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<document id="DAD0E5CB7C7841C2F94F1B5F30D2869C" ID-DOI="10.5281/zenodo.5331202" ID-ISSN="0374-1036" ID-Zenodo-Dep="5331202" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630169194360" checkinUser="diego" docAuthor="Švec, Zdeněk" docDate="2012" docId="297B87BCA275FFC2FECF61A6FCA1AF42" docLanguage="en" docName="ActaEntMusNatPra.2012.52.2.411-424.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Agathidium (Macroceble) acutangulum Švec 2012, sp. nov." docType="treatment" docVersion="4" lastPageNumber="422" masterDocId="D542FFC4A27DFFC9FFB9651CFFE7A90F" masterDocTitle="New Leiodinae (Coleoptera: Leiodidae) from India and Papua New Guinea" masterLastPageNumber="424" masterPageNumber="411" pageNumber="419" updateTime="1699172137447" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="BEDE353B7422A21E5CC9F8F6F89ADD8C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="23A03960F8FADD2798DBC64CE77ACD18">
<mods:title id="2417798060BDB4602737952B22F80D3C">New Leiodinae (Coleoptera: Leiodidae) from India and Papua New Guinea</mods:title>
</mods:titleInfo>
<mods:name id="ADB3ADA2278D8354ACACC270BFD6DB60" type="personal">
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<mods:namePart id="80DEBD13A4ECD2CEA78DACD7D26F0B9C">Švec, Zdeněk</mods:namePart>
</mods:name>
<mods:typeOfResource id="1771DDAEC03F35B996F2F38850CBEED8">text</mods:typeOfResource>
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<mods:title id="C25323C873F8C90380572DAD73A1EB90">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="57AD7750328F85BFD4EAFB8083185615">
<mods:date id="7462DF2EF85B1E3D7D1A22D9BF0ECA8F">2012</mods:date>
<mods:detail id="595D8466A2C1F60856F450021C523425" type="pubDate">
<mods:number id="70CDED0F13CBF5E35ED2E0EFECF482AF">2012-12-17</mods:number>
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<mods:number id="DF86815EF29FBA941C22D4AAD6FAEDD6">52</mods:number>
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<mods:classification id="7D75A5BFC3E800CABB0900F9BE0DAF2C">journal article</mods:classification>
<mods:identifier id="C9CA395DB02D6C7A80DAB2E62499750C" type="DOI">10.5281/zenodo.5331202</mods:identifier>
<mods:identifier id="7BE3A349CAA42A019C6A41C72A7BA4AD" type="ISSN">0374-1036</mods:identifier>
<mods:identifier id="F4CA5433693BE90141CFF552247A6BF1" type="Zenodo-Dep">5331202</mods:identifier>
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<treatment id="297B87BCA275FFC2FECF61A6FCA1AF42" ID-DOI="http://doi.org/10.5281/zenodo.5398907" ID-Zenodo-Dep="5398907" LSID="urn:lsid:plazi:treatment:297B87BCA275FFC2FECF61A6FCA1AF42" httpUri="http://treatment.plazi.org/id/297B87BCA275FFC2FECF61A6FCA1AF42" lastPageId="11" lastPageNumber="422" pageId="8" pageNumber="419">
<subSubSection id="E9C86521A275FFC1FECF61A6FC6FADDB" box="[374,904,1210,1236]" pageId="8" pageNumber="419" type="nomenclature">
<paragraph id="A16D36AAA275FFC1FECF61A6FC6FADDB" blockId="8.[374,904,1210,1236]" box="[374,904,1210,1236]" pageId="8" pageNumber="419">
<heading id="FA2581C6A275FFC1FECF61A6FC6FADDB" bold="true" box="[374,904,1210,1236]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="419" reason="3">
<emphasis id="93A6EAB8A275FFC1FECF61A6FC6FADDB" bold="true" box="[374,904,1210,1236]" pageId="8" pageNumber="419">
<taxonomicName id="66D24D29A275FFC1FECF61A6FCCBADDC" ID-CoL="65PDW" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[374,812,1210,1235]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="419" phylum="Arthropoda" rank="species" species="acutangulum" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A275FFC1FECF61A6FE1FADDC" bold="true" box="[374,504,1210,1235]" italics="true" pageId="8" pageNumber="419">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A275FFC1FDB161A6FD6EADDC" bold="true" box="[520,649,1210,1235]" italics="true" pageId="8" pageNumber="419">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A275FFC1FD2061A6FCCBADDC" bold="true" box="[665,812,1210,1235]" italics="true" pageId="8" pageNumber="419">acutangulum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A275FFC1FC8B61A0FC6FADDB" box="[818,904,1212,1236]" pageId="8" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A275FFC1FD9C61C6FD3FADE2" box="[549,728,1242,1261]" pageId="8" pageNumber="419" type="description">
<paragraph id="A16D36AAA275FFC1FD9C61C6FD3FADE2" blockId="8.[549,728,1242,1261]" box="[549,728,1242,1261]" pageId="8" pageNumber="419">
(
