698808A27CFFC8FCCA6147FC30AD7C" box="[883,983,1115,1139]" pageId="1" pageNumber="412" type="holotype">holotype</typeStatus>
): head
<quantity id="662A9B4FA27CFFC8FBA66147FF7AAD9C" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="412" unit="mm" value="0.8">0.8 mm</quantity>
, pronotum
<quantity id="662A9B4FA27CFFC8FEAE6167FE8CAD9C" box="[279,363,1147,1171]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="412" unit="mm" value="1.2">1.2 mm</quantity>
, elytra
<quantity id="662A9B4FA27CFFC8FE006167FDE9AD9C" box="[441,526,1147,1171]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="412" unit="mm" value="1.2">1.2 mm</quantity>
.
</paragraph>
<paragraph id="A16D36AAA27CFFC8FF296187FE97AC1C" blockId="1.[113,1089,1083,1587]" pageId="1" pageNumber="412">
Body oval (
<figureCitation id="39E92A2FA27CFFC8FEAB6187FE84ADBC" box="[274,355,1179,1203]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="1" pageNumber="412">Fig. 18</figureCitation>
), dark chestnut coloured with lighter pronotal margins. Venter yellow-reddish. Legs and antennomeres I-VI yellow-red, antennomeres VII-XI dark. Dorsum without microreticulation; dorsal surface punctate, punctures regularly arranged, each with very short and fine seta.
</paragraph>
<paragraph id="A16D36AAA27CFFC8FF296007FBD0AC9C" blockId="1.[113,1089,1083,1587]" pageId="1" pageNumber="412">Head. Maximum width of head at basal third of eyes. Eyes bulging, 3.7 times as long as wide in dorsal view. Clypeus flatly emarginate, clypeal line absent. Anterolateral margin of head moderately narrow, raised. Dorsal surface with very sparsely scattered fine punctures separated by more than 8× of their diameter. Relative length of antennomeres III: II = 1.5.</paragraph>
<paragraph id="A16D36AAA27CFFC8FF296080FC7BACFC" blockId="1.[113,1089,1083,1587]" pageId="1" pageNumber="412">
Pronotum. Widest at basal third. Lateral sides almost straightly tapered from posterior to anterior corners in dorsal view; rounded in lateral view (
<figureCitation id="39E92A2FA27CFFC8FD7D60A7FCF5ACDC" box="[708,786,1467,1491]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="1" pageNumber="412">Fig. 19</figureCitation>
). Pronotal puncturation fine, similar to that of head; punctures separated by about 5–6× of their diameter.
</paragraph>
<paragraph id="A16D36AAA27CFFC8FF2960E7FD0DAF3C" blockId="1.[113,1089,1083,1587]" pageId="1" pageNumber="412">Elytra. Broadest approximately at basal fourth. Punctures similar to those on head and pronotum; puncturation a little sparser. Sutural stria absent.</paragraph>
<caption id="F5AD6622A27FFFCBFF216622FD20AA88" ID-DOI="http://doi.org/10.5281/zenodo.5331206" ID-Zenodo-Dep="5331206" httpUri="https://zenodo.org/record/5331206/files/figure.png" pageId="2" pageNumber="413" startId="2.[152,191,830,849]" targetBox="[180,1097,228,803]" targetPageId="2">
<paragraph id="A16D36AAA27FFFCBFF216622FD20AA88" blockId="2.[151,1126,830,903]" pageId="2" pageNumber="413">
Figs. 1–5. Genitalia of
<taxonomicName id="66D24D29A27FFFCBFEE16623FDE9AA5E" box="[344,526,830,849]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="undefined-1">
<emphasis id="93A6EAB8A27FFFCBFEE16623FE5BAA5E" box="[344,444,831,849]" italics="true" pageId="2" pageNumber="413">Agathidium</emphasis>
species. 1
</taxonomicName>
–2 – aedeagus of
<taxonomicName id="66D24D29A27FFFCBFD1A6623FC25AA5E" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[675,962,830,849]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov." subGenus="Agathidium">
<emphasis id="93A6EAB8A27FFFCBFD1A6623FCE0AA5E" box="[675,775,831,849]" italics="true" pageId="2" pageNumber="413">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A27FFFCBFCAB6623FC91AA5E" box="[786,886,831,849]" italics="true" pageId="2" pageNumber="413">Agathidium</emphasis>
)
<emphasis id="93A6EAB8A27FFFCBFC386623FC25AA5E" box="[897,962,831,849]" italics="true" pageId="2" pageNumber="413">fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A27FFFCBFC7E6622FBEEAA5E" box="[967,1033,830,849]" pageId="2" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
(1 – lateral view, 2 – apex in dorsal view). 3–5 –
<taxonomicName id="66D24D29A27FFFCBFE6D6646FD01AA63" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[468,742,857,876]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="cochleariforme" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A27FFFCBFE6D6646FE06AA63" box="[468,481,858,876]" italics="true" pageId="2" pageNumber="413">A</emphasis>
. (
<emphasis id="93A6EAB8A27FFFCBFE4A6646FDBFAA63" box="[499,600,858,876]" italics="true" pageId="2" pageNumber="413">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A27FFFCBFDDD6646FD01AA63" box="[612,742,858,876]" italics="true" pageId="2" pageNumber="413">cochleariforme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A27FFFCBFD526645FCC9AA63" box="[747,814,857,876]" pageId="2" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
(3 – aedeagus, lateral view; 4 – apex of aedeagus, dorsal view; 5 – spermatheca). Scale bars = 0.1 mm.
