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<document id="018DEE7B0E29384B34B065295A7DFF18" ID-DOI="10.5852/ejt.2016.204" ID-GBIF-Dataset="9b814d4f-378b-4e95-8144-1e6b183711d5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3840075" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1590162337488" checkinUser="valdenar" docAuthor="Riaux-Gobin, Catherine, Compère, Pierre, Jordan, Richard W., Coste, Michel &amp; Yesilyurt, Jovita C." docDate="2016" docId="003BB414F62E3E6B575A0BB6FAC8C325" docLanguage="en" docName="ejt-2016-204.pdf.imf" docOrigin="European Journal of Taxonomy 204" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cocconeis dirupta W. Greg. (Gregory 1857" docType="treatment" docVersion="10" lastPageNumber="14" masterDocId="FC02CC6CF6243E6655420866FFB9C13C" masterDocTitle="Cocconeis molesta Kütz., C. diaphana W. Sm. and C. dirupta W. Greg. (Bacillariophyta): type material, ambiguities and possible synonymies" masterLastPageNumber="18" masterPageNumber="1" pageNumber="11" updateTime="1734455776656" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A1579A2261FE432A8D3E00007E0D8810">Cocconeis molesta Kütz., C. diaphana W. Sm. and C. dirupta W. Greg. (Bacillariophyta): type material, ambiguities and possible synonymies</mods:title>
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<mods:namePart id="5D26187CDE844394987D0610ECF90FB8">Riaux-Gobin, Catherine</mods:namePart>
<mods:affiliation id="B6286E9FC5E538CE62196D50EF697A6A">CRIOBE-USR 3278 CNRS-EPHE-UPVD, 66000 Perpignan, France. &amp; Laboratoire dExcellence CORAIL, 66000 Perpignan, France.</mods:affiliation>
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<mods:namePart id="681AB0B36B17CA56278F398188F9A317">Compère, Pierre</mods:namePart>
<mods:affiliation id="2C5A6A6991E613DFC5C5DF72BA0ACD08">Deceased 29 April 2016.</mods:affiliation>
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<mods:namePart id="64CEE51933ED1DDBFDC68500570100FC">Jordan, Richard W.</mods:namePart>
<mods:affiliation id="BEA2A6E7745AC44AAF4828E768FAF663">Department of Earth &amp; Environmental Sciences, Yamagata University, Yamagata 990 - 8560, Japan.</mods:affiliation>
<mods:nameIdentifier id="B08CCF9C9B293BE851A998746B2C1EAE" type="email">sh081@kdw.kj.yamagata-u.ac.jp</mods:nameIdentifier>
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<mods:namePart id="1A36193850E17B3D3786477921CADF62">Coste, Michel</mods:namePart>
<mods:affiliation id="0D6819E50447563C9D73E627F0FE2E25">IRSTEA, 50 avenue de Verdun, Cestas, France. &amp; The Natural History Museum, Department of Life Sciences, Cromwell Rd, SW 7 5 BD, London, United Kingdom.</mods:affiliation>
<mods:nameIdentifier id="FF6065F9F45517EC081493688D98BD87" type="email">michel.coste@irstea.fr&amp;j.yesilyurt@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="9DE2FCCD370CF95A8DDDE49B6EEA1DB4">Yesilyurt, Jovita C.</mods:namePart>
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<mods:title id="63596056BE19F422859FFC88F53EAEE7">European Journal of Taxonomy</mods:title>
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<mods:date id="18D1D35C3236716919B579A0BDAA1D51">2016</mods:date>
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<treatment id="003BB414F62E3E6B575A0BB6FAC8C325" ID-DOI="http://doi.org/10.5281/zenodo.3853040" ID-GBIF-Taxon="164249563" ID-Zenodo-Dep="3853040" LSID="urn:lsid:plazi:treatment:003BB414F62E3E6B575A0BB6FAC8C325" httpUri="http://treatment.plazi.org/id/003BB414F62E3E6B575A0BB6FAC8C325" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" scope_class="Bacillariophyceae" scope_family="Cocconeidaceae" scope_subClass="Bacillariophycidae">
<subSubSection id="C0885689F62E3E6C575A0BB6FBA2C2D6" box="[536,1051,975,1002]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="882D0502F62E3E6C575A0BB6FBA2C2D6" blockId="10.[536,1051,975,1038]" box="[536,1051,975,1002]" pageId="10" pageNumber="11">