<figureCitation id="39E92A2FA275FFC1FD9561C6FD74ADE2" box="[556,659,1242,1261]" captionStart="Figs" captionStartId="9.[113,152,885,904]" captionTargetBox="[138,1064,224,854]" captionTargetId="figure-227@9.[131,1071,219,865]" captionTargetPageId="9" captionText="Figs. 11–17. Genitalia of Agathidium and Zeadolopus. 11–13 – Agathidium (Macroceble) guttigerum sp nov. (11 – aedeagus, lateral view; 12 – apex of aedeagus, dorsal view; 13 – spermatheca). 14–16 – A. (Macroceble) acutangulum sp. nov.(14 – aedeagus, lateral view; 15 – apex of aedeagus, dorsal view; 16 – spermatheca). 17 – Zeadolopus punctiventris sp. nov., aedeagus, dorsal view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331212" httpUri="https://zenodo.org/record/5331212/files/figure.png" pageId="8" pageNumber="419">Figs. 14–16</figureCitation>
,
<figureCitation id="39E92A2FA275FFC1FD2561C6FD35ADE2" box="[668,722,1242,1261]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="8" pageNumber="419">28, 29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A275FFC1FF2E6018FEBDACE3" pageId="8" pageNumber="419" type="materials_examined">
<paragraph id="A16D36AAA275FFC1FF2E6018FEE2AC3E" blockId="8.[151,1127,1283,1516]" pageId="8" pageNumber="419">
<emphasis id="93A6EAB8A275FFC1FF2E6018FEEAAC18" bold="true" box="[151,269,1283,1303]" pageId="8" pageNumber="419">
<typeStatus id="7E698808A275FFC1FF2E6018FF24AC18" box="[151,195,1284,1303]" pageId="8" pageNumber="419">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="D9C5763AA275FFC1FEA96018FEDAAC18" box="[272,317,1284,1303]" name="India" pageId="8" pageNumber="419">India</collectingCountry>
,
<collectingRegion id="6316F848A275FFC1FEFA6018FE04AC18" box="[323,483,1284,1303]" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="419">Arunachal Pradesh</collectingRegion>
,
<quantity id="662A9B4FA275FFC1FE506018FDFBAC18" box="[489,540,1284,1303]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="419" unit="km" value="11.0">11 km</quantity>
SSE of Tenga, Eagles Nest wildlife sanctuary,
<geoCoordinate id="C4E6506DA275FFC1FC256018FC1AAC18" box="[924,1021,1283,1303]" degrees="27" direction="north" minutes="06" orientation="latitude" pageId="8" pageNumber="419" precision="15" seconds="23" value="27.10639">27º06′23″N</geoCoordinate>
<geoCoordinate id="C4E6506DA275FFC1FBBB6018FB84AC18" box="[1026,1123,1283,1303]" degrees="92" direction="east" minutes="26" orientation="longitude" pageId="8" pageNumber="419" precision="15" seconds="02" value="92.43389">92º26′02″E</geoCoordinate>
,
<quantity id="662A9B4FA275FFC1FF2E6002FF3FAC3D" box="[151,216,1310,1330]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="8" pageNumber="419" unit="m" value="2510.0">2510 m</quantity>
a.s.l.
</paragraph>
<paragraph id="A16D36AAA275FFC1FF2E6025FEBDACE3" blockId="8.[151,1127,1283,1516]" pageId="8" pageNumber="419">
<emphasis id="93A6EAB8A275FFC1FF2E6025FEF9AC43" bold="true" box="[151,286,1336,1356]" pageId="8" pageNumber="419">Type material.</emphasis>
<typeStatus id="7E698808A275FFC1FE9F6025FE9EAC44" box="[294,377,1337,1356]" pageId="8" pageNumber="419" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="C7C3AE6FA275FFC1FE1D6025FE01AC43" box="[420,486,1337,1356]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="419">NMPC</collectionCode>
): ‘
<collectingCountry id="D9C5763AA275FFC1FE436025FDDAAC43" box="[506,573,1337,1356]" name="India" pageId="8" pageNumber="419">INDIA</collectingCountry>
:
<collectingRegion id="6316F848A275FFC1FDFF6025FD0CAC43" box="[582,747,1337,1356]" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="419">Arunachal Pradesh</collectingRegion>
(15) /
<quantity id="662A9B4FA275FFC1FC926025FC84AC43" box="[811,867,1337,1356]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="419" unit="km" value="11.0">11 km</quantity>
SSE of Tenga, Eagles / Nest WL Sanctuary,
<date id="D56C106AA275FFC1FEA46048FE81AC68" box="[285,358,1364,1383]" pageId="8" pageNumber="419" value="2008-05-04">4.v.2008</date>
/
<geoCoordinate id="C4E6506DA275FFC1FEC06048FE3BAC68" box="[377,476,1363,1383]" degrees="27" direction="north" minutes="06" orientation="latitude" pageId="8" pageNumber="419" precision="15" seconds="23" value="27.10639">27º06′23″N</geoCoordinate>
<geoCoordinate id="C4E6506DA275FFC1FE5B6048FDA2AC68" box="[482,581,1363,1383]" degrees="92" direction="east" minutes="26" orientation="longitude" pageId="8" pageNumber="419" precision="15" seconds="02" value="92.43389">92º26′02″E</geoCoordinate>
,
<quantity id="662A9B4FA275FFC1FDF6604FFD76AC68" box="[591,657,1363,1383]" metricMagnitude="3" metricUnit="m" metricValue="2.51" pageId="8" pageNumber="419" unit="m" value="2510.0">2510 m</quantity>
/ Fikáček, Podskalská, Šípek lgt. // Primary evergreen mountain / cloud forest with very sparse / understory vegetation / sifting: wet leaf litter’.