</paragraph>
</caption>
<paragraph id="A16D36AAA27FFFCBFF0C66E6FEE5AD5E" blockId="2.[151,1127,1017,1585]" pageId="2" pageNumber="413">
Legs. Protarsomere I slightly enlarged in male. Femora and tibiae slender, simple (i,e. without any tooth or emagination on posterior margin). Tarsal formula: 5-5-
<quantity id="662A9B4FA27FFFCBFC126105FC33AD3E" box="[939,980,1049,1073]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="413" unit="in" value="4.0">4 in</quantity>
male. Female unknown.
</paragraph>
<paragraph id="A16D36AAA27FFFCBFF0C6145FBE0AD7E" blockId="2.[151,1127,1017,1585]" box="[181,1031,1113,1137]" pageId="2" pageNumber="413">Mesoventrite. Longitudinal mesoventral carina low; lateral lines not developed.</paragraph>
<paragraph id="A16D36AAA27FFFCBFF0C6165FE3CAD9E" blockId="2.[151,1127,1017,1585]" box="[181,475,1145,1169]" pageId="2" pageNumber="413">Metathoracic wings absent.</paragraph>
<paragraph id="A16D36AAA27FFFCBFF0C6185FC8CADDE" blockId="2.[151,1127,1017,1585]" pageId="2" pageNumber="413">Metaventrite. Femoral lines incomplete. Male with shallow subcircular depression near posterior margin between coxae bearing a group of long erect setae.</paragraph>
<paragraph id="A16D36AAA27FFFCBFF0C61C5FC7CAC1E" blockId="2.[151,1127,1017,1585]" pageId="2" pageNumber="413">
Genitalia. Tegmen with horizontally orientated spiral part and broadly rounded apex as in
<figureCitation id="39E92A2FA27FFFCBFF2E61E5FF10AC1E" box="[151,247,1273,1297]" captionStart="Figs" captionStartId="2.[152,191,830,849]" captionTargetBox="[180,1097,228,803]" captionTargetId="figure-234@2.[175,1128,220,806]" captionTargetPageId="2" captionText="Figs. 1–5. Genitalia of Agathidium species. 1–2 – aedeagus of Agathidium (Agathidium) fikaceki sp. nov. (1 – lateral view, 2 – apex in dorsal view). 3–5 – A. (Macroceble) cochleariforme sp. nov. (3 – aedeagus, lateral view; 4 – apex of aedeagus, dorsal view; 5 – spermatheca). Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331206" httpUri="https://zenodo.org/record/5331206/files/figure.png" pageId="2" pageNumber="413">Figs 1–2</figureCitation>
. Operculum very long, narrowly U-shaped. Female not known.