<heading id="D365B26EF62E3E6C575A0BB6FBA2C2D6" box="[536,1051,975,1002]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="11" reason="2">
<taxonomicName id="4F927E81F62E3E6C575A0BB6FBA2C2D6" authority="W. Greg. (Gregory 1857)" authorityName="W. Greg. (Gregory" authorityYear="1857" box="[536,1051,975,1002]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="dirupta">
<emphasis id="BAE6D910F62E3E6C575A0BB6FD4AC2D6" bold="true" box="[536,755,975,1002]" italics="true" pageId="10" pageNumber="11">Cocconeis dirupta</emphasis>
W.Greg. (
<bibRefCitation id="EC0378F3F62E3E6C562E0BB6FBAAC2D6" author="Gregory W." box="[876,1043,975,1002]" pageId="10" pageNumber="11" pagination="473 - 542" refId="ref7707" refString="Gregory W. 1857. On new forms of marine Diatomaceae, found in the Firth of Clyde and in Loch Fyne, illustrated by numerous figures drawn by R. K. Greville, L. L. D., F. R. S. E. Transactions of the Royal Society of Edinburgh 21: 473 - 542." type="journal article" year="1857">Gregory 1857</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C0885689F62E3E6C57D50B95FC24C532" box="[663,925,1011,1038]" pageId="10" pageNumber="11" type="description">
<paragraph id="882D0502F62E3E6C57D50B95FC24C532" blockId="10.[536,1051,975,1038]" box="[663,925,1011,1038]" pageId="10" pageNumber="11">
<figureCitation id="10A91987F62E3E6C57D50B95FD5BC532" box="[663,738,1011,1038]" captionStart="Figs 18" captionStartId="2.[189,241,1895,1921]" captionTargetBox="[351,1203,1002,1833]" captionTargetId="figure@2.[312,1275,990,1851]" captionTargetPageId="2" captionText="Figs 18. Original drawings. 1. Cocconeis molesta Kütz. (Kützing 1844, pl. 5, fig. 7). 23. Cocconeis diaphana W.Sm. (Smith 1853, pl. 30): 2. var. β. 3. type C. diaphana var. diaphana. 4. Cocconeis dirupta W.Greg. (Gregory 1857, pl. 9). 58. Cocconeis molesta var. crucifera Grunow ex Cleve (Van Heurck 18801885, pl. 30): 56. f. minor. 78. f. major." figureDoi="http://doi.org/10.5281/zenodo.3840079" httpUri="https://zenodo.org/record/3840079/files/figure.png" pageId="10" pageNumber="11">Figs 4</figureCitation>
,
<figureCitation id="10A91987F62E3E6C57AC0B95FC83C532" box="[750,826,1011,1038]" captionStart="Figs 3538" captionStartId="11.[189,241,1895,1921]" captionTargetBox="[189,1398,666,1854]" captionTargetId="figure@11.[189,1398,666,1854]" captionTargetPageId="11" captionText="Figs 3538. Cocconeis dirupta W.Greg. from BM 1420. 37. Lectotype illustration. SV sternum lanceolate with apices slightly bent in opposite directions (36), one SV stria lacking on one side (35, 36, arrow), RV striae strongly radiate, RV fascia narrow and extended (38, arrows), helictoglossae deflected in opposite directions (37, arrowheads) and raphe slightly sigmoid (37). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3840091" httpUri="https://zenodo.org/record/3840091/files/figure.png" pageId="10" pageNumber="11">3538</figureCitation>
,
<tableCitation id="C51030B9F62E3E6C56040B95FC24C532" box="[838,925,1011,1038]" captionStart="Table 1" captionStartId="4.[187,256,611,639]" captionTargetId="graphics@4.[187,1943,673,1710]" captionTargetPageId="4" captionText="Table 1. Comparison of morphological features and biometric data of the examined type slide taxa." httpUri="http://table.plazi.org/id/DCED558AF6203E6255F90A05FAF0C343" pageId="10" pageNumber="11" tableUuid="DCED558AF6203E6255F90A05FAF0C343">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0885689F62E3E6C55FF0C50FCDBC56D" box="[189,866,1078,1105]" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="882D0502F62E3E6C55FF0C50FCDBC56D" blockId="10.[189,866,1078,1105]" box="[189,866,1078,1105]" pageId="10" pageNumber="11">
<taxonomicName id="4F927E81F62E3E6C55FF0C50FDBDC56C" authority="W. Sm." authorityName="W. Sm. (Smith" authorityYear="1853" box="[189,516,1078,1104]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="diaphana">
<emphasis id="BAE6D910F62E3E6C55FF0C50FE17C56C" box="[189,430,1078,1104]" italics="true" pageId="10" pageNumber="11">Cocconeis diaphana</emphasis>
W.Sm.