<typeStatus id="7E698808A275FFC1FC296073FC0FAC8F" box="[912,1000,1391,1410]" pageId="8" pageNumber="419" type="paratype">PARATYPES</typeStatus>
: 5 JJ,
<specimenCount id="B7D4FD23A275FFC1FB956072FB81AC8E" box="[1068,1126,1384,1409]" pageId="8" pageNumber="419" type="female">12 ♀♀</specimenCount>
(
<collectionCode id="C7C3AE6FA275FFC1FF246095FF3AAC93" box="[157,221,1417,1436]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="419">NMPC</collectionCode>
,
<collectionCode id="C7C3AE6FA275FFC1FF516095FEC7AC93" box="[232,288,1417,1436]" pageId="8" pageNumber="419">ZSPC</collectionCode>
): same data as the
<typeStatus id="7E698808A275FFC1FE746095FDFCAC93" box="[461,539,1417,1436]" pageId="8" pageNumber="419" type="holotype">holotype</typeStatus>
;
<specimenCount id="B7D4FD23A275FFC1FD9F6095FDAFAC93" box="[550,584,1411,1436]" pageId="8" pageNumber="419" type="female">1 ♀</specimenCount>
(
<collectionCode id="C7C3AE6FA275FFC1FDEF6095FD7FAC93" box="[598,664,1417,1436]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="8" pageNumber="419">NMPC</collectionCode>
): ‘
<collectingCountry id="D9C5763AA275FFC1FD156095FD17AC93" box="[684,752,1417,1436]" name="India" pageId="8" pageNumber="419">INDIA</collectingCountry>
:
<collectingRegion id="6316F848A275FFC1FD406095FC78AC93" box="[761,927,1417,1436]" country="India" name="Arunachal Pradesh" pageId="8" pageNumber="419">Arunachal Pradesh</collectingRegion>
(12) /
<quantity id="662A9B4FA275FFC1FC596095FBF9AC93" box="[992,1054,1417,1436]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="8" pageNumber="419" unit="km" value="1.6">1.6 km</quantity>
SEE of Tawang (at petrol station) /
<date id="D56C106AA275FFC1FE3F60B8FE05ACB8" box="[390,482,1444,1463]" pageId="8" pageNumber="419" value="2008-04-30">30.iv.2008</date>
,
<geoCoordinate id="C4E6506DA275FFC1FE5260BFFDA9ACB8" box="[491,590,1443,1463]" degrees="27" direction="north" minutes="35" orientation="latitude" pageId="8" pageNumber="419" precision="15" seconds="55" value="27.598612">27º35′55″N</geoCoordinate>
<geoCoordinate id="C4E6506DA275FFC1FDEC60BFFD5FACB8" box="[597,696,1443,1463]" degrees="91" direction="east" minutes="52" orientation="longitude" pageId="8" pageNumber="419" precision="15" seconds="47" value="91.87972">91º52′47″E</geoCoordinate>
,
<quantity id="662A9B4FA275FFC1FD7860B8FCE4ACB8" box="[705,771,1444,1463]" metricMagnitude="3" metricUnit="m" metricValue="2.88" pageId="8" pageNumber="419" unit="m" value="2880.0">2880 m</quantity>
/ Fikáček, Podskalská, Šípek lgt. // small remnants of high
<emphasis id="93A6EAB8A275FFC1FE9660A3FDE1ACDE" box="[303,518,1471,1489]" italics="true" pageId="8" pageNumber="419">
<taxonomicName id="66D24D29A275FFC1FE9660A3FE67ACDE" authorityName="C.Linnaeus" authorityYear="1753" box="[303,384,1471,1489]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="419" phylum="Tracheophyta" rank="genus">Juniperus</taxonomicName>
/
<taxonomicName id="66D24D29A275FFC1FE3060A3FDE1ACDE" box="[393,518,1471,1489]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="419" phylum="Tracheophyta" rank="genus">Rhododendron</taxonomicName>
</emphasis>
forest with / sparse understory vegetation / sifting: leaf litter + fallen /
<taxonomicName id="66D24D29A275FFC1FF2E60C6FEF2ACE3" box="[151,277,1498,1516]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="8" pageNumber="419" phylum="Tracheophyta" rank="genus">
<emphasis id="93A6EAB8A275FFC1FF2E60C6FEF2ACE3" box="[151,277,1498,1516]" italics="true" pageId="8" pageNumber="419">Rhododendron</emphasis>
</taxonomicName>
petals’.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A275FFC2FF2E6318FBF4AD02" lastPageId="11" lastPageNumber="422" pageId="8" pageNumber="419" type="description">
<paragraph id="A16D36AAA275FFC1FF2E6318FCB7AF53" blockId="8.[151,1127,1540,1628]" pageId="8" pageNumber="419">
<emphasis id="93A6EAB8A275FFC1FF2E6318FEC5AF13" bold="true" box="[151,290,1540,1564]" pageId="8" pageNumber="419">Description.</emphasis>
Body length
<quantity id="662A9B4FA275FFC1FE096318FDD0AF13" box="[432,567,1540,1564]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="8" pageNumber="419" unit="mm" value="2.45" valueMax="2.7" valueMin="2.2">2.2– 2.7 mm</quantity>
(
<typeStatus id="7E698808A275FFC1FD876318FD78AF13" box="[574,671,1540,1564]" pageId="8" pageNumber="419" type="holotype">holotype</typeStatus>
:
<quantity id="662A9B4FA275FFC1FD116318FD1AAF13" box="[680,765,1540,1564]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="8" pageNumber="419" unit="mm" value="2.3">2.3 mm</quantity>
). Length of body parts (
<typeStatus id="7E698808A275FFC1FC416318FBBBAF13" box="[1016,1116,1540,1564]" pageId="8" pageNumber="419" type="holotype">holotype</typeStatus>
): head
<quantity id="662A9B4FA275FFC1FF696338FEC2AF33" box="[208,293,1572,1596]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="419" unit="mm" value="0.5">0.5 mm</quantity>
, pronotum
<quantity id="662A9B4FA275FFC1FE266338FE14AF33" box="[415,499,1572,1596]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="419" unit="mm" value="0.9">0.9 mm</quantity>
, elytra
<quantity id="662A9B4FA275FFC1FDFB6338FD71AF33" box="[578,662,1572,1596]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="419" unit="mm" value="0.9">0.9 mm</quantity>
, antenna
<quantity id="662A9B4FA275FFC1FD426338FCB7AF33" box="[763,848,1572,1596]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="8" pageNumber="419" unit="mm" value="0.7">0.7 mm</quantity>
. Maximum width of body parts (
<typeStatus id="7E698808A275FFC1FF606358FEDAAF53" box="[217,317,1604,1628]" pageId="8" pageNumber="419" type="holotype">holotype</typeStatus>
): head
<quantity id="662A9B4FA275FFC1FE316358FE3BAF53" box="[392,476,1604,1628]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="8" pageNumber="419" unit="mm" value="1.2">1.2 mm</quantity>
, pronotum
<quantity id="662A9B4FA275FFC1FDEC6358FD4EAF53" box="[597,681,1604,1628]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="419" unit="mm" value="1.5">1.5 mm</quantity>
, elytra
<quantity id="662A9B4FA275FFC1FD4E6358FCABAF53" box="[759,844,1604,1628]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="8" pageNumber="419" unit="mm" value="1.4">1.4 mm</quantity>
.