</paragraph>
<paragraph id="A16D36AAA27FFFCBFF2E6005FC01AC5E" blockId="2.[151,1127,1017,1585]" pageId="2" pageNumber="413">
<emphasis id="93A6EAB8A27FFFCBFF2E6005FEF1AC3E" bold="true" box="[151,278,1305,1329]" pageId="2" pageNumber="413">Variability.</emphasis>
The length ratio of antennal segments III:II varies between
<quantity id="662A9B4FA27FFFCBFC246005FBEEAC3E" box="[925,1033,1305,1329]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" metricValueMax="3.81" metricValueMin="3.302" pageId="2" pageNumber="413" unit="in" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 in</quantity>
the
<typeStatus id="7E698808A27FFFCBFB806006FB80AC3D" box="[1081,1127,1306,1330]" pageId="2" pageNumber="413">type</typeStatus>
series; the ratio of length: width of eyes in dorsal view varies between 3.4–3.9.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27FFFCAFF2E6045FCDBABBC" lastPageId="3" lastPageNumber="414" pageId="2" pageNumber="413" type="diagnosis">
<paragraph id="A16D36AAA27FFFCAFF2E6045FCDBABBC" blockId="2.[151,1127,1017,1585]" lastBlockId="3.[113,1089,219,820]" lastPageId="3" lastPageNumber="414" pageId="2" pageNumber="413">
<emphasis id="93A6EAB8A27FFFCBFF2E6045FE6BAC7E" bold="true" box="[151,396,1369,1393]" pageId="2" pageNumber="413">Differential diagnosis.</emphasis>
<taxonomicName id="66D24D29A27FFFCBFE2A6046FD10AC7E" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[403,759,1369,1393]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov." subGenus="Agathidium">
<emphasis id="93A6EAB8A27FFFCBFE2A6046FDEAAC7E" box="[403,525,1370,1393]" italics="true" pageId="2" pageNumber="413">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A27FFFCBFDA46046FD70AC7E" box="[541,663,1370,1393]" italics="true" pageId="2" pageNumber="413">Agathidium</emphasis>
)
<emphasis id="93A6EAB8A27FFFCBFD1E6046FD10AC7E" box="[679,759,1370,1393]" italics="true" pageId="2" pageNumber="413">fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A27FFFCBFD476046FCB7AC7E" box="[766,848,1370,1393]" pageId="2" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="66D24D29A27FFFCBFC7F6046FDA4AC9E" authority="Angelini &amp; DeMarzo, 1986" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="pseudomontanellum" subGenus="Agathidium">
<emphasis id="93A6EAB8A27FFFCBFC7F6046FC39AC7E" box="[966,990,1370,1393]" italics="true" pageId="2" pageNumber="413">A.</emphasis>
(
<emphasis id="93A6EAB8A27FFFCBFC546046FBE3AC7E" box="[1005,1028,1370,1393]" italics="true" pageId="2" pageNumber="413">A.</emphasis>
)
<emphasis id="93A6EAB8A27FFFCBFBAD6046FEC7AC9E" italics="true" pageId="2" pageNumber="413">pseudomontanellum</emphasis>
Angelini &amp; DeMarzo,
<quantity id="662A9B4FA27FFFCBFDB76065FDA4AC9E" box="[526,579,1401,1425]" metricMagnitude="1" metricUnit="m" metricValue="5.04444" pageId="2" pageNumber="413" unit="in" value="1986.0">1986</quantity>
</taxonomicName>
in body size, coloration of the antennae, the presence of fine and sparse dorsal puncturation, absence of the microsculpture on the dorsum, absence of sutural striae, and by the broadly rounded apex of the tegmen in dorsal view. The new species differs from
<taxonomicName id="66D24D29A27FFFCBFEA360C6FDEDACFE" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" box="[282,522,1498,1521]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="pseudomontanellum">
<emphasis id="93A6EAB8A27FFFCBFEA360C6FDEDACFE" box="[282,522,1498,1521]" italics="true" pageId="2" pageNumber="413">A. pseudomontanellum</emphasis>
</taxonomicName>
by the lighter coloration of the dorsum, by the absence of metathoracic wings, by the longer antennomere III (the length ratio antennomeres III: II=
<quantity id="662A9B4FA27FFFCBFBA360E5FF4CAF3E" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" metricValueMax="3.81" metricValueMin="3.302" pageId="2" pageNumber="413" unit="in" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 in</quantity>
<taxonomicName id="66D24D29A27FFFCBFF0D6306FEC5AF3E" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[180,290,1562,1585]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov.">
<emphasis id="93A6EAB8A27FFFCBFF0D6306FEC5AF3E" box="[180,290,1562,1585]" italics="true" pageId="2" pageNumber="413">A. fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A27FFFCBFE936306FE9AAF3E" box="[298,381,1562,1585]" pageId="2" pageNumber="413" rank="species">sp. nov.</taxonomicNameLabel>