</taxonomicName>
var. β (“Sidmouth material”).
</paragraph>
</subSubSection>
<subSubSection id="C0885689F62E3E6C55FF0C1EFB0AC46D" pageId="10" pageNumber="11" type="description">
<paragraph id="882D0502F62E3E6C55FF0C1EFE03C5AE" blockId="10.[189,442,1144,1170]" box="[189,442,1144,1170]" pageId="10" pageNumber="11">
<heading id="D365B26EF62E3E6C55FF0C1EFE03C5AE" bold="true" box="[189,442,1144,1170]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="BAE6D910F62E3E6C55FF0C1EFE03C5AE" bold="true" box="[189,442,1144,1170]" pageId="10" pageNumber="11">Original description</emphasis>
</heading>
</paragraph>
<paragraph id="882D0502F62E3E6C55FF0CCEFC3AC5FE" blockId="10.[189,1399,1191,1362]" box="[189,899,1191,1218]" pageId="10" pageNumber="11">
<bibRefCitation id="EC0378F3F62E3E6C55FF0CCEFEC0C5FE" author="Gregory W." box="[189,377,1191,1218]" pageId="10" pageNumber="11" pagination="473 - 542" refId="ref7707" refString="Gregory W. 1857. On new forms of marine Diatomaceae, found in the Firth of Clyde and in Loch Fyne, illustrated by numerous figures drawn by R. K. Greville, L. L. D., F. R. S. E. Transactions of the Royal Society of Edinburgh 21: 473 - 542." type="journal article" year="1857">Gregory (1857)</bibRefCitation>
: 491, 492, pl. 9, fig. 25 reproduced in
<figureCitation id="10A91987F62E3E6C567A0CC1FCC6C5FE" box="[824,895,1191,1218]" captionStart="Figs 18" captionStartId="2.[189,241,1895,1921]" captionTargetBox="[351,1203,1002,1833]" captionTargetId="figure@2.[312,1275,990,1851]" captionTargetPageId="2" captionText="Figs 18. Original drawings. 1. Cocconeis molesta Kütz. (Kützing 1844, pl. 5, fig. 7). 23. Cocconeis diaphana W.Sm. (Smith 1853, pl. 30): 2. var. β. 3. type C. diaphana var. diaphana. 4. Cocconeis dirupta W.Greg. (Gregory 1857, pl. 9). 58. Cocconeis molesta var. crucifera Grunow ex Cleve (Van Heurck 18801885, pl. 30): 56. f. minor. 78. f. major." figureDoi="http://doi.org/10.5281/zenodo.3840079" httpUri="https://zenodo.org/record/3840079/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
.
</paragraph>
<paragraph id="882D0502F62E3E6C55FF0CADFB0AC46D" blockId="10.[189,1399,1191,1362]" pageId="10" pageNumber="11">
“Broad short oval to orbicular. Length
<quantity id="4F6AA8E7F62E3E6C57CB0CADFC8DC5DA" box="[649,820,1227,1254]" metricMagnitude="-2" metricUnit="m" metricValue="4.32" metricValueMax="6.1" metricValueMin="2.54" pageId="10" pageNumber="11" unit="mm" value="43.2" valueMax="61.0" valueMin="25.4">25.4 to 61 µm</quantity>
; breadth
<quantity id="4F6AA8E7F62E3E6C56E30CAAFBDAC5DA" box="[929,1123,1227,1254]" metricMagnitude="-2" metricUnit="m" metricValue="3.5549999999999997" metricValueMax="5.33" metricValueMin="1.78" pageId="10" pageNumber="11" unit="mm" value="35.55" valueMax="53.3" valueMin="17.8">17.8 to 53.3 µm</quantity>
”. “Valve thick. Median line irregular, like a slit. Wavy longitudinal striae, except the slit. Fine transverse striae. Appearance of a long stauros, which, under a higher power, disappears as such. The striated surface seems to be torn asunder in the middle. Vertical striae about 26, transverse striae about
<quantity id="4F6AA8E7F62E3E6C56BC0D51FB80C46D" box="[1022,1081,1335,1361]" metricMagnitude="0" metricUnit="m" metricValue="1.524" pageId="10" pageNumber="11" unit="in" value="60.0">60 in</quantity>
1 c.d.m.”.