</paragraph>
<caption id="F5AD6622A274FFC0FFC86669FD63AAD7" ID-DOI="http://doi.org/10.5281/zenodo.5331212" ID-Zenodo-Dep="5331212" httpUri="https://zenodo.org/record/5331212/files/figure.png" pageId="9" pageNumber="420" startId="9.[113,152,885,904]" targetBox="[138,1064,224,854]" targetPageId="9">
<paragraph id="A16D36AAA274FFC0FFC86669FD63AAD7" blockId="9.[113,1089,885,984]" pageId="9" pageNumber="420">
Figs. 11–17. Genitalia of
<taxonomicName id="66D24D29A274FFC0FEF6666AFE54AA87" authorityName="Panzer" authorityYear="1797" box="[335,435,886,904]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
<emphasis id="93A6EAB8A274FFC0FEF6666AFE54AA87" box="[335,435,886,904]" italics="true" pageId="9" pageNumber="420">Agathidium</emphasis>
</taxonomicName>
and
<taxonomicName id="66D24D29A274FFC0FE67666AFDA5AA87" authorityName="Broun" authorityYear="1903" box="[478,578,886,904]" class="Insecta" family="Leiodidae" genus="Zeadolopus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="genus">
<emphasis id="93A6EAB8A274FFC0FE67666AFDA5AA87" box="[478,578,886,904]" italics="true" pageId="9" pageNumber="420">Zeadolopus</emphasis>
</taxonomicName>
. 11–13 –
<taxonomicName id="66D24D29A274FFC0FD20666AFC3CAA87" box="[665,987,885,904]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="guttigerum" subGenus="Macroceble">
<emphasis id="93A6EAB8A274FFC0FD20666AFD1AAA87" box="[665,765,886,904]" italics="true" pageId="9" pageNumber="420">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A274FFC0FCB2666AFC97AA87" box="[779,880,886,904]" italics="true" pageId="9" pageNumber="420">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A274FFC0FCC4666AFC3CAA87" box="[893,987,886,904]" italics="true" pageId="9" pageNumber="420">guttigerum</emphasis>
</taxonomicName>
sp nov. (11 – aedeagus, lateral view; 12 – apex of aedeagus, dorsal view; 13 – spermatheca). 14–16 –
<taxonomicName id="66D24D29A274FFC0FCD6668CFF41AAB2" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="acutangulum" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A274FFC0FCD6668CFC64AAAD" box="[879,899,912,930]" italics="true" pageId="9" pageNumber="420">A.</emphasis>
(
<emphasis id="93A6EAB8A274FFC0FC37668CFC14AAAD" box="[910,1011,912,930]" italics="true" pageId="9" pageNumber="420">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A274FFC0FC46668CFF41AAB2" italics="true" pageId="9" pageNumber="420">acutangulum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A274FFC0FF1266B6FF0AAAB2" box="[171,237,938,957]" pageId="9" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
(14 – aedeagus, lateral view; 15 – apex of aedeagus, dorsal view; 16 – spermatheca). 17 –
<taxonomicName id="66D24D29A274FFC0FC6466B7FF07AAD7" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" class="Insecta" family="Leiodidae" genus="Zeadolopus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="420" phylum="Arthropoda" rank="species" species="punctiventris" status="sp. nov.">
<emphasis id="93A6EAB8A274FFC0FC6466B7FF07AAD7" italics="true" pageId="9" pageNumber="420">Zeadolopus punctiventris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A274FFC0FF5F66D9FECCAAD7" box="[230,299,965,984]" pageId="9" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
, aedeagus, dorsal view. Scale = 0.1 mm.
</paragraph>
</caption>
<paragraph id="A16D36AAA274FFC0FF29610BFD14AD61" blockId="9.[113,1089,1047,1615]" pageId="9" pageNumber="420">
Body shortly oval (
<figureCitation id="39E92A2FA274FFC0FEDD610BFE53AD20" box="[356,436,1047,1071]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="9" pageNumber="420">Fig. 28</figureCitation>
), black-brown with lighter margins of pronotum and elytra, antennae and legs yellow-reddish. Venter light yellow-reddish. Dorsum without microreticulation, punctate, each puncture with very short and fine seta.