while 1.0 in
<taxonomicName id="66D24D29A27FFFCBFDB56306FD1BAF3E" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" box="[524,764,1562,1585]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="413" phylum="Arthropoda" rank="species" species="pseudomontanellum">
<emphasis id="93A6EAB8A27FFFCBFDB56306FD1BAF3E" box="[524,764,1562,1585]" italics="true" pageId="2" pageNumber="413">A. pseudomontanellum</emphasis>
</taxonomicName>
), and by the narrow parameres in both dorsal and lateral views (in contrast to dorsoventrally flattened parameres with broad basal part in lateral view in
<taxonomicName id="66D24D29A27EFFCAFE2E65E1FD60A81B" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" box="[407,647,253,276]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="pseudomontanellum">
<emphasis id="93A6EAB8A27EFFCAFE2E65E1FD60A81B" box="[407,647,253,276]" italics="true" pageId="3" pageNumber="414">A. pseudomontanellum</emphasis>
</taxonomicName>
). As it is sometimes difficult to recognize if the examined species belongs to the
<taxonomicName id="66D24D29A27EFFCAFDB26401FD19A83B" authorityName="Portevin" authorityYear="1937" box="[523,766,285,308]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="madurense">
<emphasis id="93A6EAB8A27EFFCAFDB26401FD19A83B" box="[523,766,285,308]" italics="true" pageId="3" pageNumber="414">Agathidium madurense</emphasis>
</taxonomicName>
species group or to a different species group, the new species was also compared with
<taxonomicName id="66D24D29A27EFFCAFD7A6421FC11A85B" authority="Portevin, 1937" authorityName="Portevin" authorityYear="1937" box="[707,1014,315,340]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="madurense">
<emphasis id="93A6EAB8A27EFFCAFD7A6421FCB4A85B" box="[707,851,317,340]" italics="true" pageId="3" pageNumber="414">A. madurense</emphasis>
Portevin, 1937
</taxonomicName>
and
<taxonomicName id="66D24D29A27EFFCAFB936421FDF6A87C" authority="Angelini &amp; DeMarzo, 1986" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="shilongense">
<emphasis id="93A6EAB8A27EFFCAFB936421FF0BA87B" italics="true" pageId="3" pageNumber="414">A. shilongense</emphasis>
Angelini &amp; DeMarzo, 1986
</taxonomicName>
(both attributed to the
<taxonomicName id="66D24D29A27EFFCAFD446441FC08A87B" authorityName="Portevin" authorityYear="1937" box="[765,1007,349,372]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="madurense">
<emphasis id="93A6EAB8A27EFFCAFD446441FC08A87B" box="[765,1007,349,372]" italics="true" pageId="3" pageNumber="414">Agathidium madurense</emphasis>
</taxonomicName>
species group by
<bibRefCitation id="C5434B5BA27EFFCAFF6C6460FE6EA89B" author="ANGELINI F." box="[213,393,380,404]" pageId="3" pageNumber="414" pagination="135 - 577" refId="ref7045" refString="ANGELINI F. 2004: Chiavi dicotomiche e catalogo degli Agathidium Panzer dell'Asia sudorientale e della Cina (Coleoptera, Leiodidae, Agathidiini). Annali del Museo Civico di Storia Naturale Giacomo Doria 95: 135 - 577." type="journal article" year="2004">ANGELINI (2004))</bibRefCitation>
. The new species is similar to both above species in the coloration of the dorsum and antennae, the presence of dorsal puncturation, absence of microsculpture on the dorsum, absence of sutural striae and clypeal line, the length ratio of the antennomeres III: II and by the same tarsal formula in males. The new species differs from
<taxonomicName id="66D24D29A27EFFCAFCC564C1FBF3A8FB" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" box="[892,1044,477,500]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="shilongense">
<emphasis id="93A6EAB8A27EFFCAFCC564C1FBF3A8FB" box="[892,1044,477,500]" italics="true" pageId="3" pageNumber="414">A. shilongense</emphasis>
</taxonomicName>
and
<taxonomicName id="66D24D29A27EFFCAFFC864E1FF18AB1B" authorityName="Portevin" authorityYear="1937" box="[113,255,509,532]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="madurense">
<emphasis id="93A6EAB8A27EFFCAFFC864E1FF18AB1B" box="[113,255,509,532]" italics="true" pageId="3" pageNumber="414">A. madurense</emphasis>
</taxonomicName>
by the absence of metathoracic wings, by smaller size (compared to 2.8–3.0 mm in