</paragraph>
</subSubSection>
<subSubSection id="C0885689F62E3E6C55FF0D1CFEE5C4D4" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="882D0502F62E3E6C55FF0D1CFED5C4A8" blockId="10.[189,364,1402,1428]" box="[189,364,1402,1428]" pageId="10" pageNumber="11">
<heading id="D365B26EF62E3E6C55FF0D1CFED5C4A8" bold="true" box="[189,364,1402,1428]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="BAE6D910F62E3E6C55FF0D1CFED5C4A8" bold="true" box="[189,364,1402,1428]" pageId="10" pageNumber="11">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="882D0502F62E3E6C55FF0DCCFEE5C4D4" blockId="10.[189,1398,1449,1512]" pageId="10" pageNumber="11">
<materialsCitation id="38FA0F5FF62E3E6C55FF0DCCFEE5C4D4" ID-GBIF-Occurrence="2625225301" collectionCode="BM" pageId="10" pageNumber="11" specimenCode="BM 1420" specimenCount="1" typeStatus="lectotype">
<typeStatus id="5729BBA0F62E3E6C55FF0DCCFE8CC4F8" box="[189,309,1450,1476]" pageId="10" pageNumber="11" type="lectotype">Lectotype</typeStatus>
, designated here: slide
<specimenCode id="D834AD79F62E3E6C57060DCCFD0FC4FF" box="[580,694,1449,1476]" collectionCode="BM" country="United Kingdom" name="Bristol Museum" pageId="10" pageNumber="11">BM 1420</specimenCode>
, labelled “Arran 56” in Greville collection, the only syntype found in
<collectionCode id="EE839DC7F62E3E6C54650DA8FEE1C4D4" box="[295,344,1486,1512]" country="United Kingdom" name="Bristol Museum" pageId="10" pageNumber="11">BM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C0885689F62E3E6C55FF0E76FE9FC631" pageId="10" pageNumber="11" type="discussion">
<paragraph id="882D0502F62E3E6C55FF0E76FEBAC716" blockId="10.[189,259,1552,1578]" box="[189,259,1552,1578]" pageId="10" pageNumber="11">
<heading id="D365B26EF62E3E6C55FF0E76FEBAC716" bold="true" box="[189,259,1552,1578]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="BAE6D910F62E3E6C55FF0E76FEBAC716" bold="true" box="[189,259,1552,1578]" pageId="10" pageNumber="11">Notes</emphasis>
</heading>
</paragraph>
<paragraph id="882D0502F62E3E6C55FF0E59FE9FC631" blockId="10.[189,1399,1599,1805]" pageId="10" pageNumber="11">
Several localities (Glenshira sand, Mr Miless Corallina gathering and several dredgings) cited by Gregory in the original description could contain
<typeStatus id="5729BBA0F62E3E6C57BF0E05FCDEC741" box="[765,871,1635,1661]" pageId="10" pageNumber="11" type="syntype">syntypes</typeStatus>
, but only the one designated as
<typeStatus id="5729BBA0F62E3E6C51950E05FAFAC741" box="[1239,1347,1635,1661]" pageId="10" pageNumber="11" type="lectotype">lectotype</typeStatus>
was found in BM. According to W. Gregory “
<taxonomicName id="4F927E81F62E3E6C57E70EE1FC8FC79D" authorityName="W. Sm. (Smith" authorityYear="1853" box="[677,822,1671,1697]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="diaphana">
<emphasis id="BAE6D910F62E3E6C57E70EE1FC8FC79D" box="[677,822,1671,1697]" italics="true" pageId="10" pageNumber="11">C. diaphana</emphasis>
</taxonomicName>
[…] which was found occurring with
<taxonomicName id="4F927E81F62E3E6C51BF0EE1FACFC79D" authorityName="W. Greg. (Gregory" authorityYear="1857" box="[1277,1398,1671,1697]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="dirupta">
<emphasis id="BAE6D910F62E3E6C51BF0EE1FACFC79D" box="[1277,1398,1671,1697]" italics="true" pageId="10" pageNumber="11">C. dirupta</emphasis>
</taxonomicName>
[…] may perhaps be an imperfect form of
<taxonomicName id="4F927E81F62E3E6C57900ECDFCF6C7F9" authorityName="W. Greg. (Gregory" authorityYear="1857" box="[722,847,1707,1733]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="10" pageNumber="11" phylum="Ochrophyta" rank="species" species="dirupta">
<emphasis id="BAE6D910F62E3E6C57900ECDFCF6C7F9" box="[722,847,1707,1733]" italics="true" pageId="10" pageNumber="11">C. dirupta</emphasis>
</taxonomicName>
[…] or possibly [...] the lower valve of the latter”, with some further notes that “it is, however equally probable that these forms belong to different species”.