</paragraph>
<paragraph id="A16D36AAA274FFC0FF29616BFC66ADE0" blockId="9.[113,1089,1047,1615]" pageId="9" pageNumber="420">Head. Maximum width of head far before posterior margin of eyes. Eyes flat, 6× as long as wide in dorsal view. Clypeus moderately emarginate, clypeal line lacking. Dorsal surface with rare and very fine punctures separated by much more than 10× their diameter. Mandibles without striking characters. Relative length of antennomeres III: II = 1.4.</paragraph>
<paragraph id="A16D36AAA274FFC0FF2961EBFC25AC41" blockId="9.[113,1089,1047,1615]" pageId="9" pageNumber="420">
Pronotum. Widest at basal third. Lateral sides flatly rounded toward anterior angles in dorsal view; with distinct very closely rounded angle in lateral view (
<figureCitation id="39E92A2FA274FFC0FCEC600BFC42AC20" box="[853,933,1303,1327]" captionStart="Figs 18–29" captionStartId="10.[151,188,1516,1535]" captionTargetBox="[154,1124,236,1485]" captionTargetId="figure-11@10.[152,1127,219,1493]" captionTargetPageId="10" captionText="Figs 18–29. Habitus of the holotypes of Agathidium species. 18–19 – A. (A.) fikaceki sp. nov.; 20–21 – A. (Macroceble) cochleariforme sp. nov.; 22–23 – A. (Macr.) armigerum sp. nov.; 24–25 – A. (Macr.) unicorne sp. nov., 26–27 – A. (Macr.) guttiferum sp. nov.; 28–29 – A. (Macr.) acutangulum sp. nov. (18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view)." figureDoi="http://doi.org/10.5281/zenodo.5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="9" pageNumber="420">Fig. 29</figureCitation>
). Puncturation sparser and finer than that of head, here and there with several larger punctures.
</paragraph>
<paragraph id="A16D36AAA274FFC0FF29604BFDE9AC80" blockId="9.[113,1089,1047,1615]" pageId="9" pageNumber="420">Elytra. Broadest approximately at anterior quarter of elytral length. Punctures very small, fine and very rare. Sutural stria absent.</paragraph>
<paragraph id="A16D36AAA274FFC0FF29608BFEFAAF00" blockId="9.[113,1089,1047,1615]" pageId="9" pageNumber="420">
Legs. Anterior tarsomeres I and II a little enlarged in male. Tarsi slender in female. Femora of both sexes with flat emagination at posterior margin situated before and behind midlength, hence forming hardly detectable unobtrusive central angle. Tarsal formula 5-5-
<quantity id="662A9B4FA274FFC0FC6960CAFC1AACE0" box="[976,1021,1494,1519]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="420" unit="in" value="4.0">4 in</quantity>
male, 4-4-
<quantity id="662A9B4FA274FFC0FF2460EBFF20AF00" box="[157,199,1527,1551]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="420" unit="in" value="4.0">4 in</quantity>
female.
</paragraph>
<paragraph id="A16D36AAA274FFC0FF29630BFBD8AF20" blockId="9.[113,1089,1047,1615]" box="[144,1087,1559,1583]" pageId="9" pageNumber="420">Mesoventrite. Longitudinal mesoventral carina low and feeble; lateral lines not developed.</paragraph>
<paragraph id="A16D36AAA274FFC0FF29632BFE52AF40" blockId="9.[113,1089,1047,1615]" box="[144,437,1591,1615]" pageId="9" pageNumber="420">Metathoracic wings absent.</paragraph>
<caption id="F5AD6622A277FFC3FF2E60F0FE53AF40" ID-DOI="http://doi.org/10.5281/zenodo.5331214" ID-Zenodo-Dep="5331214" httpUri="https://zenodo.org/record/5331214/files/figure.png" pageId="10" pageNumber="421" startId="10.[151,188,1516,1535]" targetBox="[154,1124,236,1485]" targetPageId="10">
<paragraph id="A16D36AAA277FFC3FF2E60F0FE53AF40" blockId="10.[151,1127,1516,1615]" pageId="10" pageNumber="421">
Figs 18–29. Habitus of the holotypes of
<taxonomicName id="66D24D29A277FFC3FE4960F1FD7AACF0" box="[496,669,1516,1535]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="421" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="93A6EAB8A277FFC3FE4960F1FDB3ACF0" box="[496,596,1517,1535]" italics="true" pageId="10" pageNumber="421">Agathidium</emphasis>
species.
</taxonomicName>
18–19 –
<taxonomicName id="66D24D29A277FFC3FD5560F1FC8CACF0" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[748,875,1516,1535]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="421" phylum="Arthropoda" rank="species" species="fikaceki" status="sp. nov." subGenus="Agathidium">
<emphasis id="93A6EAB8A277FFC3FD5560F1FD18ACF0" box="[748,767,1517,1535]" italics="true" pageId="10" pageNumber="421">A.</emphasis>
(
<emphasis id="93A6EAB8A277FFC3FCB260F1FCF9ACF0" box="[779,798,1517,1535]" italics="true" pageId="10" pageNumber="421">A.</emphasis>
)
<emphasis id="93A6EAB8A277FFC3FC9360F1FC8CACF0" box="[810,875,1517,1535]" italics="true" pageId="10" pageNumber="421">fikaceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A277FFC3FCC960F0FC51ACF0" box="[880,950,1516,1535]" pageId="10" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
; 20–21 –
<taxonomicName id="66D24D29A277FFC3FBB160F1FEB5AF15" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="421" phylum="Arthropoda" rank="species" species="cochleariforme" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A277FFC3FBB160F1FBFBACF0" box="[1032,1052,1517,1535]" italics="true" pageId="10" pageNumber="421">A.</emphasis>
(
<emphasis id="93A6EAB8A277FFC3FB9E60F1FF23AF15" italics="true" pageId="10" pageNumber="421">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A277FFC3FF696314FEB5AF15" box="[208,338,1544,1562]" italics="true" pageId="10" pageNumber="421">cochleariforme</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A277FFC3FEEF631BFE7DAF15" box="[342,410,1543,1562]" pageId="10" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
; 22–23 –
<emphasis id="93A6EAB8A277FFC3FE506314FE1AAF15" box="[489,509,1544,1562]" italics="true" pageId="10" pageNumber="421">A.</emphasis>
(
<emphasis id="93A6EAB8A277FFC3FDBE6314FDDDAF15" box="[519,570,1544,1562]" italics="true" pageId="10" pageNumber="421">Macr.</emphasis>
)
<taxonomicName id="66D24D29A277FFC3FDFD6314FD44AF15" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[580,675,1544,1562]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="421" phylum="Arthropoda" rank="species" species="armigerum" status="sp. nov.">
<emphasis id="93A6EAB8A277FFC3FDFD6314FD44AF15" box="[580,675,1544,1562]" italics="true" pageId="10" pageNumber="421">armigerum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A277FFC3FD1E631BFD0CAF15" box="[679,747,1543,1562]" pageId="10" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
; 24–25 –
<emphasis id="93A6EAB8A277FFC3FC836314FCA0AF15" box="[826,839,1544,1562]" italics="true" pageId="10" pageNumber="421">A</emphasis>
. (
<emphasis id="93A6EAB8A277FFC3FCE16314FC61AF15" box="[856,902,1544,1562]" italics="true" pageId="10" pageNumber="421">Macr</emphasis>
.)