<taxonomicName id="66D24D29A27EFFCAFF356701FEC2AB3B" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" box="[140,293,541,564]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="shilongense">
<emphasis id="93A6EAB8A27EFFCAFF356701FEC2AB3B" box="[140,293,541,564]" italics="true" pageId="3" pageNumber="414">A. shilongense</emphasis>
</taxonomicName>
and
<quantity id="662A9B4FA27EFFCAFEE16700FE31AB3C" box="[344,470,540,564]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="3" pageNumber="414" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.2–3.3 mm</quantity>
in
<taxonomicName id="66D24D29A27EFFCAFE4E6701FD60AB3B" authorityName="Portevin" authorityYear="1937" box="[503,647,541,564]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="madurense">
<emphasis id="93A6EAB8A27EFFCAFE4E6701FD60AB3B" box="[503,647,541,564]" italics="true" pageId="3" pageNumber="414">A. madurense</emphasis>
</taxonomicName>
), by the spiral, horizontally oriented basal part of the tegmen (compared to the irregularly twisted basal part in
<taxonomicName id="66D24D29A27EFFCAFC896721FC20AB5B" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" box="[816,967,573,596]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="shilongense">
<emphasis id="93A6EAB8A27EFFCAFC896721FC20AB5B" box="[816,967,573,596]" italics="true" pageId="3" pageNumber="414">A. shilongense</emphasis>
</taxonomicName>
) and by the broadly rounded apex of the tegmen in dorsal view (compared to abruptly rounded apex in
<taxonomicName id="66D24D29A27EFFCAFB936741FF0BAB9B" authorityName="Angelini &amp; DeMarzo" authorityYear="1986" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="shilongense">
<emphasis id="93A6EAB8A27EFFCAFB936741FF0BAB9B" italics="true" pageId="3" pageNumber="414">A. shilongense</emphasis>
</taxonomicName>
). The new species also differs by the narrow parameres in lateral view, in contrast to parameres with a broad basal part in lateral view in
<taxonomicName id="66D24D29A27EFFCAFD1F6781FCD2ABBB" authorityName="Portevin" authorityYear="1937" box="[678,821,669,692]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="species" species="madurense">
<emphasis id="93A6EAB8A27EFFCAFD1F6781FCD2ABBB" box="[678,821,669,692]" italics="true" pageId="3" pageNumber="414">A. madurense</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A27EFFCAFFC867A0FCF8AA8C" pageId="3" pageNumber="414" type="etymology">
<paragraph id="A16D36AAA27EFFCAFFC867A0FBDEABDB" blockId="3.[113,1089,219,820]" box="[113,1081,699,724]" pageId="3" pageNumber="414">
<emphasis id="93A6EAB8A27EFFCAFFC867A0FF15ABDB" bold="true" box="[113,242,700,724]" pageId="3" pageNumber="414">Etymology.</emphasis>
The new species is named after one of its collectors, Martin Fikáček (
<collectingRegion id="6316F848A27EFFCAFC6467A0FBCAABDB" box="[989,1069,700,724]" country="Czech Republic" name="Praha" pageId="3" pageNumber="414">Prague</collectingRegion>
).
</paragraph>
<paragraph id="A16D36AAA27EFFCAFFC867C7FD4CAA3B" blockId="3.[113,1089,219,820]" pageId="3" pageNumber="414">
<emphasis id="93A6EAB8A27EFFCAFFC867C7FF08ABFC" bold="true" box="[113,239,731,755]" pageId="3" pageNumber="414">Bionomics.</emphasis>
The
<typeStatus id="7E698808A27EFFCAFE9F67C0FEB3ABFB" box="[294,340,732,756]" pageId="3" pageNumber="414">type</typeStatus>
specimens were collected by sifting of a thin layer of leaf litter in the bamboo bushes growing at the margin of the secondary tropical forest on the southern slope of the
<collectingRegion id="6316F848A27EFFCAFF0C6607FECCAA3C" box="[181,299,795,819]" country="India" name="Meghalaya" pageId="3" pageNumber="414">Meghalaya</collectingRegion>
Plateau (M. Fikáček, pers. comm.).
</paragraph>
<paragraph id="A16D36AAA27EFFCAFE2B6677FCF8AA8C" blockId="3.[402,799,875,899]" box="[402,799,875,899]" pageId="3" pageNumber="414">
subgenus
<taxonomicName id="66D24D29A27EFFCAFE456670FCF8AA8C" authority="Angelini, 1993" authorityName="Angelini" authorityYear="1993" box="[508,799,875,899]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="414" phylum="Arthropoda" rank="subGenus" subGenus="Macroceble">
<emphasis id="93A6EAB8A27EFFCAFE456670FD9DAA8C" box="[508,634,876,899]" italics="true" pageId="3" pageNumber="414">Macroceble</emphasis>
Angelini, 1993
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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