</paragraph>
</subSubSection>
<subSubSection id="C0885689F62E3E6B55FF0F53FAC8C325" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11" type="description">
<paragraph id="882D0502F62E3E6C55FF0F53FC64C673" blockId="10.[189,989,1845,1871]" box="[189,989,1845,1871]" pageId="10" pageNumber="11">
<heading id="D365B26EF62E3E6C55FF0F53FC64C673" bold="true" box="[189,989,1845,1871]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="BAE6D910F62E3E6C55FF0F53FC64C673" bold="true" box="[189,989,1845,1871]" pageId="10" pageNumber="11">LM examination of the type material and completed description</emphasis>
</heading>
</paragraph>
<paragraph id="882D0502F62E3E6D55FF0F03FDE7C303" blockId="10.[189,1399,1892,2027]" lastBlockId="11.[189,1399,267,575]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<figureCitation id="10A91987F62E3E6C55FF0F03FEFAC643" box="[189,323,1892,1919]" captionStart="Figs 3538" captionStartId="11.[189,241,1895,1921]" captionTargetBox="[189,1398,666,1854]" captionTargetId="figure@11.[189,1398,666,1854]" captionTargetPageId="11" captionText="Figs 3538. Cocconeis dirupta W.Greg. from BM 1420. 37. Lectotype illustration. SV sternum lanceolate with apices slightly bent in opposite directions (36), one SV stria lacking on one side (35, 36, arrow), RV striae strongly radiate, RV fascia narrow and extended (38, arrows), helictoglossae deflected in opposite directions (37, arrowheads) and raphe slightly sigmoid (37). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3840091" httpUri="https://zenodo.org/record/3840091/files/figure.png" pageId="10" pageNumber="11">Figs 3538</figureCitation>
, from BM 1420: n = 16, valves are 18.336.5 (20 ± 6) µm long, 17.230.8 (26.6 ± 4.7) µm wide; with 13.519.5 (16.6 ± 2) SV striae in
<quantity id="4F6AA8E7F62E3E6C57980FEFFC92C69F" box="[730,811,1929,1955]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="10.0">10 µm</quantity>
, and 1623 (18.9 ± 1.9) RV striae in
<quantity id="4F6AA8E7F62E3E6C51AF0FEFFA86C69F" box="[1261,1343,1929,1955]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="11" unit="mm" value="10.0">10 µm</quantity>
. L/l: 1.12 ± 0.06. Valves thick, round-elliptic to almost discoid. Valves rarely found separate, appearing darkbrown. SV with coarse radiate striae, equidistant, composed of transapically oblong areolae arranged in a zig-zag pattern along longitudinal lines, areolae smaller near the margin. One median stria often shorter on one side of the central area (
<figureCitation id="10A91987F62F3E6D57650948FD14C075" box="[551,685,302,329]" captionStart="Figs 3538" captionStartId="11.[189,241,1895,1921]" captionTargetBox="[189,1398,666,1854]" captionTargetId="figure@11.[189,1398,666,1854]" captionTargetPageId="11" captionText="Figs 3538. Cocconeis dirupta W.Greg. from BM 1420. 37. Lectotype illustration. SV sternum lanceolate with apices slightly bent in opposite directions (36), one SV stria lacking on one side (35, 36, arrow), RV striae strongly radiate, RV fascia narrow and extended (38, arrows), helictoglossae deflected in opposite directions (37, arrowheads) and raphe slightly sigmoid (37). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3840091" httpUri="https://zenodo.org/record/3840091/files/figure.png" pageId="11" pageNumber="12">Figs 3536</figureCitation>