<taxonomicName id="66D24D29A277FFC3FC2E6314FC04AF15" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[919,995,1544,1562]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="421" phylum="Arthropoda" rank="species" species="unicorne" status="sp. nov.">
<emphasis id="93A6EAB8A277FFC3FC2E6314FC04AF15" box="[919,995,1544,1562]" italics="true" pageId="10" pageNumber="421">unicorne</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A277FFC3FC5E631BFBCCAF15" box="[999,1067,1543,1562]" pageId="10" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
, 26–27 –
<emphasis id="93A6EAB8A277FFC3FF1E633EFF5DAF3B" box="[167,186,1570,1588]" italics="true" pageId="10" pageNumber="421">A.</emphasis>
(
<emphasis id="93A6EAB8A277FFC3FF7C633EFF1FAF3B" box="[197,248,1570,1588]" italics="true" pageId="10" pageNumber="421">Macr.</emphasis>
)
<taxonomicName id="66D24D29A277FFC3FEBA633EFEBBAF3B" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[259,348,1570,1588]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="421" phylum="Arthropoda" rank="species" species="guttiferum" status="sp. nov.">
<emphasis id="93A6EAB8A277FFC3FEBA633EFEBBAF3B" box="[259,348,1570,1588]" italics="true" pageId="10" pageNumber="421">guttiferum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A277FFC3FED8633EFE41AF3A" box="[353,422,1570,1589]" pageId="10" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
; 28–29 –
<emphasis id="93A6EAB8A277FFC3FE4E633EFDECAF3B" box="[503,523,1570,1588]" italics="true" pageId="10" pageNumber="421">A.</emphasis>
(
<emphasis id="93A6EAB8A277FFC3FDAF633EFDAFAF3B" box="[534,584,1570,1588]" italics="true" pageId="10" pageNumber="421">Macr.</emphasis>
)
<taxonomicName id="66D24D29A277FFC3FDEA633EFD25AF3B" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[595,706,1570,1588]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="421" phylum="Arthropoda" rank="species" species="acutangulum" status="sp. nov.">
<emphasis id="93A6EAB8A277FFC3FDEA633EFD25AF3B" box="[595,706,1570,1588]" italics="true" pageId="10" pageNumber="421">acutangulum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A277FFC3FD71633EFCEDAF3A" box="[712,778,1570,1589]" pageId="10" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
(18, 20, 22, 24, 26, 28 – dorsal view; 19, 21, 23, 25, 27, 29 – lateral view).
</paragraph>
</caption>
<caption id="F5AD6622A276FFC2FFC867AAFEEAABEC" ID-DOI="http://doi.org/10.5281/zenodo.5331216" ID-Zenodo-Dep="5331216" httpUri="https://zenodo.org/record/5331216/files/figure.png" pageId="11" pageNumber="422" startId="11.[113,152,694,713]" targetBox="[113,1068,221,681]" targetPageId="11">
<paragraph id="A16D36AAA276FFC2FFC867AAFEEAABEC" blockId="11.[113,1089,694,740]" pageId="11" pageNumber="422">
Figs. 30–32. Habitus of the holotype of
<taxonomicName id="66D24D29A276FFC2FE6A67ABFD4AABC6" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[467,685,695,713]" class="Insecta" family="Leiodidae" genus="Zeadolopus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="punctiventris" status="sp. nov.">
<emphasis id="93A6EAB8A276FFC2FE6A67ABFD4AABC6" box="[467,685,695,713]" italics="true" pageId="11" pageNumber="422">Zeadolopus punctiventris</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A276FFC2FD0D67AAFD1EABC6" box="[692,761,694,713]" pageId="11" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
(30 – dorsal view; 31 – lateral view; 32 – ventral view)
</paragraph>
</caption>
<paragraph id="A16D36AAA276FFC2FF296609FDD9AA43" blockId="11.[113,1089,789,1613]" pageId="11" pageNumber="422">Metaventrite. Femoral lines absent. Both sexes with club-shaped ventro-caudally oriented process at posterior margin between coxae.</paragraph>
<paragraph id="A16D36AAA276FFC2FF296649FDD3AA82" blockId="11.[113,1089,789,1613]" pageId="11" pageNumber="422">
Genitalia. Aedeagus as in
<figureCitation id="39E92A2FA276FFC2FE0A6649FDD0AA62" box="[435,567,853,877]" captionStart="Figs" captionStartId="9.[113,152,885,904]" captionTargetBox="[138,1064,224,854]" captionTargetId="figure-227@9.[131,1071,219,865]" captionTargetPageId="9" captionText="Figs. 11–17. Genitalia of Agathidium and Zeadolopus. 11–13 – Agathidium (Macroceble) guttigerum sp nov. (11 – aedeagus, lateral view; 12 – apex of aedeagus, dorsal view; 13 – spermatheca). 14–16 – A. (Macroceble) acutangulum sp. nov.(14 – aedeagus, lateral view; 15 – apex of aedeagus, dorsal view; 16 – spermatheca). 17 – Zeadolopus punctiventris sp. nov., aedeagus, dorsal view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331212" httpUri="https://zenodo.org/record/5331212/files/figure.png" pageId="11" pageNumber="422">Figs. 14–15</figureCitation>
; operculum oval with short notch in middle of anterior margin; spermatheca as in
<figureCitation id="39E92A2FA276FFC2FE516669FDD7AA82" box="[488,560,885,909]" captionStart="Figs" captionStartId="9.[113,152,885,904]" captionTargetBox="[138,1064,224,854]" captionTargetId="figure-227@9.[131,1071,219,865]" captionTargetPageId="9" captionText="Figs. 11–17. Genitalia of Agathidium and Zeadolopus. 11–13 – Agathidium (Macroceble) guttigerum sp nov. (11 – aedeagus, lateral view; 12 – apex of aedeagus, dorsal view; 13 – spermatheca). 14–16 – A. (Macroceble) acutangulum sp. nov.(14 – aedeagus, lateral view; 15 – apex of aedeagus, dorsal view; 16 – spermatheca). 17 – Zeadolopus punctiventris sp. nov., aedeagus, dorsal view. Scale = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5331212" httpUri="https://zenodo.org/record/5331212/files/figure.png" pageId="11" pageNumber="422">Fig.16</figureCitation>
.
</paragraph>
<paragraph id="A16D36AAA276FFC2FFC86689FBF4AD02" blockId="11.[113,1089,789,1613]" pageId="11" pageNumber="422">
<emphasis id="93A6EAB8A276FFC2FFC86689FF0AAAA2" bold="true" box="[113,237,917,941]" pageId="11" pageNumber="422">Variability.</emphasis>
The length ratio of antennomeres III: II varies between
<quantity id="662A9B4FA276FFC2FC836689FC45AAA2" box="[826,930,917,941]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="4.064" metricValueMin="3.5559999999999996" pageId="11" pageNumber="422" unit="in" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 in</quantity>
the
<typeStatus id="7E698808A276FFC2FC766689FC1BAAA2" box="[975,1020,917,941]" pageId="11" pageNumber="422">type</typeStatus>
specimens; the ratio of length: width of the eyes in dorsal view between 4–6. The coloration of the dorsum varies from uniformly dark brown (with lighter margins of the pronotum and elytra) to a bicoloured dorsum with a light chest-nut pronotum and/or head with dark brown elytra.
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A276FFC2FFC86109FDFCAC82" pageId="11" pageNumber="422" type="diagnosis">
<paragraph id="A16D36AAA276FFC2FFC86109FDFCAC82" blockId="11.[113,1089,789,1613]" pageId="11" pageNumber="422">
<emphasis id="93A6EAB8A276FFC2FFC86109FE8DAD22" bold="true" box="[113,362,1045,1069]" pageId="11" pageNumber="422">Differential diagnosis.</emphasis>
The shape of the aedeagus in
<taxonomicName id="66D24D29A276FFC2FD26610AFBA7AD22" authority="Švec, 2012" authorityName="Švec" authorityYear="2012" box="[671,1088,1045,1069]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="acutangulum" status="sp. nov." subGenus="Macroceble">
<emphasis id="93A6EAB8A276FFC2FD26610AFCFCAD22" box="[671,795,1046,1069]" italics="true" pageId="11" pageNumber="422">Agathidium</emphasis>
(
<emphasis id="93A6EAB8A276FFC2FC90610AFC40AD22" box="[809,935,1046,1069]" italics="true" pageId="11" pageNumber="422">Macroceble</emphasis>
)
<emphasis id="93A6EAB8A276FFC2FC0C610AFBA7AD22" box="[949,1088,1046,1069]" italics="true" pageId="11" pageNumber="422">acutangulum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="889557C3A276FFC2FFC86129FF22AD43" box="[113,197,1077,1100]" pageId="11" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
is almost identical to that in
<emphasis id="93A6EAB8A276FFC2FDB9612AFDFFAD42" box="[512,536,1078,1101]" italics="true" pageId="11" pageNumber="422">A.</emphasis>
(
<emphasis id="93A6EAB8A276FFC2FD91612AFD81AD42" box="[552,614,1078,1101]" italics="true" pageId="11" pageNumber="422">Macr.</emphasis>
)
<taxonomicName id="66D24D29A276FFC2FDCF612AFC11AD42" authority="Angelini &amp; DeMarzo, 1981" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[630,1014,1077,1101]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="93A6EAB8A276FFC2FDCF612AFD21AD42" box="[630,710,1078,1101]" italics="true" pageId="11" pageNumber="422">caelebs</emphasis>
Angelini &amp; DeMarzo, 1981
</taxonomicName>
and in
<emphasis id="93A6EAB8A276FFC2FFC8614AFF6EAD62" box="[113,137,1110,1133]" italics="true" pageId="11" pageNumber="422">A.</emphasis>
(
<emphasis id="93A6EAB8A276FFC2FF28614AFF37AD62" box="[145,208,1110,1133]" italics="true" pageId="11" pageNumber="422">Macr.</emphasis>
)
<taxonomicName id="66D24D29A276FFC2FF59614AFD87AD62" authority="Angelini &amp; DeMarzo, 1981" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[224,608,1109,1133]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="sherpa">
<emphasis id="93A6EAB8A276FFC2FF59614AFECEAD62" box="[224,297,1110,1133]" italics="true" pageId="11" pageNumber="422">sherpa</emphasis>
Angelini &amp; DeMarzo, 1981
</taxonomicName>
. The new species is also similar to both the above species in the body size, the length ratio of antennomeres III: II (
<quantity id="662A9B4FA276FFC2FC306169FC2EAD82" box="[905,969,1141,1165]" metricMagnitude="-2" metricUnit="m" metricValue="3.5559999999999996" pageId="11" pageNumber="422" unit="in" value="1.4">1.4 in</quantity>
<taxonomicName id="66D24D29A276FFC2FC68616AFBA6AD82" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[977,1089,1142,1165]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="93A6EAB8A276FFC2FC68616AFBA6AD82" box="[977,1089,1142,1165]" italics="true" pageId="11" pageNumber="422">A. caelebs</emphasis>
</taxonomicName>
and
<quantity id="662A9B4FA276FFC2FF276189FF3DADA2" box="[158,218,1173,1197]" metricMagnitude="-2" metricUnit="m" metricValue="4.064" pageId="11" pageNumber="422" unit="in" value="1.6">1.6 in</quantity>
<taxonomicName id="66D24D29A276FFC2FF66618AFEA2ADA2" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[223,325,1174,1197]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="sherpa">