, arrows). Presence of intercalary short striae near the margin. SV sternum relatively large, fusiform (rarely constricted in its mid-part) with apices often deflected in opposite directions (not illustrated). RV with delicate but clearly identifiable striae, equidistant, strongly bent at the apices, with frequent intercalary short striae near the margin. Raphe almost straight to slightly sigmoid (
<figureCitation id="10A91987F62F3E6D546E09DDFE3DC0E9" box="[300,388,443,469]" captionStart="Figs 3538" captionStartId="11.[189,241,1895,1921]" captionTargetBox="[189,1398,666,1854]" captionTargetId="figure@11.[189,1398,666,1854]" captionTargetPageId="11" captionText="Figs 3538. Cocconeis dirupta W.Greg. from BM 1420. 37. Lectotype illustration. SV sternum lanceolate with apices slightly bent in opposite directions (36), one SV stria lacking on one side (35, 36, arrow), RV striae strongly radiate, RV fascia narrow and extended (38, arrows), helictoglossae deflected in opposite directions (37, arrowheads) and raphe slightly sigmoid (37). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3840091" httpUri="https://zenodo.org/record/3840091/files/figure.png" pageId="11" pageNumber="12">Fig. 37</figureCitation>
). RV axial area narrow. Distal raphe fissures close to the margin. Helictoglossae often clearly deflected in opposite directions (
<figureCitation id="10A91987F62F3E6D57D209B8FD5EC0C5" box="[656,743,478,505]" captionStart="Figs 3538" captionStartId="11.[189,241,1895,1921]" captionTargetBox="[189,1398,666,1854]" captionTargetId="figure@11.[189,1398,666,1854]" captionTargetPageId="11" captionText="Figs 3538. Cocconeis dirupta W.Greg. from BM 1420. 37. Lectotype illustration. SV sternum lanceolate with apices slightly bent in opposite directions (36), one SV stria lacking on one side (35, 36, arrow), RV striae strongly radiate, RV fascia narrow and extended (38, arrows), helictoglossae deflected in opposite directions (37, arrowheads) and raphe slightly sigmoid (37). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3840091" httpUri="https://zenodo.org/record/3840091/files/figure.png" pageId="11" pageNumber="12">Fig. 37</figureCitation>
, arrowheads); presence of a large and narrow transverse fascia (never reaching the margin, formed by one or two shorter striae,
<figureCitation id="10A91987F62F3E6D56B90A64FBE9C320" box="[1019,1104,514,540]" captionStart="Figs 3538" captionStartId="11.[189,241,1895,1921]" captionTargetBox="[189,1398,666,1854]" captionTargetId="figure@11.[189,1398,666,1854]" captionTargetPageId="11" captionText="Figs 3538. Cocconeis dirupta W.Greg. from BM 1420. 37. Lectotype illustration. SV sternum lanceolate with apices slightly bent in opposite directions (36), one SV stria lacking on one side (35, 36, arrow), RV striae strongly radiate, RV fascia narrow and extended (38, arrows), helictoglossae deflected in opposite directions (37, arrowheads) and raphe slightly sigmoid (37). Scale bars = 10 µm." figureDoi="http://doi.org/10.5281/zenodo.3840091" httpUri="https://zenodo.org/record/3840091/files/figure.png" pageId="11" pageNumber="12">Fig. 38</figureCitation>
, arrows); proximal raphe endings coarse and relatively close.