<emphasis id="93A6EAB8A276FFC2FF66618AFEA2ADA2" box="[223,325,1174,1197]" italics="true" pageId="11" pageNumber="422">A. sherpa</emphasis>
</taxonomicName>
), absence of microsculpture on the dorsum, the missing sutural stria and clypeal line. The new species differs from both above species by the shape of the pronotum, having the lateral margin almost angulate (in contrast to broadly rounded lateral margins in both species compared). The new species also differs in the shape of the spermatheca, which is almost spherical in it basal part and twisted in its proximal part (in contrast to the is simply horseshoe-shaped spermatheca in
<taxonomicName id="66D24D29A276FFC2FE63602AFDA7AC42" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[474,576,1334,1357]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="sherpa">
<emphasis id="93A6EAB8A276FFC2FE63602AFDA7AC42" box="[474,576,1334,1357]" italics="true" pageId="11" pageNumber="422">A. sherpa</emphasis>
</taxonomicName>
). The spermatheca of
<taxonomicName id="66D24D29A276FFC2FC92602AFC34AC42" authorityName="Švec" authorityYear="2012" box="[811,979,1334,1357]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="acutangulum">
<emphasis id="93A6EAB8A276FFC2FC92602AFC34AC42" box="[811,979,1334,1357]" italics="true" pageId="11" pageNumber="422">A. acutangulum</emphasis>
</taxonomicName>
is slightly similar to that of
<taxonomicName id="66D24D29A276FFC2FE95604AFE7CAC62" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[300,411,1366,1389]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="93A6EAB8A276FFC2FE95604AFE7CAC62" box="[300,411,1366,1389]" italics="true" pageId="11" pageNumber="422">A. caelebs</emphasis>
</taxonomicName>
, but both species differ by the tarsal formula in females (4-4-
<quantity id="662A9B4FA276FFC2FB8B6049FF61AC82" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="11" pageNumber="422" unit="in" value="4.0">4 in</quantity>
the new species, 5-4-
<quantity id="662A9B4FA276FFC2FED46069FE70AC82" box="[365,407,1397,1421]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="11" pageNumber="422" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="66D24D29A276FFC2FE24606AFDECAC82" authorityName="Angelini &amp; DeMarzo" authorityYear="1981" box="[413,523,1398,1421]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="422" phylum="Arthropoda" rank="species" species="caelebs">
<emphasis id="93A6EAB8A276FFC2FE24606AFDECAC82" box="[413,523,1398,1421]" italics="true" pageId="11" pageNumber="422">A. caelebs</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E9C86521A276FFC2FFC86089FCA1AF42" pageId="11" pageNumber="422" type="etymology">
<paragraph id="A16D36AAA276FFC2FFC86089FF21ACC2" blockId="11.[113,1089,789,1613]" pageId="11" pageNumber="422">
<emphasis id="93A6EAB8A276FFC2FFC86089FF15ACA2" bold="true" box="[113,242,1429,1453]" pageId="11" pageNumber="422">Etymology.</emphasis>
The name refers to the shape of the lateral margin of the pronotum of the new species.
</paragraph>
<paragraph id="A16D36AAA276FFC2FFC860C9FCA1AF42" blockId="11.[113,1089,789,1613]" pageId="11" pageNumber="422">
<emphasis id="93A6EAB8A276FFC2FFC860C9FF08ACE2" bold="true" box="[113,239,1493,1517]" pageId="11" pageNumber="422">Bionomics.</emphasis>
The specimens were collected by the sifting of wet leaf litter in a primary evergreen mountain cloud forest, with very sparse understory vegetation, and by the sifting of leaf litter and fallen
<taxonomicName id="66D24D29A276FFC2FEAE630AFE52AF22" box="[279,437,1558,1581]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="11" pageNumber="422" phylum="Tracheophyta" rank="genus">
<emphasis id="93A6EAB8A276FFC2FEAE630AFE52AF22" box="[279,437,1558,1581]" italics="true" pageId="11" pageNumber="422">Rhododendron</emphasis>
</taxonomicName>
petals in the small remnants of high
<emphasis id="93A6EAB8A276FFC2FC8B630AFBA6AF22" box="[818,1089,1558,1581]" italics="true" pageId="11" pageNumber="422">
<taxonomicName id="66D24D29A276FFC2FC8B630AFC70AF22" authorityName="C.Linnaeus" authorityYear="1753" box="[818,919,1558,1581]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="422" phylum="Tracheophyta" rank="genus">Juniperus</taxonomicName>
/
<taxonomicName id="66D24D29A276FFC2FC1D630AFBA6AF22" box="[932,1089,1558,1581]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="11" pageNumber="422" phylum="Tracheophyta" rank="genus">Rhododendron</taxonomicName>
</emphasis>
forest with sparse understory vegetation (M. Fikáček, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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