</paragraph>
<caption id="DCED558AF62F3E6D55FF0F01FBC9C6D7" ID-DOI="http://doi.org/10.5281/zenodo.3840091" ID-Zenodo-Dep="3840091" httpUri="https://zenodo.org/record/3840091/files/figure.png" pageId="11" pageNumber="12" startId="11.[189,241,1895,1921]" targetBox="[189,1398,666,1854]" targetPageId="11">
<paragraph id="882D0502F62F3E6D55FF0F01FBC9C6D7" blockId="11.[189,1399,1895,2028]" pageId="11" pageNumber="12">
<emphasis id="BAE6D910F62F3E6D55FF0F01FEFFC6BD" bold="true" box="[189,326,1895,1921]" pageId="11" pageNumber="12">Figs 3538.</emphasis>
<taxonomicName id="4F927E81F62F3E6D54090F0EFD31C6BD" authority="W. Greg." authorityName="W. Greg. (Gregory" authorityYear="1857" box="[331,648,1895,1922]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="11" pageNumber="12" phylum="Ochrophyta" rank="species" species="dirupta">
<emphasis id="BAE6D910F62F3E6D54090F0EFD99C6BD" box="[331,544,1895,1922]" italics="true" pageId="11" pageNumber="12">Cocconeis dirupta</emphasis>
W.Greg.
</taxonomicName>
from BM 1420.
<emphasis id="BAE6D910F62F3E6D56060F01FCD8C6BD" bold="true" box="[836,865,1895,1921]" pageId="11" pageNumber="12">37</emphasis>
. Lectotype illustration. SV sternum lanceolate with apices slightly bent in opposite directions (36), one SV stria lacking on one side (35, 36, arrow), RV striae strongly radiate, RV fascia narrow and extended (38, arrows), helictoglossae deflected in opposite directions (37, arrowheads) and raphe slightly sigmoid (37). Scale bars = 10 µm.
</paragraph>
</caption>
<caption id="DCED558AF6283E6A55FF0EBCFACFC6D7" ID-DOI="http://doi.org/10.5281/zenodo.3840093" ID-Zenodo-Dep="3840093" httpUri="https://zenodo.org/record/3840093/files/figure.png" pageId="12" pageNumber="13" startId="12.[189,241,1754,1780]" targetBox="[295,1293,265,1705]" targetPageId="12">
<paragraph id="882D0502F6283E6A55FF0EBCFACFC6D7" blockId="12.[189,1399,1754,2028]" pageId="12" pageNumber="13">
<emphasis id="BAE6D910F6283E6A55FF0EBCFEF5C7C8" bold="true" box="[189,332,1754,1780]" pageId="12" pageNumber="13">Figs 3944.</emphasis>
<taxonomicName id="4F927E81F6283E6A54150EBDFD1DC7C8" authority="W. Sm." authorityName="W. Sm. (Smith" authorityYear="1853" box="[343,676,1754,1781]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="species" species="diaphana">
<emphasis id="BAE6D910F6283E6A54150EBDFDF2C7C8" box="[343,587,1754,1781]" italics="true" pageId="12" pageNumber="13">Cocconeis diaphana</emphasis>
W.Sm.
</taxonomicName>
isolectotype SEM illustration. Mica labelled as “
<taxonomicName id="4F927E81F6283E6A51BE0EBDFE95C62B" authority="Riaux-Gobin &amp; Compère &amp; Jordan &amp; Coste &amp; Yesilyurt Criobe-Usr &amp; Irstea, 2016" authorityName="Riaux-Gobin &amp; Compère &amp; Jordan &amp; Coste &amp; Yesilyurt Criobe-Usr &amp; Irstea" authorityYear="2016" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="12" pageNumber="13" phylum="Ochrophyta" rank="species" species="diaphana" status="sp. nov.">
<emphasis id="BAE6D910F6283E6A51BE0EBDFE95C62B" italics="true" pageId="12" pageNumber="13">Cocconeis diaphana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1D5646BF6283E6A54720E98FED3C624" box="[304,362,1790,1816]" pageId="12" pageNumber="13" rank="species">n.sp.</taxonomicNameLabel>
, Jersey, Aug. 14. 1852”, deposited in the Van Heurck collection in Meise (BR). SV external view with a reduced central area (39, arrow) and striae composed of transversally elongated alveoli (39). SV internal view (40). Note the SV valvocopula (SVVC) with a smooth edge (40, arrowhead). Detail of the SV apex (41). Detail of a broken SV showing the double layered structure of the alveoli (42). RV external view with the central oblong-elongate central area reaching less than ¼ of the valve width (43, arrow). RV internal view with a low and straight helictoglossa (44, arrow) and the RV valvocopula (RVVC) with a smooth edge (44, arrowhead). Scale bars = 10 µm (39, 40, 4344), 2 µm (41), 1 µm (42).
</paragraph>
</caption>
<paragraph id="882D0502F6293E6B55FF096DFBEDC088" blockId="13.[189,1398,267,436]" pageId="13" pageNumber="14">
The complete frustules appear obscure while the separate RV and SV appear translucent, which probably justifies W. Gregorys hesitation about the existence of a second taxon (
<taxonomicName id="4F927E81F6293E6B516C0949FAA6C075" authority="W. Sm." authorityName="W. Sm. (Smith" authorityYear="1853" box="[1070,1311,303,329]" class="Bacillariophyceae" family="Cocconeidaceae" genus="Cocconeis" higherTaxonomySource="GBIF" kingdom="Chromista" order="Achnanthales" pageId="13" pageNumber="14" phylum="Ochrophyta" rank="species" species="diaphana">
<emphasis id="BAE6D910F6293E6B516C0949FB7AC075" box="[1070,1219,303,329]" italics="true" pageId="13" pageNumber="14">C. diaphana</emphasis>
W.Sm.
</taxonomicName>
) in his material. On the other hand the R.K. Greville drawing (
<bibRefCitation id="EC0378F3F6293E6B560C0935FC40C051" author="Gregory W." box="[846,1017,338,365]" pageId="13" pageNumber="14" pagination="473 - 542" refId="ref7707" refString="Gregory W. 1857. On new forms of marine Diatomaceae, found in the Firth of Clyde and in Loch Fyne, illustrated by numerous figures drawn by R. K. Greville, L. L. D., F. R. S. E. Transactions of the Royal Society of Edinburgh 21: 473 - 542." type="journal article" year="1857">Gregory 1857</bibRefCitation>
: fig. 25) probably superimposed the two valves. Contrarily to the latter drawing (
<bibRefCitation id="EC0378F3F6293E6B57B70910FC27C0AD" author="Gregory W." box="[757,926,374,401]" pageId="13" pageNumber="14" pagination="473 - 542" refId="ref7707" refString="Gregory W. 1857. On new forms of marine Diatomaceae, found in the Firth of Clyde and in Loch Fyne, illustrated by numerous figures drawn by R. K. Greville, L. L. D., F. R. S. E. Transactions of the Royal Society of Edinburgh 21: 473 - 542." type="journal article" year="1857">Gregory 1857</bibRefCitation>
, fig. 25 reproduced in
<figureCitation id="10A91987F6293E6B51EA0910FB49C0AC" box="[1192,1264,374,400]" captionStart="Figs 18" captionStartId="2.[189,241,1895,1921]" captionTargetBox="[351,1203,1002,1833]" captionTargetId="figure@2.[312,1275,990,1851]" captionTargetPageId="2" captionText="Figs 18. Original drawings. 1. Cocconeis molesta Kütz. (Kützing 1844, pl. 5, fig. 7). 23. Cocconeis diaphana W.Sm. (Smith 1853, pl. 30): 2. var. β. 3. type C. diaphana var. diaphana. 4. Cocconeis dirupta W.Greg. (Gregory 1857, pl. 9). 58. Cocconeis molesta var. crucifera Grunow ex Cleve (Van Heurck 18801885, pl. 30): 56. f. minor. 78. f. major." figureDoi="http://doi.org/10.5281/zenodo.3840079" httpUri="https://zenodo.org/record/3840079/files/figure.png" pageId="13" pageNumber="14">Fig. 4</figureCitation>
), the fascia does not reach the valve margin in the many specimens observed in this study.
</paragraph>
<paragraph id="882D0502F6293E6B55FF09BDFAC8C325" blockId="13.[189,1398,475,538]" pageId="13" pageNumber="14">
The slide BM 1420 from Arran 56 (Gregory in Greville collection) is designated here as the
<typeStatus id="5729BBA0F6293E6B504609BDFACAC0C9" box="[1284,1395,475,501]" pageId="13" pageNumber="14" type="lectotype">lectotype</typeStatus>
, since nothing else has been traced that could be considered as original material studied by W. Gregory.
</paragraph>
</subSubSection>
</treatment>
</document>